225 lines
35 KiB
XML
225 lines
35 KiB
XML
<document id="DB38DE9F9354381BD9CB0B947C5CF0D2" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F116FFE77640FBA357187CF9" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) lunatus Assing 2017, spec. nov." docType="treatment" docUuid="ACD6BAAB-FE21-415D-903F-1D2CDEFE0607" docUuidSource="ZooBank" docVersion="9" lastPageNumber="240" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="239" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="93EE67F9EFD8AFF24192653D0D29706C">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="CCFCB099632928DC059B36388BAAF88E">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="E0B5D372A69FCCFBEB67CE0BF846F816">text</mods:typeOfResource>
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<mods:title id="56966F3D9F9A92F14562A18D84CFDDD4">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="FAB2B9BDBF38FC62FE07E3300741FE62">
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<mods:date id="264EFD60D6C40D3C623B5CABC79336CE">2017</mods:date>
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<mods:number id="8B0816A8EAA444342D5730290781CB62">2017-12-08</mods:number>
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<mods:number id="177D4FB08FEDB327B64A1A3C74A5E0D5">67</mods:number>
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<mods:identifier id="C9CC45097C446E5852466C9804103DF8" type="DOI">10.21248/contrib.entomol.67.2.213-246</mods:identifier>
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<treatment id="03FD87A1F116FFE77640FBA357187CF9" ID-DOI="http://doi.org/10.5281/zenodo.5889031" ID-GBIF-Taxon="192500462" ID-Zenodo-Dep="5889031" LSID="urn:lsid:zoobank.org:act:ACD6BAAB-FE21-415D-903F-1D2CDEFE0607" httpUri="http://treatment.plazi.org/id/03FD87A1F116FFE77640FBA357187CF9" lastPageId="27" lastPageNumber="240" pageId="26" pageNumber="239">
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<subSubSection id="C34E653CF116FFE67640FBA351077B97" pageId="26" pageNumber="239" type="nomenclature">
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<paragraph id="8BEB36B7F116FFE67640FBA350DD7BB6" blockId="26.[869,1395,1139,1231]" box="[869,1193,1139,1166]" pageId="26" pageNumber="239">
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<heading id="D0A381DBF116FFE67640FBA350DD7BB6" box="[869,1193,1139,1166]" fontSize="12" level="2" pageId="26" pageNumber="239" reason="2">
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<taxonomicName id="4C544D34F116FFE67640FBA3504E7BB6" ID-CoL="5DDDK" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[869,1082,1139,1166]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="lunatus" status="sp. nov." subGenus="Zyras">
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<emphasis id="B920EAA5F116FFE67640FBA357EF7BB6" box="[869,923,1139,1166]" italics="true" pageId="26" pageNumber="239">Zyras</emphasis>
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(
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<emphasis id="B920EAA5F116FFE6768CFBA357AB7BB6" box="[937,991,1139,1166]" italics="true" pageId="26" pageNumber="239">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F116FFE676C8FBA3504E7BB6" box="[1005,1082,1139,1166]" italics="true" pageId="26" pageNumber="239">lunatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21357DEF116FFE67165FBA350DD7BB6" box="[1088,1193,1139,1166]" pageId="26" pageNumber="239" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BEB36B7F116FFE67640FB4B51077B97" blockId="26.[869,1395,1139,1231]" box="[869,1395,1179,1199]" pageId="26" pageNumber="239">
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<uri id="FFC53AB5F116FFE67640FB4B51077B97" box="[869,1395,1179,1199]" pageId="26" pageNumber="239">
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urn:lsid:zoobank.org:act:
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<uuid id="FFF20C62F116FFE67137FB4B51077B97" box="[1042,1395,1179,1199]" pageId="26" pageNumber="239">ACD6BAAB-FE21-415D-903F-1D2CDEFE0607</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E653CF116FFE67640FB6B50547BF7" box="[869,1056,1211,1231]" pageId="26" pageNumber="239" type="description">
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<paragraph id="8BEB36B7F116FFE67640FB6B50547BF7" blockId="26.[869,1395,1139,1231]" box="[869,1056,1211,1231]" pageId="26" pageNumber="239">
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(
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<figureCitation id="136F2A32F116FFE6764CFB6B57D67BF7" box="[873,930,1211,1231]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="26" pageNumber="239">Figs 15</figureCitation>
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,
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<figureCitation id="136F2A32F116FFE6768CFB6B57CA7BF7" box="[937,958,1211,1231]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="26" pageNumber="239">29</figureCitation>
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,
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<figureCitation id="136F2A32F116FFE676E0FB6B57AE7BF7" box="[965,986,1211,1231]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="26" pageNumber="239">47</figureCitation>
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,
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<figureCitation id="136F2A32F116FFE676C4FB6B506F7BF7" box="[993,1051,1211,1231]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="26" pageNumber="239">99–101</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF116FFE6761AFB3957F47A99" pageId="26" pageNumber="239" type="materials_examined">
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<paragraph id="8BEB36B7F116FFE6761AFB3957F47A99" blockId="26.[831,1437,1257,1441]" pageId="26" pageNumber="239">
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<emphasis id="B920EAA5F116FFE6761AFB39502E7A39" bold="true" box="[831,1114,1257,1281]" pageId="26" pageNumber="239">Type material examined</emphasis>
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:
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<materialsCitation id="3B3C3CEAF116FFE6714AFB3951227A59" ID-GBIF-Occurrence="3450187347" collectingDate="2004-03-02" collectingDateMax="2004-03-18" collectingDateMin="2004-03-02" collectionCode="FIT" collectorName="Maruyama M." country="Malaysia" determinerName="V. Assing" elevation="250" location="Ulu Gombak Field Studies Centre" municipality="Ulu Gombak Field Studies Centre" pageId="26" pageNumber="239" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
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<typeStatus id="54EF8815F116FFE6714AFB3950A67A39" box="[1135,1234,1257,1281]" pageId="26" pageNumber="239" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D52FD3EF116FFE671C7FB3A50837A39" box="[1250,1271,1258,1281]" pageId="26" pageNumber="239">♂</specimenCount>
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: “
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<collectingCountry id="F3437627F116FFE67030FB3951EE7A39" box="[1301,1434,1257,1281]" name="Malaysia" pageId="26" pageNumber="239">MALAYSIA</collectingCountry>
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:
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<collectingRegion id="4990F855F116FFE67665FAD957E97A19" box="[832,925,1289,1313]" country="Malaysia" name="Selangor" pageId="26" pageNumber="239">Selangor</collectingRegion>
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,
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<collectingMunicipality id="6B8FACCDF116FFE6768BFAD951407A19" box="[942,1332,1289,1313]" country="Malaysia" name="Ulu Gombak Field Studies Centre" pageId="26" pageNumber="239" stateProvince="Selangor">Ulu Gombak Field Studies Centre</collectingMunicipality>
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(
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<quantity id="4CAC9B52F116FFE6706CFAD951FB7A19" box="[1353,1423,1289,1313]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="239" unit="m" value="250.0">
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<elevation id="0079D184F116FFE6706CFAD951FB7A19" box="[1353,1423,1289,1313]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="239" unit="m" value="250.0">250 m</elevation>
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</quantity>
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),
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<date id="FFEA1077F116FFE67665FAF957BB7A79" box="[832,975,1321,1345]" pageId="26" pageNumber="239" value="2004-03-02" valueMax="2004-03-18" valueMin="2004-03-02">
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<collectingDate id="EFAEE99FF116FFE67665FAF957BB7A79" box="[832,975,1321,1345]" pageId="26" pageNumber="239" value="2004-03-02" valueMax="2004-03-18" valueMin="2004-03-02">2–18.III.2004</collectingDate>
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</date>
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(FIT),
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<collectorName id="26A15361F116FFE67104FAFA50C97A79" box="[1057,1213,1322,1346]" pageId="26" pageNumber="239">Maruyama M.</collectorName>
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et al. / Holotypus ♂
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<taxonomicName id="4C544D34F116FFE67665FA9A57A57A59" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[832,977,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="lunatus" status="sp. nov.">
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<emphasis id="B920EAA5F116FFE67665FA9A57A57A59" box="[832,977,1353,1377]" italics="true" pageId="26" pageNumber="239">Zyras lunatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21357DEF116FFE676FEFA9A50607A59" box="[987,1044,1354,1377]" pageId="26" pageNumber="239" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<determinerName id="7E37DA73F116FFE67170FA9A50C37A59" box="[1109,1207,1353,1377]" pageId="26" pageNumber="239">
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V.
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<bibRefCitation id="EFC54B46F116FFE67155FA9950C37A59" author="ASSING, V." box="[1136,1207,1353,1377]" pageId="26" pageNumber="239" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
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</determinerName>
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2017” (cMar)
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</materialsCitation>
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||
.
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<materialsCitation id="3B3C3CEAF116FFE67046FA9A57F47A99" ID-GBIF-Occurrence="3455148307" pageId="26" pageNumber="239" typeStatus="paratype">
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<typeStatus id="54EF8815F116FFE67046FA9A570F7ABA" pageId="26" pageNumber="239" type="paratype">Paratypes</typeStatus>
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: 1 ♂ [teneral], 1 ♀: same data as holotype (cMar, cAss).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34E653CF116FFE67665FA1951017919" pageId="26" pageNumber="239" type="etymology">
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||
<paragraph id="8BEB36B7F116FFE67665FA1951017919" blockId="26.[832,1436,1481,1569]" pageId="26" pageNumber="239">
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||
<emphasis id="B920EAA5F116FFE67665FA1957C27AD9" bold="true" box="[832,950,1481,1505]" pageId="26" pageNumber="239">Etymology</emphasis>
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: The specific epithet is the past participle of the Latin verb lunare (to bend) and alludes to the subapically distinctly angled ventral process of the aedeagus.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E653CF116FFE77665F99951527C39" lastPageId="27" lastPageNumber="240" pageId="26" pageNumber="239" type="description">
|
||
<paragraph id="8BEB36B7F116FFE67665F99950FA7859" blockId="26.[832,1437,1609,2081]" pageId="26" pageNumber="239">
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||
<emphasis id="B920EAA5F116FFE67665F99957B67959" bold="true" box="[832,962,1609,1633]" pageId="26" pageNumber="239">Description</emphasis>
|
||
: Body length
|
||
<quantity id="4CAC9B52F116FFE67174F99950A67959" box="[1105,1234,1609,1633]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.6" metricValueMin="5.4" pageId="26" pageNumber="239" unit="mm" value="6.0" valueMax="6.6" valueMin="5.4">5.4–6.6 mm</quantity>
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||
; length of forebody
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||
<quantity id="4CAC9B52F116FFE67665F9B957B579B9" box="[832,961,1641,1665]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="26" pageNumber="239" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
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||
. Coloration (
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<figureCitation id="136F2A32F116FFE6717EF9B950DE79B9" box="[1115,1194,1641,1665]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="26" pageNumber="239">Figs 15</figureCitation>
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,
|
||
<figureCitation id="136F2A32F116FFE67198F9B950AD79B9" box="[1213,1241,1641,1665]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="26" pageNumber="239">29</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F116FFE671CFF9B9517379B9" box="[1258,1287,1641,1665]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="26" pageNumber="239">47</figureCitation>
|
||
): head darkbrown to blackish; pronotum dark-brown; elytra brown to dark-brown with the suture paler; abdomen reddish with the posterior margins of tergites III–V, the posterior portions of tergites VI–VII, and all of tergite VIII yellowish-red; legs yellowish; antennae brown with antennomeres I–II reddish; maxillary palpi reddish with the apical palpomere yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F116FFE77665F8B955747E99" blockId="26.[832,1437,1609,2081]" lastBlockId="27.[151,757,233,1857]" lastPageId="27" lastPageNumber="240" pageId="26" pageNumber="239">
|
||
Head (
|
||
<figureCitation id="136F2A32F116FFE676ADF8B957A178B9" box="[904,981,1897,1921]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="26" pageNumber="239">Fig. 29</figureCitation>
|
||
) distinctly transverse; punctation moderately coarse and moderately dense, median dorsal portion extensively impunctate; pubescence long, sub-erect to erect, and brown. Eyes moderately large, slightly longer than the postocular region in dorsal view. Antenna (
|
||
<figureCitation id="136F2A32F116FFE67694F7D957897719" box="[945,1021,2057,2081]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="26" pageNumber="239">Fig. 15</figureCitation>
|
||
) 2.0–
|
||
<quantity id="4CAC9B52F116FFE67164F7D950E67719" box="[1089,1170,2057,2081]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="26" pageNumber="239" unit="mm" value="2.1">2.1 mm</quantity>
|
||
long; antennomeres IV approximately as long as broad or weakly oblong, V as long as broad or weakly transverse, VI–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI of conical shape and short, shorter than the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F117FFE775B2FE7A55C07C39" blockId="27.[151,757,233,1857]" pageId="27" pageNumber="240">
|
||
Pronotum (
|
||
<figureCitation id="136F2A32F117FFE7743EFE79551C7EF9" box="[283,360,425,449]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="27" pageNumber="240">Fig. 29</figureCitation>
|
||
) 1.11–1.12 times as broad as long and 1.23–1.25 times as broad as head, broadest in anterior half, weakly convex in cross-section; posterior angles obtusely marked; lateral margins weakly to distinctly sinuate in posterior half in dorsal view; punctation rather coarse, dense, and somewhat irregularly distributed, laterally and near posterior margin with impunctate patches; midline rather broadly impunctate; pubescence long, pale, and sub-erect; lateral and anterior margins with numerous longer, stouter, and distinctly erect dark setae.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F117FFE775B2FCD954A27CF9" blockId="27.[151,757,233,1857]" pageId="27" pageNumber="240">
|
||
Elytra (
|
||
<figureCitation id="136F2A32F117FFE775C9FCD955437C19" box="[236,311,777,801]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="27" pageNumber="240">Fig. 29</figureCitation>
|
||
) 0.81–0.87 times as long as pronotum; punctation moderately dense and moderately coarse, regularly distributed, and defined; pubescence dense, long, pale, and sub-erect. Hind wings fully developed. Metatarsomere I shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F117FFE775B2FC19553C7959" blockId="27.[151,757,233,1857]" pageId="27" pageNumber="240">
|
||
Abdomen (
|
||
<figureCitation id="136F2A32F117FFE7743EFC1955117CD9" box="[283,357,969,993]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="27" pageNumber="240">Fig. 47</figureCitation>
|
||
) nearly as broad as elytra, with rather shallow anterior impressions on tergites III–V; tergites III–V without non-setiferous punctation in anterior impressions (individual punctures may be present), with a transverse row of approximately four setiferous punctures in posterior portions, and with usually eight setiferous punctures at posterior margins; tergite VI with a transverse band of fine non-setiferous punctation anteriorly, with sparse and fine non-setiferous punctures in median third, with some setiferous punctures laterally and with 8–10 setiferous punctures at posterior margin; tergite VII with a transverse band of dense non-setiferous punctures anteriorly, with sparse non-setiferous punctures in median third, and with two transverse rows of setiferous punctures posteriorly (each composed of approximately four punctures), posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing long black setae in posterior third, posterior margin weakly concave in the middle; all sternites with long, dense, and erect yellowish pubescence (rubbed off in the
|
||
<typeStatus id="54EF8815F117FFE775FEF99955497959" box="[219,317,1609,1633]" pageId="27" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F117FFE775B2F9BA55C57839" blockId="27.[151,757,233,1857]" pageId="27" pageNumber="240">
|
||
♂: sternite VIII with convex posterior margin; median lobe of aedeagus
|
||
<quantity id="4CAC9B52F117FFE7747FF95955877999" box="[346,499,1673,1697]" metricMagnitude="-4" metricUnit="m" metricValue="7.15" metricValueMax="7.5" metricValueMin="6.8" pageId="27" pageNumber="240" unit="mm" value="0.715" valueMax="0.75" valueMin="0.68">0.68–0.75 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="136F2A32F117FFE775B2F979556D79F9" box="[151,281,1705,1729]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="27" pageNumber="240">Figs 99–100</figureCitation>
|
||
; ventral process slender and subapically distinctly angled; paramere (
|
||
<figureCitation id="136F2A32F117FFE774E6F919566F79D9" box="[451,539,1737,1761]" captionStart="Figs 87–102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87–102: Zyras parvilobatus (87–89), Z. hlavaci (90–95; 90–92: holotype; 93–95: paratype from Selangor), Z. hirtiventris, holotype (96–98), Z. lunatus (99–101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (87–88, 90–91, 93–94, 96–97, 99–100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="27" pageNumber="240">Fig. 101</figureCitation>
|
||
)
|
||
<quantity id="4CAC9B52F117FFE77702F91956B479D9" box="[551,704,1737,1761]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" metricValueMax="8.4" metricValueMin="7.8" pageId="27" pageNumber="240" unit="mm" value="0.81" valueMax="0.84" valueMin="0.78">0.78–0.84 mm</quantity>
|
||
long and with short apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F117FFE775B2F8DA55657879" blockId="27.[151,757,233,1857]" pageId="27" pageNumber="240">♀: posterior margin of sternite VIII weakly concave in the middle.</paragraph>
|
||
<paragraph id="8BEB36B7F117FFE7763FFF3951527C39" blockId="27.[794,1399,233,769]" pageId="27" pageNumber="240">
|
||
<emphasis id="B920EAA5F117FFE7763FFF3957897E39" bold="true" box="[794,1021,233,257]" pageId="27" pageNumber="240">Comparative notes</emphasis>
|
||
: Like the syntopic
|
||
<taxonomicName id="4C544D34F117FFE77022FF3A51047E39" authorityName="Assing" authorityYear="2017" box="[1287,1392,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="hlavaci">
|
||
<emphasis id="B920EAA5F117FFE77022FF3A51047E39" box="[1287,1392,233,257]" italics="true" pageId="27" pageNumber="240">Z. hlavaci</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C544D34F117FFE7763FFEDA57F27E19" authorityName="Assing" authorityYear="2017" box="[794,902,265,289]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="lunatus">
|
||
<emphasis id="B920EAA5F117FFE7763FFEDA57F27E19" box="[794,902,265,289]" italics="true" pageId="27" pageNumber="240">Z. lunatus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C544D34F117FFE7711BFEDA50EE7E19" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1086,1178,265,289]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B920EAA5F117FFE7711BFEDA50EE7E19" box="[1086,1178,265,289]" italics="true" pageId="27" pageNumber="240">Z. hirtus</emphasis>
|
||
</taxonomicName>
|
||
group. It is distinguished from
|
||
<taxonomicName id="4C544D34F117FFE776E5FEFA505D7E79" authorityName="Assing" authorityYear="2017" box="[960,1065,297,321]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="hlavaci">
|
||
<emphasis id="B920EAA5F117FFE776E5FEFA505D7E79" box="[960,1065,297,321]" italics="true" pageId="27" pageNumber="240">Z. hlavaci</emphasis>
|
||
</taxonomicName>
|
||
by smaller body size, more slender antennae, finer and less extensive non-setiferous punctation on the abdomen, and by a smaller aedeagus with a subapically more strongly angled, apically shorter, and in ventral view broader and apically less acute ventral process. It differs from the even smaller
|
||
<taxonomicName id="4C544D34F117FFE771FBFE1A506A7D39" authority="ASSING, 2016" authorityName="ASSING" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="formosanus">
|
||
<emphasis id="B920EAA5F117FFE771FBFE1A51037ED9" box="[1246,1399,457,481]" italics="true" pageId="27" pageNumber="240">Z. formosanus</emphasis>
|
||
ASSING, 2016 (
|
||
<collectingRegion id="4990F855F117FFE776E1FE39506A7D39" box="[964,1054,489,513]" country="China" name="Taiwan)" pageId="27" pageNumber="240">Taiwan)</collectingRegion>
|
||
</taxonomicName>
|
||
by a darker forebody, darker antennae with shorter antennomeres IV and V, much longer elytra, the punctation pattern of the abdomen (
|
||
<taxonomicName id="4C544D34F117FFE77606FD9A57C87D59" authorityName="ASSING" authorityYear="2016" box="[803,956,585,609]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="formosanus">
|
||
<emphasis id="B920EAA5F117FFE77606FD9A57C87D59" box="[803,956,585,609]" italics="true" pageId="27" pageNumber="240">Z. formosanus</emphasis>
|
||
</taxonomicName>
|
||
: tergites III–VI with a transverse row of numerous long black setae posteriorly; tergites VI and VII with very sparse non-setiferous punctation anteriorly), and by a smaller aedeagus (despite larger body size) with a longer, subapically distinctly angled, and apically much more slender ventral process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF117FFE7763FFCF957187CF9" pageId="27" pageNumber="240" type="distribution">
|
||
<paragraph id="8BEB36B7F117FFE7763FFCF957187CF9" blockId="27.[794,1399,809,961]" pageId="27" pageNumber="240">
|
||
<emphasis id="B920EAA5F117FFE7763FFCF950EF7C79" bold="true" box="[794,1179,809,833]" pageId="27" pageNumber="240">Distribution and natural history</emphasis>
|
||
: The type locality is situated in
|
||
<collectingRegion id="4990F855F117FFE776F7FC9950457C59" box="[978,1073,841,865]" country="Malaysia" name="Selangor" pageId="27" pageNumber="240">Selangor</collectingRegion>
|
||
, Peninsular
|
||
<collectingCountry id="F3437627F117FFE771F7FC9951417C59" box="[1234,1333,841,865]" name="Malaysia" pageId="27" pageNumber="240">Malaysia</collectingCountry>
|
||
. The specimens were collected with flight interception traps, together with
|
||
<taxonomicName id="4C544D34F117FFE77697FC5A506F7C99" authorityName="Assing" authorityYear="2017" box="[946,1051,905,929]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="240" phylum="Arthropoda" rank="species" species="hlavaci">
|
||
<emphasis id="B920EAA5F117FFE77697FC5A506F7C99" box="[946,1051,905,929]" italics="true" pageId="27" pageNumber="240">Z. hlavaci</emphasis>
|
||
</taxonomicName>
|
||
. One of the
|
||
<typeStatus id="54EF8815F117FFE77184FC5A517E7C9A" box="[1185,1290,906,930]" pageId="27" pageNumber="240" type="paratype">paratypes</typeStatus>
|
||
is slightly teneral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |