243 lines
25 KiB
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243 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice" docDate="2022" docId="EA2B83521F5A560B9F375CE2836D3A15" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) irakensis Guéorguiev & Wrase & Assmann & Muilwijk & Machard 2022, sp. nov." docType="treatment" docUuid="95E89306-626A-468D-BA68-F16489ABB9F3" docUuidSource="ZooBank" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
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<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation>Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
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<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
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<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Machard, Patrice</mods:namePart>
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<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>71</mods:start>
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<mods:end>122</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/dez.69.83840</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.83840</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-1324-1-71</mods:identifier>
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<mods:identifier type="ZooBank">7DEB0587A3734B06BAADCA6102EC6581</mods:identifier>
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<mods:identifier type="Pensoft-UUID">DF59A16543B05799A4FD7B66E721FC76</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:95E89306-626A-468D-BA68-F16489ABB9F3" httpUri="http://treatment.plazi.org/id/EA2B83521F5A560B9F375CE2836D3A15" lastPageNumber="71" pageId="0" pageNumber="71">
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<subSubSection pageId="0" pageNumber="71" type="nomenclature">
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<paragraph pageId="0" pageNumber="71">
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17.
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<taxonomicName LSID="http://zoobank.org/95E89306-626A-468D-BA68-F16489ABB9F3" authority="Guéorguiev & Wrase & Assmann & Muilwijk & Machard, 2022" authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) irakensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="description">
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<paragraph pageId="0" pageNumber="71">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4B</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche & Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">, 6C</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15D</figureCitation>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="type locality">
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<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="71">Iraq, Al Anbar Governorate, Ar Rutba District, ca. 115 km east of Ar-Rutbah Town.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="type material">
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<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
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♀, 'IRAQ, Al-Anbar / Rutba, singled / 115 km. E of town / 8.I.1978 [w, h] // No. 350 /
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<normalizedToken originalValue="Topál">Topal</normalizedToken>
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& Zilahy [w, h]' (HNHM); paratype ♀, 'IRAQ, Al-Anbar / Rutba, singled / at 30 km W of / town, 9.I.1978 [w, h] // No. 351 /
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<normalizedToken originalValue="Topál">Topal</normalizedToken>
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& Zilahy [w, h]' (NMNHS).
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</paragraph>
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<paragraph pageId="0" pageNumber="71">TME: 2 specimens. TGE: 1♀.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="etymology">
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<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, referring to the name of the country in which this new species was found.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="diagnosis">
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<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<taxonomicName authorityName="Guéorguiev & Wrase & Assmann & Muilwijk & Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus irakensis</emphasis>
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</taxonomicName>
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is distinct from all other species of "
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<taxonomicName baseAuthorityName="Reiche & Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
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</taxonomicName>
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" group by the orange-brown color of integument (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
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), mesotarsomeres and metatarsomeres dorsally flattened and slightly grooved (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Metatarsus, dorsal view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, left metatarsus, female specimen, Aziziyah, Jafara District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, left metatarsus, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) irakensis, sp. nov., left metatarsus (black arrows indicate longitudinal grooves on metatarsomeres 2, 3 and 4), holotype; D. P. (E.) jordanensis, sp. nov., right metatarsus, holotype; E. P. (E.) languidus (Reiche & Saulcy, 1855), right metatarsus, male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure6" httpUri="https://binary.pensoft.net/fig/695643" pageId="0" pageNumber="71">6C</figureCitation>
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), pronotum with sides to base slightly convex and posterior angles laterally not prominent, and distal enlargement of bursa copulatrix (Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
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); the last character may be an autapomorphy. Among Asian representatives of species group, it is the form with the highest mean values for EL/PL (= 2.69) and EL/EW (= 1.63), and, together with
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
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</taxonomicName>
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sp. nov., the one possessing lowest index for PW/PB (= 1.10).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="description">
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<paragraph pageId="0" pageNumber="71">Description.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
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.
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</emphasis>
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Specimens of moderate size for
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<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
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</taxonomicName>
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species (BL: 8.30-9.00 mm; BW: 2.90-3.20 mm), with elongate and fairly convex body (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4B</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
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.
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</emphasis>
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See Table
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<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
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.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
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.
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</emphasis>
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Body and appendages orange-brown, with head somewhat darker than the rest of the body and ventral surface lighter than dorsal surface. Integument slightly to moderately shiny, head and pronotum as shiny as elytra, ventral surface somewhat shinier than dorsal surface.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
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.
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</emphasis>
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Pronotum with evident microreticulation, sculpticells regular isodiametric. Elytral intervals, scutellum and lateral gutter with distinct isodiametric sculpticells, sculpticells of basal margin more or less reduced. Ventral surface with isodiametric or slight transverse sculpticells, less apparent on epipleura and middle coxa. Head impunctate and smooth, only frontal furrows and lateral ends of clypeus, with very fine and short wrinkles. Pronotum surface mostly smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled; basal foveae and adjacent lateral areas shallowly punctate. Elytral intervals without evident punctures. Abdominal ventrites 1-3 finely wrinkled at sides, smooth medially.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
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.
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</emphasis>
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About one time and a third as wide as pronotum (PW/HW= 1.30-1.32). Eyes long, moderately convex. Labrum subrectangular, barely shorter than clypeus, with anterior margin concave medially. Frontoclypeal suture distinct in middle, reduced at sides. Frontal furrows punctiform, shallow. Paraorbital sulci straight, fine, backward surpassing posterior margin of eye, not reaching level of posterior supraorbital pore.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
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.
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</emphasis>
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Pronotum about a tenth wider than long (PW/PL= 1.13), with widest point at medial third. Anterior transverse impression indistinct; posterior transverse impression indistinct to barely distinct. Sides not sinuate, shallowly convex anteriorly, barely convex posteriorly; anterior bead present laterally, lacking or present in medial ninth to tenth; basal bead present laterally, reduced to absent in medial third. Metepisternum slightly longer than wide, MA/MI= 0.91-0.95.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
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.
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</emphasis>
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Elongate, about one and two thirds as long as wide (EL/EW= 1.62-1.63), two and two thirds as long as pronotum (EL/PL= 2.65-2.72), and almost one and a half times as wide as pronotum (EW/PW= 1.45-1.47), with widest point at first half of third quarter. Parascutellar striole and striae 1-8 shallowly impressed, superficially to indistinctly punctate; striole short, not joining stria 1; bases of striae not reaching basal bead (except for stria 2 in paratype). Interval 3 with three discal setiferous punctures (posterior puncture lacking on left elytron of holotype), all adjoining stria 2; one additional discal puncture adjoining stria 4 at anterior quarter of right elytron of paratype. Umbilicate setiferous series with 16 punctures on left elytron and 17 on right elytron in holotype, respectively, with 16 and 15 punctures in paratype.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
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.
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</emphasis>
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Posterior side of profemur with one or two setae in basal third and one in medial third. Mesofemur mostly with 4 or 5 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with a few long setae, one in basal third and one-two in apical half.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
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.
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</emphasis>
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Unknown.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15D</figureCitation>
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). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Bursa copulatrix two-chambered, with a shorter, but wider basal part and a longer, but narrower apical enlargement. Spermathecal canal connected in basal third of receptaculum.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="comparisons">
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<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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In addition to characters mentioned in
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<normalizedToken originalValue="“Diagnosis”">"Diagnosis"</normalizedToken>
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, the new species differs from
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
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<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
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</taxonomicName>
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sp. nov., by: (1) head wider, compared to pronotum (PW/HW: 1.30-1.32, vs. PW/HW: 1.35-1.42); (2) pronotum with sides less constricted anteriorly and posteriorly (PW/PA: 1.34-1.38 and PW/PB: 1.09-1.11, vs. PW/PA: 1.39-1.52 and PW/PB: 1.15-1.22); (3) elytra longer compared to their width and the length of the pronotum (EL/EW: 1.62-1.63 and EL/PL: 2.65-2.72, vs. EL/EW: 1.56-1.60 and EL/PL: 2.41-2.60).
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</paragraph>
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<paragraph pageId="0" pageNumber="71">
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For differences between
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
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<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
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</taxonomicName>
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sp. nov. and such between
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
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<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
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</taxonomicName>
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, see
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<normalizedToken originalValue="“Comparisons”">"Comparisons"</normalizedToken>
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under the latter species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="habitat">
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<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
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<paragraph pageId="0" pageNumber="71">Nothing is known about the bionomics of this species.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="distribution">
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<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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It is currently known from two localities situated in the Ar Rutba District, which is the largest district of Al Anbar Governorate, Western Iraq (Fig.
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
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). The holotype was collected about 115 km east of Ar-Rutbah Town whilst the paratype was found about 30 km west of the same town. Most of the area of the Ar Rutba District is a high plateau.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |