259 lines
37 KiB
XML
259 lines
37 KiB
XML
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<mods:title id="36B361E519765116940970B4564B2213">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:title id="471236D147A8BA8327A7202F728ECA52">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="D44FB9D397C4DAFBFCCB8524D719A51D">2017</mods:date>
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<mods:number id="F26651637FE47F19DE78651BAFE23088">2017-12-08</mods:number>
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<mods:number id="8754F3658DFDBDD11E5576FC2CB9413E">67</mods:number>
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<treatment id="03FD87A1F103FFEC7598F98355827C79" ID-DOI="http://doi.org/10.5281/zenodo.5888794" ID-GBIF-Taxon="192500467" ID-Zenodo-Dep="5888794" LSID="urn:lsid:plazi:treatment:03FD87A1F103FFEC7598F98355827C79" httpUri="http://treatment.plazi.org/id/03FD87A1F103FFEC7598F98355827C79" lastPageId="16" lastPageNumber="229" pageId="15" pageNumber="228">
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<subSubSection id="C34E653CF103FFF37598F983566C7956" box="[189,536,1619,1646]" pageId="15" pageNumber="228" type="nomenclature">
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<paragraph id="8BEB36B7F103FFF37598F983566C7956" blockId="15.[189,536,1619,1679]" box="[189,536,1619,1646]" pageId="15" pageNumber="228">
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<heading id="D0A381DBF103FFF37598F983566C7956" box="[189,536,1619,1646]" centered="true" fontSize="12" level="2" pageId="15" pageNumber="228" reason="2">
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<taxonomicName id="4C544D34F103FFF37598F983566C7956" ID-CoL="5DDC4" authority="LAST, 1969" authorityName="Last" authorityYear="1969" box="[189,536,1619,1646]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="facundus" subGenus="Zyras">
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<emphasis id="B920EAA5F103FFF37598F98354877956" box="[189,243,1619,1646]" italics="true" pageId="15" pageNumber="228">Zyras</emphasis>
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(
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<emphasis id="B920EAA5F103FFF37424F98355437956" box="[257,311,1619,1646]" italics="true" pageId="15" pageNumber="228">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F103FFF37460F98355D07956" box="[325,420,1619,1646]" italics="true" pageId="15" pageNumber="228">facundus</emphasis>
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<bibRefCitation id="EFC54B46F103FFF3748FF983566C7956" author="LAST, H. R." box="[426,536,1619,1646]" pageId="15" pageNumber="228" pagination="279" refId="ref26523" refString="LAST, H. R. 1969: Changes in the nomenclature of the genus Zyras STEPHENS (Col., Staphylinidae). - Entomologist's Monthly Magazine 104 [1968]: 279." type="journal article" year="1969">LAST, 1969</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF103FFF37598F9AB551079B7" box="[189,356,1659,1679]" pageId="15" pageNumber="228" type="description">
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<paragraph id="8BEB36B7F103FFF37598F9AB551079B7" blockId="15.[189,536,1619,1679]" box="[189,356,1659,1679]" pageId="15" pageNumber="228">
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(
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<figureCitation id="136F2A32F103FFF375E4F9AB557979B7" box="[193,269,1659,1679]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="15" pageNumber="228">Figs 2, 24</figureCitation>
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,
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<figureCitation id="136F2A32F103FFF37431F9AB555D79B7" box="[276,297,1659,1679]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="15" pageNumber="228">36</figureCitation>
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,
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<figureCitation id="136F2A32F103FFF3740AF9AB551479B7" box="[303,352,1659,1679]" captionStart-0="Figs 56–57" captionStart-1="Figs 53–71" captionStartId-0="16.[189,233,233,257]" captionStartId-1="11.[151,192,1971,1993]" captionTargetBox-1="[155,1387,245,1958]" captionTargetId-1="figure-129@11.[154,1396,232,1963]" captionText-0="Figs 56–57; paramere (Fig. 58) longer than median lobe, 0.77 mm long, and with rather large and distinctly flattened apical lobe." captionText-1="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi-1="http://doi.org/10.5281/zenodo.5888738" httpUri-1="https://zenodo.org/record/5888738/files/figure.png" pageId="15" pageNumber="228">56–58</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF103FFF375B2F97C56B979D9" pageId="15" pageNumber="228" type="reference_group">
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<paragraph id="8BEB36B7F103FFF375B2F97C566579F8" blockId="15.[151,717,1707,1761]" box="[151,529,1707,1729]" pageId="15" pageNumber="228">
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<treatmentCitationGroup id="AB441199F103FFF375B2F97C566579F8" box="[151,529,1707,1729]" pageId="15" pageNumber="228">
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<treatmentCitation id="0AF510A6F103FFF375B2F97C567979F8" author="LAST, H. R." box="[151,525,1707,1729]" page="279" pageId="15" pageNumber="228" year="1969">
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<taxonomicName id="4C544D34F103FFF375B2F97C567979F8" ID-CoL="5DDC4" authority="LAST, 1969: 279" authorityName="Last" authorityPageNumber="279" authorityYear="1969" box="[151,525,1707,1729]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="facundus" subGenus="Zyras">
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<emphasis id="B920EAA5F103FFF375B2F97C54BF79F9" box="[151,203,1708,1729]" italics="true" pageId="15" pageNumber="228">Zyras</emphasis>
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(
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<emphasis id="B920EAA5F103FFF375FCF97C557979F9" box="[217,269,1708,1729]" italics="true" pageId="15" pageNumber="228">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F103FFF3743EF97B550479F8" box="[283,368,1707,1728]" italics="true" pageId="15" pageNumber="228">facundus</emphasis>
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||
<bibRefCitation id="EFC54B46F103FFF37450F97B567979F8" author="LAST, H. R." box="[373,525,1707,1729]" pageId="15" pageNumber="228" pagination="279" refId="ref26523" refString="LAST, H. R. 1969: Changes in the nomenclature of the genus Zyras STEPHENS (Col., Staphylinidae). - Entomologist's Monthly Magazine 104 [1968]: 279." type="journal article" year="1969">LAST, 1969: 279</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BEB36B7F103FFF375B2F91C56B979D9" blockId="15.[151,717,1707,1761]" box="[151,717,1739,1761]" pageId="15" pageNumber="228">
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||
<treatmentCitationGroup id="AB441199F103FFF375B2F91C56B979D9" box="[151,717,1739,1761]" pageId="15" pageNumber="228">
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||
<treatmentCitation id="0AF510A6F103FFF375B2F91C563379D8" author="CAMERON, M." box="[151,583,1739,1761]" page="18" pageId="15" pageNumber="228" year="1939">
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||
<taxonomicName id="4C544D34F103FFF375B2F91C563379D8" authority="CAMERON, 1939 b: 18" authorityName="Cameron" authorityPageNumber="18" authorityYear="1939" box="[151,583,1739,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="semirufus" subGenus="Zyras">
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<emphasis id="B920EAA5F103FFF375B2F91C54BF79D9" box="[151,203,1740,1761]" italics="true" pageId="15" pageNumber="228">Zyras</emphasis>
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||
(
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<emphasis id="B920EAA5F103FFF375FCF91C557979D9" box="[217,269,1740,1761]" italics="true" pageId="15" pageNumber="228">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F103FFF3743EF91B550379D8" box="[283,375,1739,1760]" italics="true" pageId="15" pageNumber="228">semirufus</emphasis>
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||
<bibRefCitation id="EFC54B46F103FFF37459F91B563379D8" author="CAMERON, M." box="[380,583,1739,1761]" pageId="15" pageNumber="228" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON, 1939b: 18</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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; preoccupied.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E653CF103FFF375B2F8D955F07739" pageId="15" pageNumber="228" type="materials_examined">
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<paragraph id="8BEB36B7F103FFF375B2F8D955F07739" blockId="15.[151,756,1801,2049]" pageId="15" pageNumber="228">
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<emphasis id="B920EAA5F103FFF375B2F8D955D97819" bold="true" box="[151,429,1801,1825]" pageId="15" pageNumber="228">Type material examined</emphasis>
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||
:
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<materialsCitation id="3B3C3CEAF103FFF37498F8DA55CD78D9" ID-GBIF-Occurrence="3455148301" pageId="15" pageNumber="228" typeStatus="lectotype">
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<typeStatus id="54EF8815F103FFF37498F8DA565C781A" box="[445,552,1802,1826]" pageId="15" pageNumber="228" type="lectotype">Lectotype</typeStatus>
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♂, present designation: “F. C. Drescher, G. Tangkoeban Prahoe, 4000–5000 Voet, Preanger,
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<collectingRegion id="4990F855F103FFF374E2F89A55827859" box="[455,502,1866,1889]" country="Indonesia" name="Jawa Timur" pageId="15" pageNumber="228">Java</collectingRegion>
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,
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<date id="FFEA1077F103FFF37727F89956F77859" box="[514,643,1865,1889]" pageId="15" pageNumber="228" value="1930-07-20">20.VII.1930</date>
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/
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<taxonomicName id="4C544D34F103FFF377B8F89A54B078B9" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="semirufa">
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<emphasis id="B920EAA5F103FFF377B8F89A54B078B9" italics="true" pageId="15" pageNumber="228">Z. semirufa</emphasis>
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</taxonomicName>
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[sic]
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<collectingRegion id="4990F855F103FFF37426F8B9554E78B9" box="[259,314,1897,1921]" country="Myanmar" name="Chin" pageId="15" pageNumber="228">Cam</collectingRegion>
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. Type /
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<typeStatus id="54EF8815F103FFF374B4F8B9559378B9" box="[401,487,1897,1921]" pageId="15" pageNumber="228" type="syntype">Syntype</typeStatus>
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||
/ M. Cameron. Bequest. B.M. 1955-147 /
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<typeStatus id="54EF8815F103FFF37473F85A55B9789A" box="[342,461,1930,1954]" pageId="15" pageNumber="228" type="lectotype">Lectotypus</typeStatus>
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♂
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<taxonomicName id="4C544D34F103FFF374D0F85A54CB78F9" authority="Cameron" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="semirufus">
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<emphasis id="B920EAA5F103FFF374D0F85A56D57899" box="[501,673,1929,1953]" italics="true" pageId="15" pageNumber="228">Zyras semirufus</emphasis>
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Cameron
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||
</taxonomicName>
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||
, desig. V.
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<bibRefCitation id="EFC54B46F103FFF37411F87955C878F9" author="ASSING, V." box="[308,444,1961,1985]" pageId="15" pageNumber="228" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing 2017</bibRefCitation>
|
||
/
|
||
<taxonomicName id="4C544D34F103FFF374FEF87A56C878F9" authority="Last" authorityName="Last" authorityYear="1969" box="[475,700,1961,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="facundus">
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||
<emphasis id="B920EAA5F103FFF374FEF87A56F678F9" box="[475,642,1961,1985]" italics="true" pageId="15" pageNumber="228">Zyras facundus</emphasis>
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||
Last
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||
</taxonomicName>
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, det. V.
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<bibRefCitation id="EFC54B46F103FFF37596F819554878D9" author="ASSING, V." box="[179,316,1993,2017]" pageId="15" pageNumber="228" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing 2017</bibRefCitation>
|
||
” (
|
||
<collectionCode id="ED45AE72F103FFF37473F81955D978D9" box="[342,429,1993,2017]" pageId="15" pageNumber="228">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B3C3CEAF103FFF374E7F81955F07739" ID-GBIF-Occurrence="3455148304" pageId="15" pageNumber="228" typeStatus="paralectotype">
|
||
<typeStatus id="54EF8815F103FFF374E7F819562278D9" box="[450,598,1993,2017]" pageId="15" pageNumber="228" type="paralectotype">Paralectotype</typeStatus>
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♀: same data as
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<typeStatus id="54EF8815F103FFF37596F83955667739" box="[179,274,2025,2049]" pageId="15" pageNumber="228" type="holotype">holotype</typeStatus>
|
||
(
|
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<collectionCode id="ED45AE72F103FFF37404F839550C7739" box="[289,376,2025,2049]" pageId="15" pageNumber="228">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF103FFF3763FFF3957B17D39" pageId="15" pageNumber="228" type="discussion">
|
||
<paragraph id="8BEB36B7F103FFF3763FFF3957B17D39" blockId="15.[794,1399,233,513]" pageId="15" pageNumber="228">
|
||
<emphasis id="B920EAA5F103FFF3763FFF3957FD7E39" bold="true" box="[794,905,233,257]" pageId="15" pageNumber="228">Comment</emphasis>
|
||
: The original description is based on an unspecified number of
|
||
<typeStatus id="54EF8815F103FFF37105FEDA50087E1A" box="[1056,1148,266,290]" pageId="15" pageNumber="228" type="syntype">syntypes</typeStatus>
|
||
from “G. Tangkoeban Prahoe” (
|
||
<bibRefCitation id="EFC54B46F103FFF376A5FEF9504F7E79" author="CAMERON, M." box="[896,1083,297,321]" pageId="15" pageNumber="228" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON 1939b</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFC54B46F103FFF3716BFEF950A77E79" author="LAST, H. R." box="[1102,1235,297,321]" pageId="15" pageNumber="228" pagination="279" refId="ref26523" refString="LAST, H. R. 1969: Changes in the nomenclature of the genus Zyras STEPHENS (Col., Staphylinidae). - Entomologist's Monthly Magazine 104 [1968]: 279." type="journal article" year="1969">LAST (1969)</bibRefCitation>
|
||
replaced
|
||
<taxonomicName id="4C544D34F103FFF37018FEFA57877E59" authority="CAMERON" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="semirufus">
|
||
<emphasis id="B920EAA5F103FFF37018FEFA57F47E59" italics="true" pageId="15" pageNumber="228">Zyras semirufus</emphasis>
|
||
CAMERON
|
||
</taxonomicName>
|
||
, a name preoccupied by
|
||
<taxonomicName id="4C544D34F103FFF371DBFE9A505B7EB9" authority="(BERNHAUER, 1902)" baseAuthorityName="BERNHAUER" baseAuthorityYear="1902" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="semirufus">
|
||
<emphasis id="B920EAA5F103FFF371DBFE9A57257EB9" italics="true" pageId="15" pageNumber="228">Zyras semirufus</emphasis>
|
||
(BERNHAUER, 1902)
|
||
</taxonomicName>
|
||
in the subgenus
|
||
<taxonomicName id="4C544D34F103FFF371C9FEB9500F7E99" authority="GEMMINGER & HAROLD, 1968" authorityName="GEMMINGER & HAROLD" authorityYear="1968" class="Insecta" family="Staphylinidae" genus="Glossacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EAA5F103FFF371C9FEB951027EB9" box="[1260,1398,361,385]" italics="true" pageId="15" pageNumber="228">Glossacantha</emphasis>
|
||
GEMMINGER & HAROLD, 1968
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC54B46F103FFF371B0FE5A511F7E99" author="HLAVAC, P. & NEWTON, A. F. & MARUYAMA, M." box="[1173,1387,393,421]" pageId="15" pageNumber="228" pagination="1 - 151" refId="ref26453" refString="HLAVAC, P.; NEWTON, A. F. & MARUYAMA, M. 2011: World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae). - Zootaxa 3075: 1 - 151." type="journal article" year="2011">HLAVÁČ et al. 2011</bibRefCitation>
|
||
), with the nomen novum
|
||
<taxonomicName id="4C544D34F103FFF37107FE7A50E87EF9" authorityName="Last" authorityYear="1969" box="[1058,1180,425,449]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="228" phylum="Arthropoda" rank="species" species="facundus">
|
||
<emphasis id="B920EAA5F103FFF37107FE7A50E87EF9" box="[1058,1180,425,449]" italics="true" pageId="15" pageNumber="228">Z. facundus</emphasis>
|
||
</taxonomicName>
|
||
. Two
|
||
<typeStatus id="54EF8815F103FFF371C5FE7A51487EFA" box="[1248,1340,426,450]" pageId="15" pageNumber="228" type="syntype">syntypes</typeStatus>
|
||
were located in the Cameron collection. A male is designated as the
|
||
<typeStatus id="54EF8815F103FFF37678FE3957B57D39" box="[861,961,489,513]" pageId="15" pageNumber="228" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF103FFEC763FFDF955B87DD9" lastPageId="16" lastPageNumber="229" pageId="15" pageNumber="228" type="description">
|
||
<paragraph id="8BEB36B7F103FFF3763FFDF951577C19" blockId="15.[794,1399,553,2081]" pageId="15" pageNumber="228">
|
||
<emphasis id="B920EAA5F103FFF3763FFDF957C77D79" bold="true" box="[794,947,553,577]" pageId="15" pageNumber="228">Redescription</emphasis>
|
||
: Body length
|
||
<quantity id="4CAC9B52F103FFF37168FDF950BA7D79" box="[1101,1230,553,577]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.6" metricValueMin="6.4" pageId="15" pageNumber="228" unit="mm" value="7.0" valueMax="7.6" valueMin="6.4">6.4–7.6 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CAC9B52F103FFF3767EFD9957A87D59" box="[859,988,585,609]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="15" pageNumber="228" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="136F2A32F103FFF37154FD9950AD7D59" box="[1137,1241,585,609]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="15" pageNumber="228">Figs 2, 24</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F103FFF371C6FD9951747D59" box="[1251,1280,585,609]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="15" pageNumber="228">36</figureCitation>
|
||
): forebody dark-brown to blackish-brown; abdomen pale-reddish with the posterior portions of tergites VII and VIII weakly infuscate; legs yellowish; antennae blackish-brown with antennomeres I–III slightly to distinctly paler and XI at least apically slightly paler; maxillary palpi brown to dark-brown with the apical palpomere yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F103FFF3763FFCF950ED7B79" blockId="15.[794,1399,553,2081]" pageId="15" pageNumber="228">
|
||
Head (
|
||
<figureCitation id="136F2A32F103FFF3764EFCF957C17C79" box="[875,949,809,833]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="15" pageNumber="228">Fig. 24</figureCitation>
|
||
) distinctly transverse, median portion extensively impunctate; punctures in lateral portions sparse and rather fine to moderately coarse. Eyes longer than postocular region in dorsal view. Antenna (
|
||
<figureCitation id="136F2A32F103FFF37014FC59511A7C99" box="[1329,1390,905,929]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="15" pageNumber="228">Fig. 2</figureCitation>
|
||
) 2.0 mm long; antennomeres IV approximately as long as broad, V weakly transverse, VI–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F103FFF3763FFB9A57FF7A19" blockId="15.[794,1399,553,2081]" pageId="15" pageNumber="228">
|
||
Pronotum (
|
||
<figureCitation id="136F2A32F103FFF376B8FB99579C7B59" box="[925,1000,1097,1121]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="15" pageNumber="228">Fig. 24</figureCitation>
|
||
) 1.25–1.26 times as broad as long and 1.16–1.23 times as broad as head, broadest in anterior half, moderately convex in cross-section; punctation rather fine, sparse, and irregularly distributed, with extensive impunctate areas on either side of midline, on either side of the broadly impunctate midline with a series of punctures.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F103FFF3763FFAF957CA7AF9" blockId="15.[794,1399,553,2081]" pageId="15" pageNumber="228">
|
||
Elytra (
|
||
<figureCitation id="136F2A32F103FFF3764AFAF957CD7A79" box="[879,953,1321,1345]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="15" pageNumber="228">Fig. 24</figureCitation>
|
||
) 0.88–0.91 times as long as pronotum; punctation sparse and rather fine, regularly or slightly irregularly distributed. Hind wings present. Metatarsomere I slender, approximately as long as the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F103FFF3763FFA19571F78D9" blockId="15.[794,1399,553,2081]" pageId="15" pageNumber="228">
|
||
Abdomen (
|
||
<figureCitation id="136F2A32F103FFF376B7FA1957A87AD9" box="[914,988,1481,1505]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="15" pageNumber="228">Fig. 36</figureCitation>
|
||
) narrower than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of weakly defined non-setiferous punctures, with a lateral setiferous puncture on either side and with four (tergites III–IV) or six (tergite V) setiferous punctures bearing long black setae near posterior margin; tergite VI with a transverse row of sparse non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side and with six setiferous punctures bearing long dark setae near posterior margin; tergite VII with or without few scattered non-setiferous punctures anteriorly and with two transverse series of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing long black setae in posterior twofifths, posterior margin convex, in the middle truncate to convex.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F103FFEC763FF83A55027E79" blockId="15.[794,1399,553,2081]" lastBlockId="16.[189,794,233,385]" lastPageId="16" lastPageNumber="229" pageId="15" pageNumber="228">
|
||
♂: posterior margin of sternite VIII strongly convex; median lobe of aedeagus
|
||
<quantity id="4CAC9B52F103FFF3710CF7D950FC7719" box="[1065,1160,2057,2081]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="228" unit="mm" value="0.65">0.65 mm</quantity>
|
||
long and shaped as in Figs 56–57; paramere (
|
||
<figureCitation id="136F2A32F11CFFEC7494FF3955887E39" box="[433,508,233,257]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="16" pageNumber="229">Fig. 58</figureCitation>
|
||
) longer than median lobe, 0.77 mm long, and with rather large and distinctly flattened apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11CFFEC7598FE9A55437EB9" blockId="16.[189,794,233,385]" pageId="16" pageNumber="229">♀: posterior margin of sternite VIII weakly concave in the middle.</paragraph>
|
||
<paragraph id="8BEB36B7F11CFFEC7598FE7955B87DD9" blockId="16.[189,794,425,737]" pageId="16" pageNumber="229">
|
||
<emphasis id="B920EAA5F11CFFEC7598FE7955E57EF9" bold="true" box="[189,401,425,449]" pageId="16" pageNumber="229">Comparative notes</emphasis>
|
||
:
|
||
<taxonomicName id="4C544D34F11CFFEC7485FE7A56367EF9" authorityName="Last" authorityYear="1969" box="[416,578,425,449]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="facundus">
|
||
<emphasis id="B920EAA5F11CFFEC7485FE7A56367EF9" box="[416,578,425,449]" italics="true" pageId="16" pageNumber="229">Zyras facundus</emphasis>
|
||
</taxonomicName>
|
||
is characterized by rather massive antennae, a rather strongly transverse pronotum with distinctive punctation, rather sparse and fine punctation of the elytra, the coloration and punctation pattern of the pronotum, and by the morphology of the aedeagus. Regarding the modifications of the apical lobe of the paramere, this species is similar to
|
||
<taxonomicName id="4C544D34F11CFFEC7598FD5A55A97D99" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[189,477,649,673]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="ambulans">
|
||
<emphasis id="B920EAA5F11CFFEC7598FD5A554B7D99" box="[189,319,649,673]" italics="true" pageId="16" pageNumber="229">Z. ambulans</emphasis>
|
||
<bibRefCitation id="EFC54B46F11CFFEC746FFD5955A97D99" author="ASSING, V." box="[330,477,649,673]" pageId="16" pageNumber="229" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3437627F11CFFEC770DFD5956FE7D99" box="[552,650,649,673]" name="Thailand" pageId="16" pageNumber="229">Thailand</collectingCountry>
|
||
, from which it is distinguished by completely different coloration and much longer elytra alone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF11CFFEC7598FCD955827C79" pageId="16" pageNumber="229" type="distribution">
|
||
<paragraph id="8BEB36B7F11CFFEC7598FCD955827C79" blockId="16.[189,794,777,834]" pageId="16" pageNumber="229">
|
||
<emphasis id="B920EAA5F11CFFEC7598FCD955327C19" bold="true" box="[189,326,777,801]" pageId="16" pageNumber="229">Distribution</emphasis>
|
||
:
|
||
<taxonomicName id="4C544D34F11CFFEC747DFCDA55897C19" authorityName="Last" authorityYear="1969" box="[344,509,777,801]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="facundus">
|
||
<emphasis id="B920EAA5F11CFFEC747DFCDA55897C19" box="[344,509,777,801]" italics="true" pageId="16" pageNumber="229">Zyras facundus</emphasis>
|
||
</taxonomicName>
|
||
is currently known only from the
|
||
<typeStatus id="54EF8815F11CFFEC7405FCFA553A7C7A" box="[288,334,810,834]" pageId="16" pageNumber="229">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="4990F855F11CFFEC74E6FCFA55857C79" box="[451,497,810,833]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="229">Java</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |