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<document id="6AEAC5C181C7194FC777500CA04FD864" ID-CLB-Dataset="8333" ID-DOI="10.11646/zootaxa.4920.3.4" ID-GBIF-Dataset="37eefe05-4ff9-4762-a724-07deef527831" ID-ISSN="1175-5326" ID-Zenodo-Dep="4478107" ID-ZooBank="845CF8B2-07CD-45DC-8128-2EA56C825356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611900587808" checkinUser="plazi" docAuthor="Sommer, David, Hillert, Oliver, Hrůzová, Lucie &amp; Král, David" docDate="2021" docId="039387DBFF9DAD24D8DA8120FD1AFEFB" docLanguage="en" docName="zootaxa.4920.3.4.pdf" docOrigin="Zootaxa 4920 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolbelasmus (Bolbelasmus) nireus" docType="treatment" docVersion="9" lastPageNumber="386" masterDocId="FFAAFFA3FF9BAD22D84D8542FFA6F90D" masterDocTitle="Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus" masterLastPageNumber="394" masterPageNumber="380" pageNumber="386" updateTime="1698925811253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E625436B92D5F9767E7A134FB2E7679">Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus</mods:title>
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<mods:namePart id="F6C80CA55B9BBAAF871B3B56ED82AB25">Sommer, David</mods:namePart>
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<mods:affiliation id="C60ED2EC780E211D53BECF7889DB7F25">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 43 Praha 2, Czech Republic. dejv. sommer @ gmail. com; https: // orcid. org / 0000 - 0003 - 4519 - 9677</mods:affiliation>
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<mods:namePart id="12E2D84AB809C0EB139E459CAAD81BE4">Hillert, Oliver</mods:namePart>
<mods:affiliation id="655AF4186CCFEADFF324E71D17A07923">Kieferndamm 10, D- 15566 Schöneiche bei Berlin, Germany.</mods:affiliation>
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<mods:namePart id="0670997A9953A1D1215A33D9D93F48A4">Hrůzová, Lucie</mods:namePart>
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<mods:affiliation id="DE45F215ABAE66D9813CD275F8E48ADF">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 43 Praha 2, Czech Republic. dejv. sommer @ gmail. com; https: // orcid. org / 0000 - 0003 - 4519 - 9677</mods:affiliation>
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<mods:namePart id="87408DBC9A3E077E6F016A01B6F77B26">Král, David</mods:namePart>
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<mods:affiliation id="7BAC4478EFA1983A355517CE4BF3E153">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 43 Praha 2, Czech Republic. dejv. sommer @ gmail. com; https: // orcid. org / 0000 - 0003 - 4519 - 9677</mods:affiliation>
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<mods:date id="1B70CA365CDB28D10A4821B296B9EA5F">2021</mods:date>
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<treatment id="039387DBFF9DAD24D8DA8120FD1AFEFB" ID-DOI="http://doi.org/10.5281/zenodo.4531753" ID-GBIF-Taxon="177316867" ID-Zenodo-Dep="4531753" LSID="urn:lsid:plazi:treatment:039387DBFF9DAD24D8DA8120FD1AFEFB" httpUri="http://treatment.plazi.org/id/039387DBFF9DAD24D8DA8120FD1AFEFB" lastPageNumber="386" pageId="6" pageNumber="386">
<subSubSection id="C3206546FF9DAD24D8DA8120FD53FD71" box="[151,757,1122,1149]" pageId="6" pageNumber="386" type="nomenclature">
<paragraph id="8B8536CDFF9DAD24D8DA8120FD53FD71" blockId="6.[151,757,1122,1186]" box="[151,757,1122,1149]" pageId="6" pageNumber="386">
<heading id="D0CD81A1FF9DAD24D8DA8120FD53FD71" bold="true" box="[151,757,1122,1149]" fontSize="11" level="1" pageId="6" pageNumber="386" reason="1">
<taxonomicName id="4C3A4D4EFF9DAD24D8DA8120FD53FD71" authority="(Reitter, 1895)" baseAuthorityName="Reitter" baseAuthorityYear="1895" box="[151,757,1122,1149]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="nireus" subGenus="Bolbelasmus">
<emphasis id="B94EEADFFF9DAD24D8DA8120FD53FD71" bold="true" box="[151,757,1122,1149]" pageId="6" pageNumber="386">
<emphasis id="B94EEADFFF9DAD24D8DA8120FE94FD70" bold="true" box="[151,306,1122,1149]" italics="true" pageId="6" pageNumber="386">Bolbelasmus</emphasis>
(
<emphasis id="B94EEADFFF9DAD24D90E8120FE78FD70" bold="true" box="[323,478,1122,1149]" italics="true" pageId="6" pageNumber="386">Bolbelasmus</emphasis>
)
<emphasis id="B94EEADFFF9DAD24D9A28121FD9AFD70" bold="true" box="[495,572,1123,1149]" italics="true" pageId="6" pageNumber="386">nireus</emphasis>
(
<bibRefCitation id="EFAB4B3CFF9DAD24DA068120FD4BFD71" author="Reitter, E." box="[587,749,1122,1148]" pageId="6" pageNumber="386" pagination="79 - 88" refId="ref8665" refString="Reitter, E. (1895) Beschreibung neuer oder wenig gekannter Coleoptera aus der Umgebung von Akbes in Syrien. Wiener Entomologische Zeitung, 14, 79 - 88. https: // doi. org / 10.5962 / bhl. part. 13210" type="journal article" year="1895">Reitter, 1895</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3206546FF9DAD24D8DA81C5FD4CFDAC" box="[151,746,1159,1186]" pageId="6" pageNumber="386" type="description">
<paragraph id="8B8536CDFF9DAD24D8DA81C5FD4CFDAC" blockId="6.[151,757,1122,1186]" box="[151,746,1159,1186]" pageId="6" pageNumber="386">
(
<figureCitation id="13012A48FF9DAD24D8D281C5FF4EFDAC" box="[159,232,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[166,1419,193,1903]" captionTargetId="figure-17@2.[153,1435,179,1918]" captionTargetPageId="2" captionText="FIGURE 1AD. Habitus of males in dorsal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478111" httpUri="https://zenodo.org/record/4478111/files/figure.png" pageId="6" pageNumber="386">Figs. 1</figureCitation>
CD, 2CD, 3CD, 4D, 5C, 7AC, 8AB, 9)
</paragraph>
</subSubSection>
<subSubSection id="C3206546FF9DAD24D8DA8189FC67FDEE" box="[151,961,1227,1251]" pageId="6" pageNumber="386" type="reference_group">
<paragraph id="8B8536CDFF9DAD24D8DA8189FC67FDEE" blockId="6.[151,961,1227,1251]" box="[151,961,1227,1251]" pageId="6" pageNumber="386">
<treatmentCitationGroup id="AB2A11E3FF9DAD24D8DA8189FC67FDEE" box="[151,961,1227,1251]" pageId="6" pageNumber="386">
<treatmentCitation id="0A9B10DCFF9DAD24D8DA8189FE59FDEE" author="Reitter, E." box="[151,511,1227,1251]" page="81" pageId="6" pageNumber="386" year="1895">
<taxonomicName id="4C3A4D4EFF9DAD24D8DA8189FE59FDEE" authority="Reitter, 1895: 81" authorityName="Reitter" authorityPageNumber="81" authorityYear="1895" box="[151,511,1227,1251]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="nireus">
<emphasis id="B94EEADFFF9DAD24D8DA8189FEE9FDEE" box="[151,335,1227,1251]" italics="true" pageId="6" pageNumber="386">Bolboceras nireus</emphasis>
<bibRefCitation id="EFAB4B3CFF9DAD24D91B8189FE59FDEE" author="Reitter, E." box="[342,511,1227,1251]" pageId="6" pageNumber="386" pagination="79 - 88" refId="ref8665" refString="Reitter, E. (1895) Beschreibung neuer oder wenig gekannter Coleoptera aus der Umgebung von Akbes in Syrien. Wiener Entomologische Zeitung, 14, 79 - 88. https: // doi. org / 10.5962 / bhl. part. 13210" type="journal article" year="1895">Reitter, 1895: 81</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="5481886FFF9DAD24DA408189FD9CFDEE" box="[525,570,1227,1251]" pageId="6" pageNumber="386">type</typeStatus>
locality: “Akbes [= Maydân Akbis]”).
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</paragraph>
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<subSubSection id="C3206546FF9DAD24D8DA804EFD1AFEFB" pageId="6" pageNumber="386" type="materials_examined">
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<emphasis id="B94EEADFFF9DAD24D8DA804EFE67FC2B" bold="true" box="[151,449,1292,1318]" pageId="6" pageNumber="386">Type material examined.</emphasis>
<materialsCitation id="3B523C90FF9DAD24D984804EFAE6FC62" ID-GBIF-Occurrence="3033773309" collectionCode="MNHN" country="Syria" determinerName="J. Krikken &amp; D. Kral &amp; D. Sommer" location="Syria" pageId="6" pageNumber="386" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B94EEADFFF9DAD24D984804EFDB4FC2B" bold="true" box="[457,530,1292,1318]" pageId="6" pageNumber="386">
<collectingCountry id="F32D765DFF9DAD24D984804EFDABFC2B" box="[457,525,1292,1318]" name="Syria" pageId="6" pageNumber="386">Syria</collectingCountry>
:
</emphasis>
<typeStatus id="5481886FFF9DAD24DA56804EFD20FC2B" box="[539,646,1292,1318]" pageId="6" pageNumber="386" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D3CFD44FF9DAD24DADF804FFD01FC2A" box="[658,679,1293,1319]" pageId="6" pageNumber="386" type="female"></specimenCount>
(
<collectionCode id="ED2BAE08FF9DAD24DAFA804EFCB6FC2B" box="[695,784,1292,1318]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="386">MNHN</collectionCode>
) (
<figureCitation id="13012A48FF9DAD24DB65804EFCD3FC2B" box="[808,885,1292,1318]" captionStart="FIGURE 7" captionStartId="8.[151,250,1944,1968]" captionTargetBox="[191,1411,194,1906]" captionTargetId="figure-17@8.[167,1420,181,1920]" captionTargetPageId="8" captionText="FIGURE 7AC. Holotype of Bolbelasmus nireus (Reitter, 1895). A, habitus in dorsal view; B, original labels of holotype; C, habitus in lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478123" httpUri="https://zenodo.org/record/4478123/files/figure.png" pageId="6" pageNumber="386">Figs. 7</figureCitation>
AC, 8AB) “
<taxonomicName id="4C3A4D4EFF9DAD24DC55804EFB3FFC2B" box="[1048,1177,1292,1318]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="genus">Bolboceras</taxonomicName>
| Nireus m | n. sp. Akbes [hw, Reitters hand] || Typus [p, red label] || jkF720329 || VIO ||
<taxonomicName id="4C3A4D4EFF9DAD24DBE08072FB98FC47" authorityName="Boucomont" authorityYear="1911" box="[941,1086,1328,1354]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="genus">Bolbelasmus</taxonomicName>
|
<taxonomicName id="4C3A4D4EFF9DAD24DC1B8072FB73FC46" authority="Rttr." authorityName="Reitter" authorityYear="1895" box="[1110,1237,1328,1355]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="nireus">nireus Rttr.</taxonomicName>
|| det.
<determinerName id="7E59DA09FF9DAD24DD688073FA3AFC47" box="[1317,1436,1328,1355]" pageId="6" pageNumber="386">J. Krikken</determinerName>
197[p]2[hw, Krikkens hand] ||
<taxonomicName id="4C3A4D4EFF9DAD24D9BB8016FD74FC63" baseAuthorityName="Reitter" baseAuthorityYear="1895" box="[502,722,1364,1390]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="386" phylum="Arthropoda" rank="species" species="nireus">Bolbelasmus nireus</taxonomicName>
| (
<bibRefCitation id="EFAB4B3CFF9DAD24DAA08016FC27FC62" author="Reitter, E." box="[749,897,1364,1391]" pageId="6" pageNumber="386" pagination="79 - 88" refId="ref8665" refString="Reitter, E. (1895) Beschreibung neuer oder wenig gekannter Coleoptera aus der Umgebung von Akbes in Syrien. Wiener Entomologische Zeitung, 14, 79 - 88. https: // doi. org / 10.5962 / bhl. part. 13210" type="journal article" year="1895">Reitter, 1895</bibRefCitation>
) |
<determinerName id="7E59DA09FF9DAD24DBD08017FC57FC63" box="[925,1009,1364,1391]" pageId="6" pageNumber="386">D. Král</determinerName>
&amp;
<determinerName id="7E59DA09FF9DAD24DC588017FB30FC63" box="[1045,1174,1364,1391]" pageId="6" pageNumber="386">D. Sommer</determinerName>
det. 2019 [p]”.
</materialsCitation>
</paragraph>
<paragraph id="8B8536CDFF9DAD24D88A803AFE60FF66" blockId="6.[151,1437,1292,2039]" pageId="6" pageNumber="386">
<emphasis id="B94EEADFFF9DAD24D88A803AFD95FC9F" bold="true" box="[199,563,1400,1426]" pageId="6" pageNumber="386">Additional material examined.</emphasis>
<materialsCitation id="3B523C90FF9DAD24DA77803AFC85FCBB" ID-GBIF-Occurrence="3033773306" collectingDate="2017-03-28" collectingDateMax="2017-04-16" collectingDateMin="2017-03-28" collectionCode="LSCN, OHCB" collectorName="L. Schmidt" country="Greece" latitude="36.41697" location="Lalyssos" longLatPrecision="1" longitude="28.145563" pageId="6" pageNumber="386" specimenCount="1">
<emphasis id="B94EEADFFF9DAD24DA77803AFD5BFC9F" bold="true" box="[570,765,1400,1426]" pageId="6" pageNumber="386">
<collectingCountry id="F32D765DFF9DAD24DA77803AFD29FC9F" box="[570,655,1400,1426]" name="Greece" pageId="6" pageNumber="386">Greece</collectingCountry>
,
<collectorName id="26CF531BFF9DAD24DAD6803AFD5EFC9F" box="[667,760,1400,1426]" pageId="6" pageNumber="386">Rhodes</collectorName>
:
</emphasis>
1 ♁ (
<collectionCode id="ED2BAE08FF9DAD24DB76803AFC2EFC9F" box="[827,904,1400,1426]" pageId="6" pageNumber="386">LSCN</collectionCode>
), 1 ♁ (
<collectionCode id="ED2BAE08FF9DAD24DB9E803AFB82FC9F" box="[979,1060,1400,1426]" pageId="6" pageNumber="386">OHCB</collectionCode>
) (
<figureCitation id="13012A48FF9DAD24DC76803AFB3AFC9F" box="[1083,1180,1400,1426]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[166,1419,193,1903]" captionTargetId="figure-17@2.[153,1435,179,1918]" captionTargetPageId="2" captionText="FIGURE 1AD. Habitus of males in dorsal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478111" httpUri="https://zenodo.org/record/4478111/files/figure.png" pageId="6" pageNumber="386">Figs. 1C</figureCitation>
,
<figureCitation id="13012A48FF9DAD24DCE5803AFB6DFC9F" box="[1192,1227,1400,1426]" captionStart="FIGURE 2" captionStartId="3.[151,250,1151,1175]" captionTargetBox="[157,1426,190,1120]" captionTargetId="figure-20@3.[151,1436,181,1127]" captionTargetPageId="3" captionText="FIGURE 2AD. Pronotum of males in frontal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478113" httpUri="https://zenodo.org/record/4478113/files/figure.png" pageId="6" pageNumber="386">2C</figureCitation>
,
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, [without date], [E.]
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(
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),
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,
<date id="FF84100DFF9DAD24D8DA80A6FEA9FCF3" box="[151,271,1508,1534]" pageId="6" pageNumber="386" value="1973-04-27">
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</date>
, K.
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lgt
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.;
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<collectingDate id="EFC0E9E5FF9DAD24DAA280A6FCCEFCF3" box="[751,872,1508,1534]" pageId="6" pageNumber="386" value="1991-06-11">11.vi.1991</collectingDate>
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,
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lgt.; 1 ♁ (
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,
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, Sebil,
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, J.
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lgt
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.;
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(
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S of
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<collectingDate id="EFC0E9E5FF9DAD24DB33834AFC6BFF2F" box="[894,973,1544,1570]" pageId="6" pageNumber="386" value="2001-05">v.2001</collectingDate>
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, [without collectors name]; 1 ♁ (
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<figureCitation id="13012A48FF9DAD24D95F836EFE91FF4B" box="[274,311,1580,1606]" captionStart="FIGURE 2" captionStartId="3.[151,250,1151,1175]" captionTargetBox="[157,1426,190,1120]" captionTargetId="figure-20@3.[151,1436,181,1127]" captionTargetPageId="3" captionText="FIGURE 2AD. Pronotum of males in frontal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4478113" httpUri="https://zenodo.org/record/4478113/files/figure.png" pageId="6" pageNumber="386">2D</figureCitation>
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NW of
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NE of
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<date id="FF84100DFF9DAD24DB4F836EFCCBFF4A" box="[770,877,1580,1607]" pageId="6" pageNumber="386" value="2015-04-08">
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, M.
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lgt.; 2 ♁♁ (
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[without date and collectors name]
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.
</paragraph>
<paragraph id="8B8536CDFF9DAD24D88A8336FD5EFFDA" blockId="6.[151,1437,1292,2039]" pageId="6" pageNumber="386">
<emphasis id="B94EEADFFF9DAD24D88A8336FEF1FF83" bold="true" box="[199,343,1652,1678]" pageId="6" pageNumber="386">Disribution.</emphasis>
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,
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and
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(
<bibRefCitation id="EFAB4B3CFF9DAD24DA2A8336FCFFFF82" author="Nikolajev, G. V. &amp; Kral, D. &amp; Bezdek, A." box="[615,857,1652,1679]" pageId="6" pageNumber="386" pagination="1802" refId="ref8582" refString="Nikolajev, G. V., Kral, D. &amp; Bezdek, A. (2016) Family Geotrupidae Latreille, 1802. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea-Scirtoidea-Dasciloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 33 - 52. https: // doi. org / 10.1163 / 9789004309142 _ 003" type="book chapter" year="2016">
Nikolajev
<emphasis id="B94EEADFFF9DAD24DA938337FCB0FF83" box="[734,790,1652,1678]" italics="true" pageId="6" pageNumber="386">et al.</emphasis>
2016
</bibRefCitation>
). New country record for
<collectingCountry id="F32D765DFF9DAD24DC358336FB61FF83" box="[1144,1223,1652,1678]" name="Greece" pageId="6" pageNumber="386">Greece</collectingCountry>
(island of Rhodes) (see map,
<figureCitation id="13012A48FF9DAD24D94983DAFEE3FFBF" box="[260,325,1688,1714]" captionStart="FIGURE 9" captionStartId="10.[151,250,940,964]" captionTargetBox="[170,1418,187,912]" captionTargetId="figure-17@10.[164,1424,181,916]" captionTargetPageId="10" captionText="FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species (red). Squares represents the type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.4478127" httpUri="https://zenodo.org/record/4478127/files/figure.png" pageId="6" pageNumber="386">Fig. 9</figureCitation>
). The record of the species
<emphasis id="B94EEADFFF9DAD24DA2683DBFDDAFFBF" box="[619,636,1689,1714]" italics="true" pageId="6" pageNumber="386">B</emphasis>
.
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(
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.)
<emphasis id="B94EEADFFF9DAD24DAFB83DBFC86FFBF" box="[694,800,1689,1714]" italics="true" pageId="6" pageNumber="386">unicornis</emphasis>
(Schrank, 1789)
</taxonomicName>
from the island of Rhodes by
<bibRefCitation id="EFAB4B3CFF9DAD24DD5583DAFF44FFDA" author="Schatzmayr, A." pageId="6" pageNumber="386" pagination="9 - 23" refId="ref8710" refString="Schatzmayr, A. (1936) Risultati scientifici delle cacce entomologiche di S. A. S. il Princiope Alessandro della Torre et Tasso nelle isole dell'Egeo. VI. - Scarabaeidae (1). Bollettino del Regio Laboratorio di Entomologia Agraria di Portici, 30, 9 - 23." type="journal article" year="1936">Schatzmayr (1936)</bibRefCitation>
most likely relates to the species
<taxonomicName id="4C3A4D4EFF9DAD24DA1683FFFD57FFDB" baseAuthorityName="Reitter" baseAuthorityYear="1895" box="[603,753,1724,1751]" class="Insecta" family="Bolboceratidae" genus="Bolbelasmus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="381" phylum="Arthropoda" rank="species" species="nireus" subGenus="Bolbelasmus">
<emphasis id="B94EEADFFF9DAD24DA1683FFFDCAFFDB" box="[603,620,1725,1750]" italics="true" pageId="6" pageNumber="386">B</emphasis>
. (
<emphasis id="B94EEADFFF9DAD24DACE83FFFD32FFDB" box="[643,660,1725,1750]" italics="true" pageId="6" pageNumber="386">B</emphasis>
.)
<emphasis id="B94EEADFFF9DAD24DAE183FFFD57FFDB" box="[684,753,1725,1750]" italics="true" pageId="6" pageNumber="386">nireus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8536CDFF9DAD24D88A83A2FD1AFEFB" blockId="6.[151,1437,1292,2039]" pageId="6" pageNumber="386">
<emphasis id="B94EEADFFF9DAD24D88A83A2FEA0FFF7" bold="true" box="[199,262,1760,1786]" pageId="6" pageNumber="386">Note.</emphasis>
The distribution in
<collectingCountry id="F32D765DFF9DAD24D9BD83A3FDBBFFF6" box="[496,541,1761,1787]" name="Iraq" pageId="6" pageNumber="386">Iraq</collectingCountry>
is based on a single female specimen from Assur tentatively assigned to this species by
<bibRefCitation id="EFAB4B3CFF9DAD24D9588246FE60FE13" author="Krikken, J." box="[277,454,1796,1822]" pageId="6" pageNumber="386" pagination="277 - 292" refId="ref8248" refString="Krikken, J. (1977) The genus Bolbelasmus Boucomont in Asia with notes on species occurring in other regions. Zoologische Mededeelingen, 51, 277 - 292." type="journal article" year="1977">Krikken (1977)</bibRefCitation>
. We have not studied this specimen, so we cannot confirm with certainty what species it belongs to. In addition, this Iraqi locality of the species is very far from its distribution in
<collectingCountry id="F32D765DFF9DAD24DCE8826AFB53FE4F" box="[1189,1269,1832,1858]" name="Turkey" pageId="6" pageNumber="386">Turkey</collectingCountry>
(see map,
<figureCitation id="13012A48FF9DAD24DD3D826AFF01FE6B" captionStart="FIGURE 9" captionStartId="10.[151,250,940,964]" captionTargetBox="[170,1418,187,912]" captionTargetId="figure-17@10.[164,1424,181,916]" captionTargetPageId="10" captionText="FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species (red). Squares represents the type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.4478127" httpUri="https://zenodo.org/record/4478127/files/figure.png" pageId="6" pageNumber="386">Fig. 9</figureCitation>
). Their distribution in
<collectingCountry id="F32D765DFF9DAD24D9EB820EFE47FE6B" box="[422,481,1868,1894]" name="Syria" pageId="6" pageNumber="386">Syria</collectingCountry>
is based on two females (including the
<typeStatus id="5481886FFF9DAD24DBEC820EFBA3FE6A" box="[929,1029,1868,1895]" pageId="6" pageNumber="386" type="holotype">holotype</typeStatus>
) collected at “Akbes”. This locality, nowadays a small village Maydân Akbis belongs to the territory of
<collectingCountry id="F32D765DFF9DAD24DBC98232FC19FE87" box="[900,959,1904,1930]" name="Syria" pageId="6" pageNumber="386">Syria</collectingCountry>
(
<collectingRegion id="49FEF82FFF9DAD24DB838232FB10FE87" box="[974,1206,1904,1930]" country="Syria" name="Halab" pageId="6" pageNumber="386">Aleppo Governorate</collectingRegion>
) and is situated very close to the border with
<collectingCountry id="F32D765DFF9DAD24D9E482D6FE5FFEA3" box="[425,505,1940,1966]" name="Turkey" pageId="6" pageNumber="386">Turkey</collectingCountry>
(see map,
<figureCitation id="13012A48FF9DAD24DA3982D6FD1EFEA3" box="[628,696,1940,1966]" captionStart="FIGURE 9" captionStartId="10.[151,250,940,964]" captionTargetBox="[170,1418,187,912]" captionTargetId="figure-17@10.[164,1424,181,916]" captionTargetPageId="10" captionText="FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová &amp; Král, new species (red). Squares represents the type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.4478127" httpUri="https://zenodo.org/record/4478127/files/figure.png" pageId="6" pageNumber="386">Fig. 9</figureCitation>
). Between 18811926 there was the Trappist monastery “Abbaye dAkbès” (
<bibRefCitation id="EFAB4B3CFF9DAD24D95E82FAFE51FEDF" author="Le Bras, G. &amp; Hourlier, J. &amp; Cocheril, M." box="[275,503,1976,2003]" pageId="6" pageNumber="386" refId="ref8315" refString="Le Bras, G., Hourlier, J. &amp; Cocheril, M. (1979) Les Ordres religieux, la Vie et l'Art. Vol. 1. Monasteres et communautes; les Benedictins; les Cisterciens; les Chartreux; les ordres militaires. Flammarion, Paris, 266 pp." type="book" year="1979">
Le Bras
<emphasis id="B94EEADFFF9DAD24D93582FBFE14FEDF" box="[376,434,1976,2002]" italics="true" pageId="6" pageNumber="386">et al.</emphasis>
1979
</bibRefCitation>
). The site is not in modern
<collectingCountry id="F32D765DFF9DAD24DB7A82FAFC2EFEDF" box="[823,904,1976,2002]" name="Turkey" pageId="6" pageNumber="386">Turkey</collectingCountry>
, as stated in some papers, see
<bibRefCitation id="EFAB4B3CFF9DAD24DCA482FAFF42FEFB" author="Krystufek, B. &amp; Spitzenberger, F. &amp; Kefelioglu, H." pageId="6" pageNumber="386" pagination="135 - 143" refId="ref8279" refString="Krystufek, B., Spitzenberger, F. &amp; Kefelioglu, H. (2001) Description, taxonomy, and distribution of Talpa davidiana. Mammalian Biology, 66, 135 - 143." type="journal article" year="2001">
Kryštufek
<emphasis id="B94EEADFFF9DAD24DD2F82FBFA33FEDF" box="[1378,1429,1976,2002]" italics="true" pageId="6" pageNumber="386">et al</emphasis>
. (2001)
</bibRefCitation>
,
<bibRefCitation id="EFAB4B3CFF9DAD24D8BD829EFE26FEFA" author="Keith, D." box="[240,384,2012,2039]" pageId="6" pageNumber="386" pagination="51 - 52" refId="ref8224" refString="Keith, D. (2005) First record of the genus Bolbelasmus from Iran. Kogane, 6, 51 - 52." type="journal article" year="2005">Keith (2005)</bibRefCitation>
or
<bibRefCitation id="EFAB4B3CFF9DAD24D9EE829EFD1EFEFB" author="Arnone, M. &amp; Massa, B." box="[419,696,2012,2039]" pageId="6" pageNumber="386" pagination="401 - 414" refId="ref7961" refString="Arnone, M. &amp; Massa, B. (2010) A new species of Bolbelasmus Boucomont, 1911 from Sicily (Italy). Il Naturalista Siciliano, 34, 401 - 414." type="journal article" year="2010">Arnone &amp; Massa (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>