264 lines
34 KiB
XML
264 lines
34 KiB
XML
<document id="2397200CC5D97596BC1B4367B4FD7065" ID-DOI="10.11646/zootaxa.5447.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="11150249" ID-ZooBank="367106D8-6248-49DA-8ABE-3125FC46BE08" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1715181318044" checkinUser="plazi" docAuthor="Chen, Jia-Heng, Yin, Wen-Qi & Shi, Hong-Liang" docDate="2024" docId="03FD87AC654B966AFF1FFBF329E25342" docLanguage="en" docName="zootaxa.5447.4.1.pdf" docOrigin="Zootaxa 5447 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) strumatus Chen & Yin & Shi 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="461" masterDocId="FFC4FFD465439660FF88FFCD2D25501F" masterDocTitle="On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)" masterLastPageNumber="472" masterPageNumber="451" pageNumber="459" updateTime="1715181786403" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="122DBDB60AC3F95E39A3D2CC40F97151">On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)</mods:title>
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<mods:name id="02502CD72771B0FB31D3C53BC3B8B179" type="personal">
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<mods:roleTerm id="55DF3D805FF519B0D73DCA73B175E6FB">Author</mods:roleTerm>
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<mods:namePart id="25308CA2F2A752F72AB8FC7B10A72853">Chen, Jia-Heng</mods:namePart>
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<mods:affiliation id="6A286D717CE2F38E8CF719D60948DEDD">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="CA249F0CADEE326E01BFEB03CAA3A2A2">Yin, Wen-Qi</mods:namePart>
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<mods:affiliation id="0FB0619DB2BB003325943667D41850D2">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="05E251944DCC52B86196C30F21CDDE85">Shi, Hong-Liang</mods:namePart>
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<mods:affiliation id="D5E563B57285A0025D4157C8901A1B1D">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:title id="8D788422E96ACD3C42C610BB793C6518">Zootaxa</mods:title>
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<mods:part id="BA52A19EC0B2C5360EE6755CADD5EF4C">
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<mods:date id="3363CB253884EF8E2A5EA6535BA1AF09">2024</mods:date>
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<mods:number id="EC05076026F57A829DCC277E87DDC671">2024-05-08</mods:number>
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<mods:number id="0CA06A5DB943C339C3BD236CE2517DDD">5447</mods:number>
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<mods:url id="E6E84ABD19254F7F7BAEFDC4E943F9D2">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:identifier id="BA166BC282DBE00308EDAF86ED790BAC" type="DOI">10.11646/zootaxa.5447.4.1</mods:identifier>
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<subSubSection id="C34E6531654B9668FF1FFBF32F4C5465" pageId="8" pageNumber="459" type="nomenclature">
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<paragraph id="8BEB36BA654B9668FF1FFBF32F9C5446" blockId="8.[151,697,1086,1185]" box="[151,697,1086,1113]" pageId="8" pageNumber="459">
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<heading id="D0A381D6654B9668FF1FFBF32F9C5446" bold="true" box="[151,697,1086,1113]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
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<emphasis id="B920EAA8654B9668FF1FFBF32F9C5446" bold="true" box="[151,697,1086,1113]" pageId="8" pageNumber="459">
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<taxonomicName id="4C544D39654B9668FF1FFBF32F715446" authority="Chen & Yin & Shi, 2024" authorityName="Chen & Yin & Shi" authorityYear="2024" box="[151,596,1086,1113]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="strumatus" status="sp. nov." subGenus="Wraseiellus">
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<emphasis id="B920EAA8654B9668FF1FFBF32C085446" bold="true" box="[151,301,1086,1113]" italics="true" pageId="8" pageNumber="459">Pterostichus</emphasis>
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(
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||
<emphasis id="B920EAA8654B9668FEB6FBF32CEF5446" bold="true" box="[318,458,1086,1113]" italics="true" pageId="8" pageNumber="459">Wraseiellus</emphasis>
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)
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<emphasis id="B920EAA8654B9668FE53FBF22F715446" bold="true" box="[475,596,1087,1113]" italics="true" pageId="8" pageNumber="459">strumatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21357D3654B9668FDD3FBF22F9C5446" box="[603,697,1087,1113]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEB36BA654B9668FF1FFBAE2F4C5465" blockId="8.[151,697,1086,1185]" box="[151,617,1123,1149]" pageId="8" pageNumber="459">
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Chinese vernacular name:
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||
<emphasis id="B920EAA8654B9668FE49FBAE2F4C5465" bold="true" box="[449,617,1123,1146]" pageId="8" pageNumber="459">OiẸaeữfflm</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34E6531654B9668FF1FFB4A2C4E54BE" box="[151,363,1159,1185]" pageId="8" pageNumber="459" type="description">
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<paragraph id="8BEB36BA654B9668FF1FFB4A2C4E54BE" blockId="8.[151,697,1086,1185]" box="[151,363,1159,1185]" pageId="8" pageNumber="459">
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<figureCitation id="136F2A3F654B9668FF1FFB4A2C2754BE" box="[151,258,1159,1185]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="8" pageNumber="459">Figures 4</figureCitation>
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,
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<figureCitation id="136F2A3F654B9668FE86FB4A2C1454BE" box="[270,305,1159,1185]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="8" pageNumber="459">9C</figureCitation>
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,
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<figureCitation id="136F2A3F654B9668FEB5FB4A2C4E54BE" box="[317,363,1159,1185]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="8" pageNumber="459">10C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6531654B9668FF1FFB022CD5552E" pageId="8" pageNumber="459" type="materials_examined">
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<paragraph id="8BEB36BA654B9668FF1FFB022CD5552E" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="459">
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<emphasis id="B920EAA8654B9668FF1FFB022C6F54F7" bold="true" box="[151,330,1230,1257]" pageId="8" pageNumber="459">Type materials</emphasis>
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.
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||
<materialsCitation id="3B3C3CE7654B9668FED2FB022F9E5512" collectingDate="2012-08-17" collectionCode="IZAS" collectorName="Hao Huang" elevation="1900" location="Emeishan Mt." municipality="Jiulinggang" pageId="8" pageNumber="459" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8818654B9668FED2FB022CE754F6" box="[346,450,1231,1257]" pageId="8" pageNumber="459" type="holotype">Holotype</typeStatus>
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||
(
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||
<collectionCode id="ED45AE7F654B9668FE5BFB022F3354F6" box="[467,534,1231,1257]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="8" pageNumber="459">IZAS</collectionCode>
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), male, body length =
|
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<quantity id="4CAC9B5F654B9668FC90FB022EA554F6" box="[792,896,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="8" pageNumber="459" unit="mm" value="13.8">13.8 mm</quantity>
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, “
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<location id="8E8B6061654B9668FC1FFB02291E54F6" LSID="urn:lsid:plazi:treatment:03FD87AC654B966AFF1FFBF329E25342:8E8B6061654B9668FC1FFB02291E54F6" box="[919,1083,1231,1257]" municipality="Jiulinggang" name="Emeishan Mt." pageId="8" pageNumber="459">Emeishan Mt.</location>
|
||
,
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||
<collectingMunicipality id="6B8FACC0654B9668FBC0FB0229EB54F6" box="[1096,1230,1231,1257]" pageId="8" pageNumber="459">Jiulinggang</collectingMunicipality>
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,
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<date id="FFEA107A654B9668FB52FB02286854F6" box="[1242,1357,1231,1257]" pageId="8" pageNumber="459" value="2012-08-17">
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<collectingDate id="EFAEE992654B9668FB52FB02286854F6" box="[1242,1357,1231,1257]" pageId="8" pageNumber="459" value="2012-08-17">2012.8.17</collectingDate>
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||
</date>
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||
.
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||
<collectingMethod id="52154EAD654B9668FAD1FB022DE05512" pageId="8" pageNumber="459">pitfall trap</collectingMethod>
|
||
,
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||
<quantity id="4CAC9B5F654B9668FF5CFB3E2C015512" box="[212,292,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="459" unit="m" value="1900.0">
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<elevation id="0079D189654B9668FF5CFB3E2C015512" box="[212,292,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="459" unit="m" value="1900.0">1900m</elevation>
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</quantity>
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,
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||
<collectorName id="26A1536C654B9668FEBBFB3E2C925512" box="[307,439,1267,1293]" pageId="8" pageNumber="459">Hao Huang</collectorName>
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||
leg.” [in Chinese] // “
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||
</materialsCitation>
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||
<typeStatus id="54EF8818654B9668FD33FB3E2E775512" box="[699,850,1267,1293]" pageId="8" pageNumber="459" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C544D39654B9668FCD3FB3E29DB5513" authority="Chen & Yin & Shi, 2024" authorityName="Chen & Yin & Shi" authorityYear="2024" box="[859,1278,1267,1293]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="strumatus" status="sp. nov." subGenus="Wraseiellus">
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||
<emphasis id="B920EAA8654B9668FCD3FB3E2EC35512" box="[859,998,1267,1293]" italics="true" pageId="8" pageNumber="459">Pterostichus</emphasis>
|
||
(
|
||
<emphasis id="B920EAA8654B9668FC72FB3E29585512" box="[1018,1149,1267,1293]" italics="true" pageId="8" pageNumber="459">Wraseiellus</emphasis>
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||
)
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||
<emphasis id="B920EAA8654B9668FB18FB3E29DB5513" box="[1168,1278,1267,1292]" italics="true" pageId="8" pageNumber="459">strumatus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A21357D3654B9668FA81FB3E28405512" box="[1289,1381,1267,1293]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
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, det. Chen & Shi. 2024” [red label].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6531654B9669FF4FFAF72CB950CA" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="diagnosis">
|
||
<paragraph id="8BEB36BA654B9668FF4FFAF72DEF564E" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="459">
|
||
<emphasis id="B920EAA8654B9668FF4FFAF72C1F554B" bold="true" box="[199,314,1338,1364]" pageId="8" pageNumber="459">Diagnosis</emphasis>
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||
. Frontal furrows shallow and short, extending a little behind level of anterior margin of eyes; vertex finely punctate; temporae short (TL/EYL 0.27–0.30). Maximum width of pronotum a little before middle; lateral margins strongly sinuate before posterior angles; posterior angles distinct, nearly rectangular, not forming prominent denticle; pronotal base finely and sparsely punctate; area between inner and outer basal foveal grooves depressed, ridge between outer groove and lateral margin distinct. Median lobe of aedeagus with a tubercle near base of ventral margin, apical lamella apically thickened and turned dorsally (
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||
<figureCitation id="136F2A3F654B9668FCC1FA222E855616" box="[841,928,1519,1545]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="8" pageNumber="459">Fig. 4B</figureCitation>
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); in dorsal view, apical lamella elongate, apex capitate and rounded, slightly bent to left (
|
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<figureCitation id="136F2A3F654B9668FDE5F9DE2F9A5632" box="[621,703,1555,1581]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="8" pageNumber="459">Fig 4C</figureCitation>
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||
); right paramere slender and long, apical portion acinaciform (
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<figureCitation id="136F2A3F654B9668FAFDF9DE2D9B564E" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="8" pageNumber="459">Fig 4D</figureCitation>
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||
).
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||
</paragraph>
|
||
<paragraph id="8BEB36BA654B9668FF4FF997297F571A" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="459">
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||
<emphasis id="B920EAA8654B9668FF4FF9972C43566B" bold="true" box="[199,358,1626,1652]" pageId="8" pageNumber="459">Comparisons</emphasis>
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||
. The new species is most similar to
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<taxonomicName id="4C544D39654B9668FC88F9962EB2566B" authorityName="Shi & Sciaky" authorityYear="2013" box="[768,919,1627,1653]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="crassiapex">
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||
<emphasis id="B920EAA8654B9668FC88F9962E34566B" box="[768,785,1627,1652]" italics="true" pageId="8" pageNumber="459">P</emphasis>
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||
.
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||
<emphasis id="B920EAA8654B9668FC97F9962EB2566B" box="[799,919,1627,1652]" italics="true" pageId="8" pageNumber="459">crassiapex</emphasis>
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</taxonomicName>
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. These two species can be distinguished from other
|
||
<taxonomicName id="4C544D39654B9668FF55F9B22C6A5686" box="[221,335,1663,1689]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="andrewesi">
|
||
<emphasis id="B920EAA8654B9668FF55F9B22C6A5686" box="[221,335,1663,1689]" italics="true" pageId="8" pageNumber="459">andrewesi</emphasis>
|
||
</taxonomicName>
|
||
-group species by the combination of following characters: (1) pronotum cordiform, posterior margin narrower than or subequal to anterior margin; (2) pronotum with one mid-lateral seta on each side; (3) pronotal basal fovea not convex between inner and outer grooves; (4) median lobe of aedeagus with apical lamella elongate (AL/AW> 1.40), apex capitate and rounded; (5) right paramere acinaciform.
|
||
</paragraph>
|
||
<paragraph id="8BEB36BA654B9669FF4FF8C22CB950CA" blockId="8.[151,1437,1230,2013]" lastBlockId="9.[151,1437,151,213]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
|
||
Comparing to
|
||
<taxonomicName id="4C544D39654B9668FEE4F8C22F215737" authorityName="Shi & Sciaky" authorityYear="2013" box="[364,516,1807,1833]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="crassiapex">
|
||
<emphasis id="B920EAA8654B9668FEE4F8C22C585737" box="[364,381,1807,1832]" italics="true" pageId="8" pageNumber="459">P</emphasis>
|
||
.
|
||
<emphasis id="B920EAA8654B9668FE04F8C22F215737" box="[396,516,1807,1832]" italics="true" pageId="8" pageNumber="459">crassiapex</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C544D39654B9668FD9BF8C22FBB5737" authority="Chen & Yin & Shi, 2024" authorityName="Chen & Yin & Shi" authorityYear="2024" box="[531,670,1807,1832]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="strumatus" status="sp. nov.">
|
||
<emphasis id="B920EAA8654B9668FD9BF8C22FBB5737" box="[531,670,1807,1832]" italics="true" pageId="8" pageNumber="459">P. strumatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EAA8654B9668FD2FF8C22E245736" bold="true" box="[679,769,1807,1833]" pageId="8" pageNumber="459">
|
||
<taxonomicNameLabel id="A21357D3654B9668FD2FF8C22E245736" box="[679,769,1807,1833]" pageId="8" pageNumber="459" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is different in: (1) pronotal posterior angles not prominent (versus forming a prominent denticle in
|
||
<taxonomicName id="4C544D39654B9668FDE2F8FE2E215753" authorityName="Shi & Sciaky" authorityYear="2013" box="[618,772,1843,1869]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="crassiapex">
|
||
<emphasis id="B920EAA8654B9668FDE2F8FE2F5E5753" box="[618,635,1843,1868]" italics="true" pageId="8" pageNumber="459">P</emphasis>
|
||
.
|
||
<emphasis id="B920EAA8654B9668FD04F8FE2E215753" box="[652,772,1843,1868]" italics="true" pageId="8" pageNumber="459">crassiapex</emphasis>
|
||
</taxonomicName>
|
||
); (2) basal foveae and pronotal mid-basal area finely and sparsely punctate (versus strongly and densely punctate in
|
||
<taxonomicName id="4C544D39654B9668FCC1F89A2EC1576F" authorityName="Shi & Sciaky" authorityYear="2013" box="[841,996,1879,1905]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="crassiapex">
|
||
<emphasis id="B920EAA8654B9668FCC1F89A2E7F576F" box="[841,858,1879,1904]" italics="true" pageId="8" pageNumber="459">P</emphasis>
|
||
.
|
||
<emphasis id="B920EAA8654B9668FCE4F89A2EC1576F" box="[876,996,1879,1904]" italics="true" pageId="8" pageNumber="459">crassiapex</emphasis>
|
||
</taxonomicName>
|
||
); (3) proepisterna finely and sparsely punctate (versus densely and coarsely punctate in
|
||
<taxonomicName id="4C544D39654B9668FD5AF8B62E4F578B" authorityName="Shi & Sciaky" authorityYear="2013" box="[722,874,1915,1941]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="crassiapex">
|
||
<emphasis id="B920EAA8654B9668FD5AF8B62FC6578B" box="[722,739,1915,1940]" italics="true" pageId="8" pageNumber="459">P</emphasis>
|
||
.
|
||
<emphasis id="B920EAA8654B9668FD7AF8B62E4F578B" box="[754,874,1915,1940]" italics="true" pageId="8" pageNumber="459">crassiapex</emphasis>
|
||
</taxonomicName>
|
||
); (4) median lobe of aedeagus with a tubercle on ventral margin; (5) in dorsal view, apical lamella less bent to left, less constricted at base, AL about one and a half times as AW (versus more strongly bent to left, more constricted at base, AL about twice as AW in
|
||
<taxonomicName id="4C544D39654B9668FB7CF80E28AE57C3" authorityName="Shi & Sciaky" authorityYear="2013" box="[1268,1419,1987,2013]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="crassiapex">
|
||
<emphasis id="B920EAA8654B9668FB7CF80E282057C3" box="[1268,1285,1987,2012]" italics="true" pageId="8" pageNumber="459">P</emphasis>
|
||
.
|
||
<emphasis id="B920EAA8654B9668FA9BF80E28AE57C3" box="[1299,1419,1987,2012]" italics="true" pageId="8" pageNumber="459">crassiapex</emphasis>
|
||
</taxonomicName>
|
||
); (6) ventral surface of apical lamella strongly concaved, forming ridged margin (versus weakly concaved, without ridge in
|
||
<taxonomicName id="4C544D39654A9669FF7CFF762CA950CB" authorityName="Shi & Sciaky" authorityYear="2013" box="[244,396,187,213]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="crassiapex">
|
||
<emphasis id="B920EAA8654A9669FF7CFF762C2050CB" box="[244,261,187,212]" italics="true" pageId="9" pageNumber="460">P</emphasis>
|
||
.
|
||
<emphasis id="B920EAA8654A9669FE9CFF762CA950CB" box="[276,396,187,212]" italics="true" pageId="9" pageNumber="460">crassiapex</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2B6632654A9669FF1FF9D82C94566A" ID-DOI="http://doi.org/10.5281/zenodo.11150263" ID-Zenodo-Dep="11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="9" pageNumber="460" startId="9.[151,250,1557,1581]" targetBox="[159,1427,282,1525]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BEB36BA654A9669FF1FF9D82C94566A" blockId="9.[151,1437,1557,1653]" pageId="9" pageNumber="460">
|
||
<emphasis id="B920EAA8654A9669FF1FF9D82C295632" bold="true" box="[151,268,1557,1581]" pageId="9" pageNumber="460">FIGURE 4</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C544D39654A9669FE1EF9D82F585632" authority="Chen & Yin & Shi, 2024" authorityName="Chen & Yin & Shi" authorityYear="2024" box="[406,637,1557,1581]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="strumatus" status="sp. nov.">
|
||
<emphasis id="B920EAA8654A9669FE1EF9D82F585632" box="[406,637,1557,1581]" italics="true" pageId="9" pageNumber="460">Pterostichus strumatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EAA8654A9669FD0AF9DB2FF75631" bold="true" box="[642,722,1558,1582]" pageId="9" pageNumber="460">
|
||
<taxonomicNameLabel id="A21357D3654A9669FD0AF9DB2FF75631" box="[642,722,1558,1582]" pageId="9" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations:
|
||
<emphasis id="B920EAA8654A9669FA94F9F72801564D" bold="true" box="[1308,1316,1594,1618]" pageId="9" pageNumber="460">t</emphasis>
|
||
tubercle on ventral margin of aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34E6531654A966AFF4FF96E2F0252EE" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460" type="description">
|
||
<paragraph id="8BEB36BA654A966AFF4FF96E2F0252EE" blockId="9.[151,1437,1699,2014]" lastBlockId="10.[151,1437,151,861]" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460">
|
||
<emphasis id="B920EAA8654A9669FF4FF96E2C7456A2" bold="true" box="[199,337,1699,1725]" pageId="9" pageNumber="460">Description</emphasis>
|
||
. BL
|
||
<quantity id="4CAC9B5F654A9669FE0FF9692CC856A1" box="[391,493,1700,1726]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="460" unit="mm" value="13.8">13.8 mm</quantity>
|
||
; dorsal surface dark brown, without metallic luster (
|
||
<figureCitation id="136F2A3F654A9669FBA6F96929AD56A1" box="[1070,1160,1700,1726]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="9" pageNumber="460">Fig. 4A</figureCitation>
|
||
).
|
||
<emphasis id="B920EAA8654A9669FB12F96929FF56A1" bold="true" box="[1178,1242,1700,1726]" pageId="9" pageNumber="460">Head</emphasis>
|
||
. Frons finely and sparsely punctate, relatively densely in frontal furrows, frontal furrows short and shallow, extending a little behind level of anterior margin of eyes; vertex finely punctate, with faint longitudinal wrinkles; temporae short (TL/EYL 0.28–0.31), not swollen behind eyes; paraocular grooves deep, impunctate and nearly straight, reaching posterior margin of eyes; antennomere 3 without accessory setae.
|
||
<emphasis id="B920EAA8654A9669FC84F8F92EA05751" bold="true" box="[780,901,1844,1870]" pageId="9" pageNumber="460">Pronotum</emphasis>
|
||
cordiform (PW/PL 1.32); anterior and posterior margins almost straight, lateral margins evenly curved from anterior angles to posterior third, abruptly sinuate before posterior angles; anterior margin slightly wider than posterior margin; anterior angles obtusely rounded, not projecting; posterior angles nearly rectangular, apex sharp, not forming prominent denticle; discal area glabrous; basal foveae relatively short and narrow, deep and areal punctate, area between inner and outer basal foveal grooves rugose and depressed, mid-basal area between two basal foveae sparsely punctate, ridge between outer groove and lateral margin distinct; a few very fine and sparse punctures present in lateral channels and, sometimes, on lateral sides of disc; one mid-lateral seta present on each side.
|
||
<emphasis id="B920EAA86549966AFC9CFF122E4550E6" bold="true" box="[788,864,223,249]" pageId="10" pageNumber="461">Elytra</emphasis>
|
||
oblong (EL/EW 1.42), lateral margins subparallel before middle, slightly expanded behind middle; humeral teeth small; intervals slightly convex; striae moderately deep, without punctures; parascutellar striae joined to stria 1, angular base of stria 1 short, apex free; umbilicate series on interval 9 composed of approximately 14–17 pores, sparse in middle and dense in anterior and posterior areas; male with microsculpture typically transverse on basal half, gradually turned to isodiametric towards apex.
|
||
<emphasis id="B920EAA86549966AFF1FFE5E2C0E51B3" bold="true" box="[151,299,402,429]" pageId="10" pageNumber="461">Ventral side</emphasis>
|
||
. Proepisterna and metaepisterna sparsely punctate, punctures on metaepisterna relatively coarser; sternite VII of males with a lunate transverse ridge at about anterior third, median region of sternite VII shallowly depressed posterior to the ridge.
|
||
<emphasis id="B920EAA86549966AFD8FFE162F1A51EA" bold="true" box="[519,575,475,501]" pageId="10" pageNumber="461">Legs</emphasis>
|
||
. Tarsomere 5 with three or four pairs of setae beneath.
|
||
<emphasis id="B920EAA86549966AFB38FE16287E51EB" bold="true" box="[1200,1371,474,501]" pageId="10" pageNumber="461">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="136F2A3F6549966AFAE2FE162D9F5206" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="10" pageNumber="461">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A3F6549966AFF4FFE322DDC5206" box="[199,249,511,537]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="10" pageNumber="461">10C</figureCitation>
|
||
). Median lobe of aedeagus relatively stout, tapered to apex; in lateral view, ventral margin with a distinct tubercle close to the basal bend (
|
||
<figureCitation id="136F2A3F6549966AFD8CFDEE2F785222" box="[516,605,547,573]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for B–D. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="10" pageNumber="461">Fig. 4B</figureCitation>
|
||
), ventral margin weakly bent to the ventral side before apex, apical orifice large, slightly turned to left, apical lamella apically thickened and turned dorsally; in dorsal view, apical lamella long (AL/AW 1.47), base slightly constricted, apex rounded and capitate, slightly bent to left; ventral surface of apical lamella strongly concaved on left side, marginal ridge present on the apical and right border of the concavity (
|
||
<figureCitation id="136F2A3F6549966AFF17FD7E2C2352D2" box="[159,262,691,717]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="10" pageNumber="461">Fig. 10C</figureCitation>
|
||
). Right paramere acinaciform, narrow and long, apex somewhat blunt. Endophallus of male genitalia not studied. Female genitalia unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E65316549966AFF4FFD362E2B5326" pageId="10" pageNumber="461" type="etymology">
|
||
<paragraph id="8BEB36BA6549966AFF4FFD362E2B5326" blockId="10.[151,1437,151,861]" pageId="10" pageNumber="461">
|
||
<emphasis id="B920EAA86549966AFF4FFD362C6D530A" bold="true" box="[199,328,763,789]" pageId="10" pageNumber="461">Etymology</emphasis>
|
||
. The scientific name of the new species is derived from the Latin root “
|
||
<emphasis id="B920EAA86549966AFBF0FD36299D530B" box="[1144,1208,763,788]" italics="true" pageId="10" pageNumber="461">strum</emphasis>
|
||
-” [tumor], referring to the median lobe of aedeagus with a peculiar swelling.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E65316549966AFF4FFC8F29E25342" box="[199,1223,834,861]" pageId="10" pageNumber="461" type="distribution">
|
||
<paragraph id="8BEB36BA6549966AFF4FFC8F29E25342" blockId="10.[151,1437,151,861]" box="[199,1223,834,861]" pageId="10" pageNumber="461">
|
||
<emphasis id="B920EAA86549966AFF4FFC8F2C7C5343" bold="true" box="[199,345,834,860]" pageId="10" pageNumber="461">Distribution</emphasis>
|
||
. Only known by the
|
||
<typeStatus id="54EF88186549966AFDCEFC8E2F8D5342" box="[582,680,835,861]" pageId="10" pageNumber="461" type="holotype">holotype</typeStatus>
|
||
collected on Emei Mountain,
|
||
<collectingRegion id="4990F8586549966AFC73FC8E29E65342" box="[1019,1219,835,861]" country="China" name="Sichuan" pageId="10" pageNumber="461">Sichuan Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |