215 lines
18 KiB
XML
215 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.85.68658" ID-GBIF-Dataset="3117a885-cd0e-45e7-8868-924db5870919" ID-Pensoft-Pub="1314-2607-85-119" ID-Pensoft-UUID="8935A382F3A3566AAD7C650E899F46D5" ID-Zenodo-Dep="5491729" ID-ZooBank="9D5B00CA75F04E1FB84711D490F769CD" ModsDocID="1314-2607-85-119" checkinTime="1630544391347" checkinUser="pensoft" docAuthor="Tribull, Carly Melissa, Pankowski, Madeline V. & Colombo, Wesley Dondoni" docDate="2021" docId="A3D9AA3EF5F359D6870D62EFCEB278D2" docLanguage="en" docName="JourHymenoptRes 85: 119-133" docOrigin="Journal of Hymenoptera Research 85" docPubDate="2021-08-31" docSource="http://dx.doi.org/10.3897/jhr.85.68658" docTitle="Archeonesia Tribull, Pankowski & Colombo 2021, gen. nov." docType="treatment" docUuid="6764DE3C-095F-4067-B96D-79706FF609C5" docUuidSource="ZooBank" docVersion="4" id="8935A382F3A3566AAD7C650E899F46D5" lastPageNumber="119" masterDocId="8935A382F3A3566AAD7C650E899F46D5" masterDocTitle="A new genus and species of Pristocerinae (Hymenoptera, Bethylidae) from upper Eocene Baltic amber with a review of conspecific association from insect fossils" masterLastPageNumber="133" masterPageNumber="119" pageNumber="119" updateTime="1678756359737" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and species of Pristocerinae (Hymenoptera, Bethylidae) from upper Eocene Baltic amber with a review of conspecific association from insect fossils</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tribull, Carly Melissa</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1044-0486</mods:nameIdentifier>
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<mods:affiliation>Farmingdale State College (State University of New York), Biology Department, Farmingdale NY, USA</mods:affiliation>
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<mods:nameIdentifier type="email">cmtribull@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pankowski, Madeline V.</mods:namePart>
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<mods:affiliation>Rockville, Maryland, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Colombo, Wesley Dondoni</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8729-4635</mods:nameIdentifier>
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<mods:affiliation>Universidade Federal do Espirito Santo, Departamento de Ciencias Biologicas, Vitoria ES, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>85</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>119</mods:start>
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<mods:end>133</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.85.68658</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.85.68658</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-85-119</mods:identifier>
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<mods:identifier type="ZooBank">9D5B00CA75F04E1FB84711D490F769CD</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8935A382F3A3566AAD7C650E899F46D5</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5491729</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="185459853" LSID="urn:lsid:zoobank.org:act:6764DE3C-095F-4067-B96D-79706FF609C5" httpUri="http://treatment.plazi.org/id/A3D9AA3EF5F359D6870D62EFCEB278D2" lastPageNumber="119" pageId="0" pageNumber="119">
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<subSubSection pageId="0" pageNumber="119" type="nomenclature">
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<paragraph pageId="0" pageNumber="119">
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Genus †
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<taxonomicName LSID="http://zoobank.org/6764DE3C-095F-4067-B96D-79706FF609C5" authority="Tribull, Pankowski & Colombo" authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus" status="gen. nov.">Archeonesia Tribull, Pankowski & Colombo</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="119">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="type-species">
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<paragraph pageId="0" pageNumber="119">Type-species.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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†
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">A. eocena</emphasis>
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</taxonomicName>
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Tribull, Pankowski & Colombo, sp. nov. by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Description.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Male</emphasis>
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Archeonesia eocena Tribull, Pankowski & Colombo, gen. et. sp. nov. A dorsal habitus of male holotype and female allotype B ventral habitus of male holotype and female allotype. Scale bars: 1 mm (A, B)." figureDoi="10.3897/jhr.85.68658.figure1" httpUri="https://binary.pensoft.net/fig/582442" pageId="0" pageNumber="119">1</figureCitation>
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-
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. † Archeonesia eocena Tribull, Pankowski & Colombo, gen. et. sp. nov., male holotype A head, ventro-frontal view B metasoma, ventral view C prothorax and mesothorax, dorsal view D mesosoma, ventral view E wings, dorsal view. Scale bars: 500 μm (C); 1 mm (A, B, D, E)." figureDoi="10.3897/jhr.85.68658.figure4" httpUri="https://binary.pensoft.net/fig/582445" pageId="0" pageNumber="119">4</figureCitation>
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). Head, pronotum, mesoscutum, metapectal-propodeal complex, petiole, antenna, and metasoma dark castaneous to black; wings hyaline. Head as long as wide and subquadrate, not globoid in lateral view. Clypeus with triangular median lobe, visible dorsally, lateral lobe reduced. Median clypeal carina delimited, lower than frons. Flagellomere longer than wide, with first flagellomeres larger than distal ones; flagellar pubescence erected; pedicel shorter than flagellomere I, apex dilated. Eye located touching mandibular base, glabrous, bulging. Frons weakly coriaceous, punctures large and sparse. Frontal line not visible. Ocellus large, salient. Frontal angle of ocellar triangle in obtuse angle. Anterior ocellus posterior to supra-ocular line. Occipital carina present. Dorsal pronotal area wider than long, weakly coriaceous, punctures small and sparse. Metafurcal pit oval. Posterior mesofurcal pit crown-shaped. Notaulus present, large, converging posteriorly, smooth. Parapsidal signum shorter than notauli. Forewing with three cells closed (C, R, 1Cu), distal flexion line visible, 2r-rs&Rs vein tubular, long, well pigmented, angled, not converging posteriorly to anterior margin, R1 vein tubular, long. Pterostigma enlarged, lanceolate. Mesoscutum-mesoscutellar sulcus present, posterior margin strongly incurved medially. Metanotum well-developed medially. Metapectal-propodeal disc not visible. Mesotibia without spines. Metasoma polished. Ninth abdominal segment with margin weakly incurved, undivided.
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</paragraph>
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<caption ID-Zenodo-Dep="5491731" doi="10.3897/jhr.85.68658.figure1" httpUri="https://binary.pensoft.net/fig/582442" pageId="0" pageNumber="119" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 1.</emphasis>
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†
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<taxonomicName authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">Archeonesia eocena</emphasis>
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</taxonomicName>
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Tribull, Pankowski & Colombo, gen. et. sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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dorsal habitus of male holotype and female allotype
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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ventral habitus of male holotype and female allotype. Scale bars: 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="119">A, B</emphasis>
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).
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</paragraph>
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</caption>
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<caption ID-Zenodo-Dep="5491733" doi="10.3897/jhr.85.68658.figure2" httpUri="https://binary.pensoft.net/fig/582443" pageId="0" pageNumber="119" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 2.</emphasis>
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†
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<taxonomicName authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">Archeonesia eocena</emphasis>
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</taxonomicName>
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Tribull, Pankowski & Colombo, gen. et. sp. nov. Illustration of dorsal habitus of male holotype and female allotype. Scale bar: 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Female</emphasis>
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Archeonesia eocena Tribull, Pankowski & Colombo, gen. et. sp. nov. A dorsal habitus of male holotype and female allotype B ventral habitus of male holotype and female allotype. Scale bars: 1 mm (A, B)." figureDoi="10.3897/jhr.85.68658.figure1" httpUri="https://binary.pensoft.net/fig/582442" pageId="0" pageNumber="119">1</figureCitation>
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-
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Archeonesia eocena Tribull, Pankowski & Colombo, gen. et. sp. nov. Illustration of ventral habitus of male holotype and female allotype. Scale bar: 1 mm." figureDoi="10.3897/jhr.85.68658.figure3" httpUri="https://binary.pensoft.net/fig/582444" pageId="0" pageNumber="119">3</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Archeonesia eocena Tribull, Pankowski & Colombo, gen. et. sp. nov., female allotype A habitus, dorsal view B habitus, ventral view C head, ventro-lateral view D eye, with red arrow pointing to lower margin. Scale bars: 650 μm (D), 1 mm (A-C)." figureDoi="10.3897/jhr.85.68658.figure5" httpUri="https://binary.pensoft.net/fig/582446" pageId="0" pageNumber="119">5</figureCitation>
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). Head, pronotum, mesoscutum, metapectal-propodeal complex, petiole, antenna, and metasoma light to dark brown. Head longer than wide and rectangular, not globoid in lateral view. Clypeus with median lobe trapezoidal, visible dorsally, lateral lobe reduced. Median clypeal carina not visible. Flagellomere as long as wide, with first flagellomeres wider than distal ones; flagellar pubescence appressed; pedicel barrel shaped, as long as flagellomere I. Eye located almost touching mandibular base, glabrous, reduced, flat. Frons weakly coriaceous, punctures small and sparse. Frontal line not visible. Ocelli absent. Dorsal pronotal area smooth, longer than mesoscutellum medially. Metafurcal pit oval. Posterior mesofurcal pit oval. Anteromesoscutum with anterior margin straight. Notaulus absent. Parapsidal signum absent. Mesoscutum-mesoscutellar sulcus absent. Mesopleuron visible dorsally, broad. Apterous. Mesotibia spinose. Metapectal-propodeal disc long, broadly in contact with anteromesoscutum, anterior margin wider than posterior one, lateral margin almost parallel in dorsal view, weak constriction at spiracles present. Second abdominal segment without flap expanded laterally.
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</paragraph>
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<caption ID-Zenodo-Dep="5491735" doi="10.3897/jhr.85.68658.figure3" httpUri="https://binary.pensoft.net/fig/582444" pageId="0" pageNumber="119" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 3.</emphasis>
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†
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<taxonomicName authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">Archeonesia eocena</emphasis>
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</taxonomicName>
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Tribull, Pankowski & Colombo, gen. et. sp. nov. Illustration of ventral habitus of male holotype and female allotype. Scale bar: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="etymology">
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<paragraph pageId="0" pageNumber="119">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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The name †
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<taxonomicName authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Archeonesia</emphasis>
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</taxonomicName>
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comes from the genus
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<taxonomicName authorityName="Alencar & Azevedo" authorityYear="2018" class="Insecta" family="Bethylidae" genus="Acrenesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acrenesia" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="119">Acrenesia</emphasis>
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</taxonomicName>
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, which the female is closest to. The prefix
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<normalizedToken originalValue="‘Archeo’">'Archeo'</normalizedToken>
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represents the extinct nature of the genus. Gender feminine.
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</paragraph>
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<caption ID-Zenodo-Dep="5491737" doi="10.3897/jhr.85.68658.figure4" httpUri="https://binary.pensoft.net/fig/582445" pageId="0" pageNumber="119" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 4.</emphasis>
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†
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<taxonomicName authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">Archeonesia eocena</emphasis>
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</taxonomicName>
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Tribull, Pankowski & Colombo, gen. et. sp. nov., male holotype
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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head, ventro-frontal view
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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metasoma, ventral view
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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prothorax and mesothorax, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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mesosoma, ventral view
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<emphasis bold="true" pageId="0" pageNumber="119">E</emphasis>
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wings, dorsal view. Scale bars: 500
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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); 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="119">A, B, D, E</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="included species">
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<paragraph pageId="0" pageNumber="119">Included species.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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†
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">A. eocena</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="distribution">
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<paragraph pageId="0" pageNumber="119">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="119">Baltic amber, Russian Federation.</paragraph>
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<caption ID-Zenodo-Dep="5491739" doi="10.3897/jhr.85.68658.figure5" httpUri="https://binary.pensoft.net/fig/582446" pageId="0" pageNumber="119" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 5.</emphasis>
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†
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<taxonomicName authorityName="Tribull, Pankowski & Colombo" authorityYear="2021" class="Insecta" family="Bethylidae" genus="Archeonesia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archeonesia eocena" order="Hymenoptera" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="eocena">
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<emphasis italics="true" pageId="0" pageNumber="119">Archeonesia eocena</emphasis>
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</taxonomicName>
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Tribull, Pankowski & Colombo, gen. et. sp. nov., female allotype
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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habitus, ventral view
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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head, ventro-lateral view
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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eye, with red arrow pointing to lower margin. Scale bars: 650
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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), 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="119">A-C</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |