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<document id="B7C069C6D65CD977CAB206F27B6B91BD" ID-CLB-Dataset="299719" ID-DOI="10.11646/zootaxa.5474.5.8" ID-GBIF-Dataset="09a20124-10af-4376-a2f8-cedb504e1b52" ID-ISSN="1175-5326" ID-Zenodo-Dep="12731004" ID-ZooBank="170DE3E1-360B-4B15-A29B-C1813F04E7CC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720374090561" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="03FD87C7EC107A13FF6CFD1875B5FAEA" docLanguage="en" docName="zootaxa.5474.5.8.pdf" docOrigin="Zootaxa 5474 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia cachemirica Coiffait 1982" docType="treatment" docVersion="4" lastPageNumber="587" masterDocId="FFC4FFBFEC117A10FFFBFFE27115FFE7" masterDocTitle="On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="592" masterPageNumber="584" pageNumber="585" updateTime="1720775253469" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="84319736B98080B557E2A11441191C12">On some Himalayan species of the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<treatment id="03FD87C7EC107A13FF6CFD1875B5FAEA" ID-DOI="http://doi.org/10.5281/zenodo.12731006" ID-GBIF-Taxon="235046007" ID-Zenodo-Dep="12731006" LSID="urn:lsid:plazi:treatment:03FD87C7EC107A13FF6CFD1875B5FAEA" httpUri="http://treatment.plazi.org/id/03FD87C7EC107A13FF6CFD1875B5FAEA" lastPageId="3" lastPageNumber="587" pageId="1" pageNumber="585">
<subSubSection id="C34E655AEC107A11FF6CFD18738CFCF3" box="[151,665,762,789]" pageId="1" pageNumber="585" type="nomenclature">
<paragraph id="8BEB36D1EC107A11FF6CFD18738CFCF3" blockId="1.[151,665,762,825]" box="[151,665,762,789]" pageId="1" pageNumber="585">
<heading id="D0A381BDEC107A11FF6CFD18738CFCF3" bold="true" box="[151,665,762,789]" fontSize="11" level="1" pageId="1" pageNumber="585" reason="1">
<taxonomicName id="4C544D52EC107A11FF6CFD18738CFCF3" ID-CoL="6R6J9" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[151,665,762,789]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FF6CFD18738CFCF3" bold="true" box="[151,665,762,789]" pageId="1" pageNumber="585">
<emphasis id="B920EAC3EC107A11FF6CFD1870F2FCF2" bold="true" box="[151,487,762,789]" italics="true" pageId="1" pageNumber="585">Mannerheimia cachemirica</emphasis>
<bibRefCitation id="EFC54B20EC107A11FE14FD18738CFCF3" author="Coiffait, H." box="[495,665,762,788]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC107A11FF6CFCFD7056FCDE" box="[151,323,799,825]" pageId="1" pageNumber="585" type="description">
<paragraph id="8BEB36D1EC107A11FF6CFCFD7056FCDE" blockId="1.[151,665,762,825]" box="[151,323,799,825]" pageId="1" pageNumber="585">
(
<figureCitation id="136F2A54EC107A11FF64FCFD7011FCDE" box="[159,260,799,825]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Figs 1–2</figureCitation>
,
<figureCitation id="136F2A54EC107A11FEEBFCFD702EFCDE" box="[272,315,799,825]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="1" pageNumber="585">6–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC107A11FF6CFC81732CFC5E" pageId="1" pageNumber="585" type="reference_group">
<paragraph id="8BEB36D1EC107A11FF6CFC81722AFC9C" blockId="1.[151,831,867,953]" box="[151,831,867,891]" pageId="1" pageNumber="585">
<treatmentCitationGroup id="AB4411FFEC107A11FF6CFC81722AFC9C" box="[151,831,867,891]" pageId="1" pageNumber="585">
<treatmentCitation id="0AF510C0EC107A11FF6CFC81736FFC9C" author="Coiffait, H." box="[151,634,867,891]" page="146" pageId="1" pageNumber="585" year="1982">
<taxonomicName id="4C544D52EC107A11FF6CFC81736FFC9C" ID-CoL="6R6J9" authority="Coiffait, 1982: 146" authorityName="Coiffait" authorityPageNumber="146" authorityYear="1982" box="[151,634,867,891]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FF6CFC8170BAFC9C" box="[151,431,867,891]" italics="true" pageId="1" pageNumber="585">Mannerheimia cachemirica</emphasis>
<bibRefCitation id="EFC54B20EC107A11FE4DFC81736FFC9C" author="Coiffait, H." box="[438,634,867,891]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982: 146</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF510C0EC107A11FD7FFC81722AFC9C" author="Shavrin, A. V." box="[644,831,867,891]" page="307" pageId="1" pageNumber="585" year="2021">
<bibRefCitation id="EFC54B20EC107A11FD7FFC81722AFC9C" author="Shavrin, A. V." box="[644,831,867,891]" pageId="1" pageNumber="585" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021: 307</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEB36D1EC107A11FF6CFC607396FC7D" blockId="1.[151,831,867,953]" box="[151,643,898,922]" pageId="1" pageNumber="585">
<treatmentCitationGroup id="AB4411FFEC107A11FF6CFC607396FC7D" box="[151,643,898,922]" pageId="1" pageNumber="585">
<treatmentCitation id="0AF510C0EC107A11FF6CFC60732DFC7D" author="Coiffait, H." box="[151,568,898,922]" page="146" pageId="1" pageNumber="585" year="1982">
<taxonomicName id="4C544D52EC107A11FF6CFC60732DFC7D" ID-CoL="3XVMT" authority="Coiffait, 1982: 146" authorityName="Coiffait" authorityPageNumber="146" authorityYear="1982" box="[151,568,898,922]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="vicina" status="syn. nov.">
<emphasis id="B920EAC3EC107A11FF6CFC607065FC7E" box="[151,368,898,922]" italics="true" pageId="1" pageNumber="585">Mannerheimia vicina</emphasis>
<bibRefCitation id="EFC54B20EC107A11FE8DFC60732DFC7D" author="Coiffait, H." box="[374,568,898,922]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982: 146</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="B920EAC3EC107A11FDC5FC607396FC7D" bold="true" box="[574,643,898,922]" pageId="1" pageNumber="585">
<taxonomicNameLabel id="A21357B8EC107A11FDC5FC607396FC7D" box="[574,643,898,922]" pageId="1" pageNumber="585">syn. n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEB36D1EC107A11FF6CFC42732CFC5E" blockId="1.[151,831,867,953]" box="[151,569,928,953]" pageId="1" pageNumber="585">
<treatmentCitationGroup id="AB4411FFEC107A11FF6CFC42732CFC5E" box="[151,569,928,953]" pageId="1" pageNumber="585">
<taxonomicName id="4C544D52EC107A11FF6CFC427065FC5F" ID-CoL="3XVMT" authorityName="Coiffait" authorityYear="1982" box="[151,368,928,952]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC107A11FF6CFC427065FC5F" box="[151,368,928,952]" italics="true" pageId="1" pageNumber="585">Mannerheimia vicina</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AF510C0EC107A11FE86FC42732CFC5E" author="Shavrin, A. V." box="[381,569,928,953]" page="307" pageId="1" pageNumber="585" year="2021">
<bibRefCitation id="EFC54B20EC107A11FE86FC42732CFC5E" author="Shavrin, A. V." box="[381,569,928,953]" pageId="1" pageNumber="585" pagination="301 - 333" refId="ref4932" refString="Shavrin, A. V. (2021) On the genus Mannerheimia Maklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species. Zootaxa, 5040 (3), 301 - 333. https: // doi. org / 10.11646 / zootaxa. 5040.3.1" type="journal article" year="2021">Shavrin 2021: 307</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC107A11FF6CFC05707AFAC6" pageId="1" pageNumber="585" type="materials_examined">
<paragraph id="8BEB36D1EC107A11FF6CFC0573FDFB76" blockId="1.[151,1437,998,2033]" pageId="1" pageNumber="585">
<emphasis id="B920EAC3EC107A11FF6CFC0570D5FBE7" bold="true" box="[151,448,998,1025]" pageId="1" pageNumber="585">Type material examined.</emphasis>
<typeStatus id="54EF8873EC107A11FE3CFC05733AFBE6" box="[455,559,999,1025]" pageId="1" pageNumber="585" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FDAEFC057525FBE6" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[597,1072,999,1025]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FDAEFC05729EFBE6" box="[597,907,999,1025]" italics="true" pageId="1" pageNumber="585">Mannerheimia cachemirica</emphasis>
<bibRefCitation id="EFC54B20EC107A11FC68FC057525FBE6" author="Coiffait, H." box="[915,1072,999,1025]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</taxonomicName>
<specimenCount id="9D52FD58EC107A11FBCCFC057559FBE6" box="[1079,1100,999,1025]" count="1" pageId="1" pageNumber="585" type="male">♂</specimenCount>
(
<figureCitation id="136F2A54EC107A11FBA7FC05758AFBE6" box="[1116,1183,999,1025]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Fig. 1</figureCitation>
; dissected): ‘Kashmir, Aru | lg.H.Franz,
<date id="FFEA1011EC107A11FEA4FBE970C6FBC2" box="[351,467,1035,1061]" pageId="1" pageNumber="585" value="1977-10">Okt. 1977</date>
’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘
<taxonomicName id="4C544D52EC107A11FC8AFBE9750FFBC2" authorityName="Maklin" authorityYear="1880" box="[881,1050,1035,1061]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
| cachemirica | H. COIFFAIT det. 19[printed]76’ &lt;handwritten in black&gt; (
<collectionCode id="ED45AE14EC107A11FDA8FBCD73B4FBAE" box="[595,673,1071,1097]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="585">NMW</collectionCode>
).
<typeStatus id="54EF8873EC107A11FD48FBCD7200FBAE" box="[691,789,1071,1097]" pageId="1" pageNumber="585" type="paratype">Paratype</typeStatus>
<specimenCount id="9D52FD58EC107A11FCE7FBCD7224FBAE" box="[796,817,1071,1097]" count="1" pageId="1" pageNumber="585" type="male">♂</specimenCount>
(dissected; aedeagus missing): first labels as that in the
<typeStatus id="54EF8873EC107A11FF6CFBB171E9FB8A" box="[151,252,1107,1133]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
, with red printed label: ‘PARATYPE’ (
<collectionCode id="ED45AE14EC107A11FD47FBB1721FFB8A" box="[700,778,1107,1133]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="585">NMW</collectionCode>
). All types with additional printed label: ‘
<taxonomicName id="4C544D52EC107A11FB12FBB17498FB8A" authorityName="Maklin" authorityYear="1880" box="[1257,1421,1107,1133]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC3EC107A11FB12FBB17498FB8A" box="[1257,1421,1107,1133]" italics="true" pageId="1" pageNumber="585">Mannerheimia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C544D52EC107A11FF6CFB9570D0FB76" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[151,453,1143,1169]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FF6CFB957034FB76" box="[151,289,1143,1169]" italics="true" pageId="1" pageNumber="585">cachemirica</emphasis>
<bibRefCitation id="EFC54B20EC107A11FED2FB9570D0FB76" author="Coiffait, H." box="[297,453,1143,1169]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</taxonomicName>
| Shavrin A.
<collectionCode id="ED45AE14EC107A11FDB4FB957370FB76" box="[591,613,1143,1169]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="585" type="Museum">V</collectionCode>
. det. 2024’.
</paragraph>
<paragraph id="8BEB36D1EC107A11FF3CFB79707AFAC6" blockId="1.[151,1437,998,2033]" pageId="1" pageNumber="585">
<typeStatus id="54EF8873EC107A11FF3CFB79703AFB52" box="[199,303,1179,1205]" pageId="1" pageNumber="585" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FEACFB7973FBFB52" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[343,750,1179,1205]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC107A11FEACFB79735DFB53" box="[343,584,1179,1205]" italics="true" pageId="1" pageNumber="585">Mannerheimia vicina</emphasis>
<bibRefCitation id="EFC54B20EC107A11FDABFB7973FBFB52" author="Coiffait, H." box="[592,750,1179,1205]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</taxonomicName>
<specimenCount id="9D52FD58EC107A11FD0DFB79721EFB52" box="[758,779,1179,1205]" count="1" pageId="1" pageNumber="585" type="male">♂</specimenCount>
(
<figureCitation id="136F2A54EC107A11FCE0FB797277FB52" box="[795,866,1179,1205]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Fig. 2</figureCitation>
; dissected): ‘Kashmir, Chandanwari | b.Pahalgam,
<quantity id="4CAC9B34EC107A11FF6CFB5D7025FB3E" box="[151,304,1215,1241]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="1" pageNumber="585" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800-3100 m</quantity>
| lg.H.Franz,
<date id="FFEA1011EC107A11FE3CFB5D7321FB3E" box="[455,564,1215,1241]" pageId="1" pageNumber="585" value="1977-10">Okt.1977</date>
’ &lt;printed&gt;, ‘TYPE’ &lt;red, printed&gt;, ‘
<taxonomicName id="4C544D52EC107A11FC27FB5D7590FB3E" authorityName="Maklin" authorityYear="1880" box="[988,1157,1215,1241]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
| vicina | H. COIFFAIT det. 19[printed]76’ &lt;handwritten in black&gt;, ‘
<taxonomicName id="4C544D52EC107A11FD69FB017223FB1A" authorityName="Maklin" authorityYear="1880" box="[658,822,1251,1277]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC3EC107A11FD69FB017223FB1A" box="[658,822,1251,1277]" italics="true" pageId="1" pageNumber="585">Mannerheimia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C544D52EC107A11FCB6FB017568FB1A" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[845,1149,1251,1277]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FCB6FB0172C2FB1A" box="[845,983,1251,1277]" italics="true" pageId="1" pageNumber="585">cachemirica</emphasis>
<bibRefCitation id="EFC54B20EC107A11FC24FB017568FB1A" author="Coiffait, H." box="[991,1149,1251,1277]" pageId="1" pageNumber="585" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</taxonomicName>
| Shavrin A.
<collectionCode id="ED45AE14EC107A11FAF1FB017435FB1A" box="[1290,1312,1251,1277]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="585" type="Museum">V</collectionCode>
. det. 2024’ &lt;printed&gt; (
<collectionCode id="ED45AE14EC107A11FEEEFAE57076FAC6" box="[277,355,1287,1313]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="585">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC107A13FF3CFAC87050FC66" lastPageId="3" lastPageNumber="587" pageId="1" pageNumber="585" type="description">
<paragraph id="8BEB36D1EC107A11FF3CFAC871F9FA56" blockId="1.[151,1437,998,2033]" pageId="1" pageNumber="585">
<emphasis id="B920EAC3EC107A11FF3CFAC87060FAA3" bold="true" box="[199,373,1322,1348]" pageId="1" pageNumber="585">Redescription.</emphasis>
Measurements (n=3): HW: 0.47–0.55; HL: 0.32–0.42; OL: 0.12; AL (
<typeStatus id="54EF8873EC107A11FB87FAC975F0FAA2" box="[1148,1253,1323,1349]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
): 0.92; PL: 0.37; PW: 0.62–0.72; ESL: 0.75–0.90; EW: 0.84–0.96; MTbL (
<typeStatus id="54EF8873EC107A11FCE1FAAD7296FA8E" box="[794,899,1359,1385]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
): 0.54; MTrL (
<typeStatus id="54EF8873EC107A11FBDDFAAD759AFA8E" box="[1062,1167,1359,1385]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
): 0.30 (MTrL 1–4: 0.18; MTrL 5: 0.12); AW: 0.85–1.03; AedL: 0.35–0.39; BL: 2.55 (
<typeStatus id="54EF8873EC107A11FCAFFA9172ADFA6A" box="[852,952,1395,1421]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FC18FA917580FA6A" authorityName="Coiffait" authorityYear="1982" box="[995,1173,1395,1421]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FC18FA917580FA6A" box="[995,1173,1395,1421]" italics="true" pageId="1" pageNumber="585">M. cachemirica</emphasis>
</taxonomicName>
)–2.65 (
<typeStatus id="54EF8873EC107A11FB14FA917446FA6A" box="[1263,1363,1395,1421]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FA85FA9171CEFA57" authorityName="Coiffait" authorityYear="1982" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC107A11FA85FA9171CEFA57" italics="true" pageId="1" pageNumber="585">M. vicina</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BEB36D1EC107A11FF3CFA597292F886" blockId="1.[151,1437,998,2033]" pageId="1" pageNumber="585">
Habitus as in
<figureCitation id="136F2A54EC107A11FE9BFA5970D1FA32" box="[352,452,1467,1493]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1758,1782]" captionTargetBox="[157,1429,185,1726]" captionTargetId="figure-16@2.[151,1435,179,1732]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus of Mannerheimia (holotypes): 1–M. cachemirica, 2–M. vicina, 3–M. emodensa, 4–M. obscura, 5–M. scutellata, Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="1" pageNumber="585">Figs 1–2</figureCitation>
. Body reddish-brown, with distinctly darker head and abdomen (pronotum and elytra of
<typeStatus id="54EF8873EC107A11FF6CFA3D71ECFA1E" box="[151,249,1503,1529]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FEE7FA3D70DFFA1E" authorityName="Coiffait" authorityYear="1982" box="[284,458,1503,1529]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FEE7FA3D70DFFA1E" box="[284,458,1503,1529]" italics="true" pageId="1" pageNumber="585">M. cachemirica</emphasis>
</taxonomicName>
distinctly paler; paratergites of abdomen paler); antennomeres 6–11 brown; mouthparts, antennomeres 1–5, legs and paratergites of abdomen yellow. Body shiny, without microsculpture except for middle portion of head (
<typeStatus id="54EF8873EC107A11FEA7F9C570D5F9A6" box="[348,448,1575,1601]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FE16F9C573B5F9A6" authorityName="Coiffait" authorityYear="1982" box="[493,672,1575,1601]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FE16F9C573B5F9A6" box="[493,672,1575,1601]" italics="true" pageId="1" pageNumber="585">M. cachemirica</emphasis>
</taxonomicName>
) with very indistinct longitudinal meshes and abdomen with fine and dense transverse microreticulation (other specimens without meshes on head), distinctly coarser in abdominal tergite III–IV. Head with irregular and sparse, moderately large and deep punctation, denser in middle; middle portion of neck with very sparse and fine punctation; punctation of pronotum denser, larger and deeper than that on head, finer and sparser in middle (
<typeStatus id="54EF8873EC107A11FDE5F9557397F936" box="[542,642,1719,1745]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FD50F955724EF936" authorityName="Coiffait" authorityYear="1982" box="[683,859,1719,1745]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FD50F955724EF936" box="[683,859,1719,1745]" italics="true" pageId="1" pageNumber="585">M. cachemirica</emphasis>
</taxonomicName>
), with impunctate middle and mediobasal portion; scutellum without (
<typeStatus id="54EF8873EC107A11FE94F93970C6F912" box="[367,467,1755,1781]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FE02F93973BDF912" authorityName="Coiffait" authorityYear="1982" box="[505,680,1755,1781]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FE02F93973BDF912" box="[505,680,1755,1781]" italics="true" pageId="1" pageNumber="585">M. cachemirica</emphasis>
</taxonomicName>
) or with several very fine punctures in middle; punctation of elytra about as that on pronotum or denser, larger and deeper (
<typeStatus id="54EF8873EC107A11FCEEF91D726CF8FE" box="[789,889,1791,1817]" pageId="1" pageNumber="585" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC107A11FC5AF91D7547F8FE" authorityName="Coiffait" authorityYear="1982" box="[929,1106,1791,1817]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="585" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC107A11FC5AF91D7547F8FE" box="[929,1106,1791,1817]" italics="true" pageId="1" pageNumber="585">M. cachemirica</emphasis>
</taxonomicName>
), finer and sparser in middle along suture, punctures not forming disitnct longitudinal rows except for
<typeStatus id="54EF8873EC107A11FC1AF8C17554F8DA" box="[993,1089,1827,1853]" pageId="1" pageNumber="585" type="paratype">paratype</typeStatus>
with fused five rows of serial punctures; abdominal tergites with very fine and sparse punctation.
</paragraph>
<paragraph id="8BEB36D1EC107A13FF3CF88972BAFF32" blockId="1.[151,1437,998,2033]" lastBlockId="3.[151,1437,151,1293]" lastPageId="3" lastPageNumber="587" pageId="1" pageNumber="585">Head 1.3–1.4 times as broad as long, indistinctly elevated in middle, with very short and shallow anteocellar foveae about as long as diameter of ocellus; anterior portion between antennal insertion and anterior margins of eyes slightly concave. Eyes very large, convex. Ocelli large, well visible, situated at about level of posterior margins of eyes; distance between ocelli slightly greater than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi distinctly more than twice as long as preceding segment, from widest basal third gradually narrowed toward subacute apex. Antennae reaching basal portion of elytra when reclined; basal antennomere ovoid, slightly less than three times as long as broad, antennomere 2 distinctly shorter than basal antennomere, 3 narrower and slightly longer than 2, 4 small, about as long as broad, 5–6 distinctly longer and broader than 4, 7–8 slightly broader than 6, 9–10 indistinctly longer and broader than 8, apical antennomere about as long as two preceding antennomeres, from middle gradually narrowed toward subacute apex.</paragraph>
<caption id="DF2B6659EC137A12FF6CF93C70D3F8FD" ID-DOI="http://doi.org/10.5281/zenodo.12731008" ID-Zenodo-Dep="12731008" httpUri="https://zenodo.org/record/12731008/files/figure.png" pageId="2" pageNumber="586" startId="2.[151,264,1758,1782]" targetBox="[157,1429,185,1726]" targetPageId="2" targetType="figure">
<paragraph id="8BEB36D1EC137A12FF6CF93C70D3F8FD" blockId="2.[151,1437,1758,1819]" pageId="2" pageNumber="586">
<emphasis id="B920EAC3EC137A12FF6CF93C702EF911" bold="true" box="[151,315,1758,1782]" pageId="2" pageNumber="586">FIGURES 1–5.</emphasis>
Habitus of
<taxonomicName id="4C544D52EC137A12FE4FF93D735CF910" authorityName="Maklin" authorityYear="1880" box="[436,585,1759,1783]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC3EC137A12FE4FF93D735CF910" box="[436,585,1759,1783]" italics="true" pageId="2" pageNumber="586">Mannerheimia</emphasis>
</taxonomicName>
(holotypes): 1–
<taxonomicName id="4C544D52EC137A12FD11F93D729FF910" authorityName="Coiffait" authorityYear="1982" box="[746,906,1759,1783]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC137A12FD11F93D729FF910" box="[746,906,1759,1783]" italics="true" pageId="2" pageNumber="586">M. cachemirica</emphasis>
</taxonomicName>
, 2–
<taxonomicName id="4C544D52EC137A12FC4AF93D7504F911" authorityName="Coiffait" authorityYear="1982" box="[945,1041,1759,1782]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC137A12FC4AF93D7504F911" box="[945,1041,1759,1782]" italics="true" pageId="2" pageNumber="586">M. vicina</emphasis>
</taxonomicName>
, 3–
<taxonomicName id="4C544D52EC137A12FBC3F93D75AAF910" authorityName="Coiffait" authorityYear="1977" box="[1080,1215,1759,1783]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="emodensa">
<emphasis id="B920EAC3EC137A12FBC3F93D75AAF910" box="[1080,1215,1759,1783]" italics="true" pageId="2" pageNumber="586">M. emodensa</emphasis>
</taxonomicName>
, 4–
<taxonomicName id="4C544D52EC137A12FB1DF93D744EF910" authorityName="Coiffait" authorityYear="1982" box="[1254,1371,1759,1783]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B920EAC3EC137A12FB1DF93D744EF910" box="[1254,1371,1759,1783]" italics="true" pageId="2" pageNumber="586">M. obscura</emphasis>
</taxonomicName>
, 5–
<taxonomicName id="4C544D52EC137A12FA7AF93D7028F8FC" authority=", Scale" authorityName="Scale" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="586" phylum="Arthropoda" rank="species" species="scutellata">
<emphasis id="B920EAC3EC137A12FA7AF93D71EFF8FC" italics="true" pageId="2" pageNumber="586">M. scutellata</emphasis>
, Scale
</taxonomicName>
bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BEB36D1EC127A13FF3CFF3D72EEFE82" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">Pronotum 1.6–1.9 times as broad as long, 1.3 times as broad as head, widest in about middle, markedly more narrowed posteriad than anteriad; apical margin rounded, about as long as somewhat straight posterior margin; anterior angles widely rounded, indistincly protruded anteriad; posterior angles obtuse, with slightly sinuate laterobasal margins; each laterobasal portions with distinct wide impressions.</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFE8D73A9FE36" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">
Elytra slightly broader than long, slightly widened apicad, twice longer than pronotum; apical margins straight or rounded. Surface of elytra in middle with distinct, irregular and transverse elevations between punctures (
<typeStatus id="54EF8873EC127A13FAC3FE717489FE4A" box="[1336,1436,403,429]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FF4DFE557070FE36" authorityName="Coiffait" authorityYear="1982" box="[182,357,439,465]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FF4DFE557070FE36" box="[182,357,439,465]" italics="true" pageId="3" pageNumber="587">M. cachemirica</emphasis>
</taxonomicName>
). Hind wings fully developed.
</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFE397389FE12" blockId="3.[151,1437,151,1293]" box="[199,668,475,501]" pageId="3" pageNumber="587">Metatarsus 1.8 times as long as metatibia.</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFE1D744AFDDA" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">Abdomen about as broad as elytra or slightly broader, with two small transverse tomentose wing-folding spots in middle of abdominal tergite V; apical margin of abdominal tergite VII with distinct narrow palisade fringe.</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFDA5751EFD2A" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">
Male. Apical margins of abdominal tergite VIII and sternite VIII widely and rather deeply concave. Aedeagus with wide basal portion, gradually narrowed apically, with narrow median lobe and small rounded apex; parameres moderately wide, distinctly broadened in apical portions, slightly not reaching apex of median lobe; internal sac narrow and moderately long (
<figureCitation id="136F2A54EC127A13FE18FD517356FD2A" box="[483,579,691,717]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="3" pageNumber="587">Fig. 6, 8</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="136F2A54EC127A13FC38FD517512FD2A" box="[963,1031,691,717]" captionStart="FIGURES 6–10" captionStartId="4.[151,264,1954,1978]" captionTargetBox="[185,1404,459,1918]" captionTargetId="figure-141@4.[177,1411,453,1931]" captionTargetPageId="4" captionText="FIGURES 6–10. Aedeagus of Mannerheimia (Figs 6–7: M. cachemirica (holotype); Fig. 8: M. vicina (holotype); Figs. 9–10: M. emodensa (Nepal: Maharigaon): 6, 8–9–parameral view, 7, 10–lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12731014" httpUri="https://zenodo.org/record/12731014/files/figure.png" pageId="3" pageNumber="587">Fig. 7</figureCitation>
.
</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFD357080FD16" blockId="3.[151,1437,151,1293]" box="[199,405,727,753]" pageId="3" pageNumber="587">Female unstudied.</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFD187050FC66" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">
<emphasis id="B920EAC3EC127A13FF3CFD1870A7FCF2" bold="true" box="[199,434,762,789]" pageId="3" pageNumber="587">Comparative notes.</emphasis>
Based on the general shape of the body and the aedeagus,
<taxonomicName id="4C544D52EC127A13FBA1FD19741EFCF2" authorityName="Coiffait" authorityYear="1982" box="[1114,1291,763,789]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FBA1FD19741EFCF2" box="[1114,1291,763,789]" italics="true" pageId="3" pageNumber="587">M. cachemirica</emphasis>
</taxonomicName>
is similar to Nepalese
<taxonomicName id="4C544D52EC127A13FEFCFCFD708AFCDE" authorityName="Coiffait" authorityYear="1977" box="[263,415,799,825]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="emodensa">
<emphasis id="B920EAC3EC127A13FEFCFCFD708AFCDE" box="[263,415,799,825]" italics="true" pageId="3" pageNumber="587">M. emodensa</emphasis>
</taxonomicName>
, from which can be distinguished by the slightly narrower elytra, finer punctation of the pronotum and the elytra, slightly longer preapical antennomeres, and shorter parameres with markedly narrower apical portions.
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC127A13FF3CFC687073FC0A" pageId="3" pageNumber="587" type="distribution">
<paragraph id="8BEB36D1EC127A13FF3CFC6875A2FC42" blockId="3.[151,1437,151,1293]" box="[199,1207,906,933]" pageId="3" pageNumber="587">
<emphasis id="B920EAC3EC127A13FF3CFC687074FC43" bold="true" box="[199,353,906,932]" pageId="3" pageNumber="587">Distribution.</emphasis>
<taxonomicName id="4C544D52EC127A13FE9CFC697388FC42" authorityName="Coiffait" authorityYear="1982" box="[359,669,907,933]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FE9CFC697388FC42" box="[359,669,907,933]" italics="true" pageId="3" pageNumber="587">Mannerheimia cachemirica</emphasis>
</taxonomicName>
is known from two localities in Kashmir,
<collectingCountry id="F3437641EC127A13FB8DFC6975A6FC42" box="[1142,1203,907,933]" name="India" pageId="3" pageNumber="587">India</collectingCountry>
.
</paragraph>
<paragraph id="8BEB36D1EC127A13FF3CFC4C7073FC0A" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">
<emphasis id="B920EAC3EC127A13FF3CFC4C705EFC2F" bold="true" box="[199,331,942,968]" pageId="3" pageNumber="587">Bionomics.</emphasis>
The
<typeStatus id="54EF8873EC127A13FE7AFC4D70F6FC2E" box="[385,483,943,969]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FDFDFC4D737AFC2F" authorityName="Coiffait" authorityYear="1982" box="[518,623,943,968]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC127A13FDFDFC4D737AFC2F" box="[518,623,943,968]" italics="true" pageId="3" pageNumber="587">M. vicina</emphasis>
</taxonomicName>
was collected at altitudes from
<quantity id="4CAC9B34EC127A13FC37FC4D756FFC2E" box="[972,1146,943,969]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="3" pageNumber="587" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800 to 3100 m</quantity>
a.s.l. Detailed bionomical data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34E655AEC127A13FF3CFC1575B5FAEA" pageId="3" pageNumber="587" type="discussion">
<paragraph id="8BEB36D1EC127A13FF3CFC1575B5FAEA" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="587">
<emphasis id="B920EAC3EC127A13FF3CFC15702EFBF6" bold="true" box="[199,315,1015,1041]" pageId="3" pageNumber="587">Remarks.</emphasis>
<taxonomicName id="4C544D52EC127A13FEB0FC15739DFBF6" authorityName="Coiffait" authorityYear="1982" box="[331,648,1015,1041]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FEB0FC15739DFBF6" box="[331,648,1015,1041]" italics="true" pageId="3" pageNumber="587">Mannerheimia cachemirica</emphasis>
</taxonomicName>
was originally described based on
<specimenCount id="9D52FD58EC127A13FBBEFC157417FBF6" box="[1093,1282,1015,1041]" count="3" pageId="3" pageNumber="587" type="generic">three specimens</specimenCount>
from “
<collectingCountry id="F3437641EC127A13FA99FC15748DFBF6" box="[1378,1432,1015,1041]" name="India" pageId="3" pageNumber="587">Inde</collectingCountry>
, Cachemire, Aru…”.
<taxonomicName id="4C544D52EC127A13FE7CFBF9736FFBD3" authorityName="Coiffait" authorityYear="1982" box="[391,634,1051,1077]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC127A13FE7CFBF9736FFBD3" box="[391,634,1051,1077]" italics="true" pageId="3" pageNumber="587">Mannerheimia vicina</emphasis>
</taxonomicName>
was originally described based on a male and a female from “
<collectingCountry id="F3437641EC127A13FA99FBF9748DFBD2" box="[1378,1432,1051,1077]" name="India" pageId="3" pageNumber="587">Inde</collectingCountry>
, Cachemire, Chandanwari, près Pahalgam…”. I studied the
<typeStatus id="54EF8873EC127A13FCC7FBDD728BFBBE" box="[828,926,1087,1113]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
and the
<typeStatus id="54EF8873EC127A13FBFFFBDD7571FBBE" box="[1028,1124,1087,1113]" pageId="3" pageNumber="587" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FB75FBDD742AFBBE" authorityName="Coiffait" authorityYear="1982" box="[1166,1343,1087,1113]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FB75FBDD742AFBBE" box="[1166,1343,1087,1113]" italics="true" pageId="3" pageNumber="587">M. cachemirica</emphasis>
</taxonomicName>
and the
<typeStatus id="54EF8873EC127A13FF6CFB8171ECFB9A" box="[151,249,1123,1149]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FED9FB817098FB9B" authorityName="Coiffait" authorityYear="1982" box="[290,397,1123,1148]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC127A13FED9FB817098FB9B" box="[290,397,1123,1148]" italics="true" pageId="3" pageNumber="587">M. vicina</emphasis>
</taxonomicName>
, and during the study I have not found sufficient morphological differences between them. The
<typeStatus id="54EF8873EC127A13FF36FB65703AFB46" box="[205,303,1159,1185]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FEA1FB657319FB46" authorityName="Coiffait" authorityYear="1982" box="[346,524,1159,1185]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FEA1FB657319FB46" box="[346,524,1159,1185]" italics="true" pageId="3" pageNumber="587">M. cachemirica</emphasis>
</taxonomicName>
has slightly more transverse pronotum and distinctly elevated surface between punctures in the middle of the elytra, but the aedeagus of it is similar to the type of
<taxonomicName id="4C544D52EC127A13FB9CFB4975C6FB23" authorityName="Coiffait" authorityYear="1982" box="[1127,1235,1195,1220]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC127A13FB9CFB4975C6FB23" box="[1127,1235,1195,1220]" italics="true" pageId="3" pageNumber="587">M. vicina</emphasis>
</taxonomicName>
. The body of the
<typeStatus id="54EF8873EC127A13FF6CFB2D71E2FB0E" box="[151,247,1231,1257]" pageId="3" pageNumber="587" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FEE4FB2D70DAFB0E" authorityName="Coiffait" authorityYear="1982" box="[287,463,1231,1257]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FEE4FB2D70DAFB0E" box="[287,463,1231,1257]" italics="true" pageId="3" pageNumber="587">M. cachemirica</emphasis>
</taxonomicName>
is almost identical to that of the
<typeStatus id="54EF8873EC127A13FCBCFB2D72BCFB0E" box="[839,937,1231,1257]" pageId="3" pageNumber="587" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D52EC127A13FC2BFB2D752EFB0F" authorityName="Coiffait" authorityYear="1982" box="[976,1083,1231,1256]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC127A13FC2BFB2D752EFB0F" box="[976,1083,1231,1256]" italics="true" pageId="3" pageNumber="587">M. vicina</emphasis>
</taxonomicName>
. Thus, I synonymize
<taxonomicName id="4C544D52EC127A13FACAFB2D7489FB0F" authorityName="Coiffait" authorityYear="1982" box="[1329,1436,1231,1256]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B920EAC3EC127A13FACAFB2D7489FB0F" box="[1329,1436,1231,1256]" italics="true" pageId="3" pageNumber="587">M. vicina</emphasis>
</taxonomicName>
with
<taxonomicName id="4C544D52EC127A13FF2BFB11706AFAEA" authorityName="Coiffait" authorityYear="1982" box="[208,383,1267,1293]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="species" species="cachemirica">
<emphasis id="B920EAC3EC127A13FF2BFB11706AFAEA" box="[208,383,1267,1293]" italics="true" pageId="3" pageNumber="587">M. cachemirica</emphasis>
</taxonomicName>
, the species which has page priority in the same article (
<bibRefCitation id="EFC54B20EC127A13FC03FB117581FAEA" author="Coiffait, H." box="[1016,1172,1267,1293]" pageId="3" pageNumber="587" pagination="21 - 179" refId="ref4851" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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