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<mods:title id="1767412C1E2B49EDC46A0FE79FEE0DED">A taxonomic review and phylogeny of the genera Hoplopisthius Senna, 1892 and Carcinopisthius Kolbe, 1892 (Coleoptera: Brentidae, Hoplopisthiini)</mods:title>
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<paragraph id="E65936BEFF997865A4B9FD3A3BBAB9EB" blockId="3.[151,478,738,764]" box="[151,478,738,764]" pageId="3" pageNumber="4">
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<taxonomicName id="21E64D3DFF997865A4B9FD3A3BBAB9EB" ID-CoL="4YP5" authority="Senna, 1892" authorityName="Senna" authorityYear="1892" box="[151,478,738,764]" class="Insecta" family="Brentidae" genus="Hoplopisthius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="D492EAACFF997865A4B9FD3A3BBAB9EB" bold="true" box="[151,478,738,764]" pageId="3" pageNumber="4">
<emphasis id="D492EAACFF997865A4B9FD3A3B24B9EB" bold="true" box="[151,320,738,764]" italics="true" pageId="3" pageNumber="4">Hoplopisthius</emphasis>
<bibRefCitation id="82774B4FFF997865A569FD3A3BBAB9EB" author="Senna" box="[327,478,738,764]" pageId="3" pageNumber="26" refString="Senna, A. (1892). Viaggio di Leonardo Fea in Birmania e regioni vicine. XLV. Brentidi. Annali del Museo civico di Storia naturale, Genova, 12, 429 - 494. [published on May 19 th 1892]" type="journal article" year="1892">Senna, 1892</bibRefCitation>
</emphasis>
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<taxonomicName id="21E64D3DFF997865A4B9FCF23B94B856" ID-CoL="4YP5" authority="Senna, 1892: 451" authorityName="Senna" authorityPageNumber="451" authorityYear="1892" box="[151,496,810,833]" class="Insecta" family="Brentidae" genus="Hoplopisthius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="D492EAACFF997865A4B9FCF23B49B856" box="[151,301,810,833]" italics="true" pageId="3" pageNumber="4">Hoplopisthius</emphasis>
<treatmentCitation id="674710AFFF997865A51DFCF23B94B856" author="Senna" box="[307,496,810,833]" page="451" pageId="3" pageNumber="4" year="1892">
<bibRefCitation id="82774B4FFF997865A51DFCF23BDFB856" author="Senna" box="[307,443,810,833]" pageId="3" pageNumber="26" refString="Senna, A. (1892). Viaggio di Leonardo Fea in Birmania e regioni vicine. XLV. Brentidi. Annali del Museo civico di Storia naturale, Genova, 12, 429 - 494. [published on May 19 th 1892]" type="journal article" year="1892">Senna, 1892</bibRefCitation>
: 451
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.
<typeStatus id="395D881CFF997865A5D4FCF2384BB856" box="[506,559,810,833]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName id="21E64D3DFF997865A6BFFCF23E5EB856" ID-CoL="3MMQJ" authority="Senna, 1892" authorityName="Senna" authorityYear="1892" box="[657,1082,810,833]" class="Insecta" family="Brentidae" genus="Hoplopisthius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="trichemerus">
<emphasis id="D492EAACFF997865A6BFFCF239C8B856" box="[657,940,810,833]" italics="true" pageId="3" pageNumber="4">Hoplopisthius trichemerus</emphasis>
<bibRefCitation id="82774B4FFF997865A79DFCF23E5EB856" author="Senna" box="[947,1082,810,833]" pageId="3" pageNumber="26" refString="Senna, A. (1892). Viaggio di Leonardo Fea in Birmania e regioni vicine. XLV. Brentidi. Annali del Museo civico di Storia naturale, Genova, 12, 429 - 494. [published on May 19 th 1892]" type="journal article" year="1892">Senna, 1892</bibRefCitation>
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, by monotypy.
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<subSubSection id="AEFC6535FF997865A4B9FCB73B19BFE8" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="E65936BEFF997865A4B9FCB73B19BFE8" blockId="3.[151,1437,879,1354]" pageId="3" pageNumber="4">
<emphasis id="D492EAACFF997865A4B9FCB73B71B89E" bold="true" box="[151,277,879,905]" pageId="3" pageNumber="4">Diagnosis</emphasis>
. Body oblong. Head flattened, especially the prorostrum, without well marked vertex-frontal median groove. Venter of mesorostrum with two small projections below insertion of antennae (
<figureCitation id="7EDD2A3BFF997865A12BFC4D3F04B8B8" box="[1285,1376,917,943]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Figs 21</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A143FC4D3FEAB8B8" box="[1389,1422,917,943]" captionStart="FIGURES 35 47" captionStartId="16.[151,273,1905,1929]" captionTargetBox="[205,1382,227,1867]" captionTargetId="figure@16.[189,1404,194,1887]" captionTargetPageId="16" captionText="FIGURES 35 47. Hoplopisthius (Carcinopisthius) fruhstorferi (Kolbe, 1892). 35, Head (Ψ), dorsal; 36, Head (Ψ), venter; 37, Antenna; 38, Metatarsi and apex of metatibia; 39, Apex of elytra (Ψ); 40, Apex of elytra (♂); 41, Abdomen (♂), venter; 42, Tergite VII (Ψ); 43, Spiculum gastrale; 44, Tegmen; 45, Apex of gonopods IX; 46, Spermatheca; 47, Apex of tergite VIII (Ψ). Scale: 1 mm except fig. 45: 0.17 mm and fig. 46 47: 0.33 mm." httpUri="https://zenodo.org/record/196138/files/figure.png" pageId="3" pageNumber="4">36</figureCitation>
). Metarostrum without well marked lateral grooves. Three last antennomeres slightly flattened, forming a club (
<figureCitation id="7EDD2A3BFF997865A4B1FC383A98B8ED" box="[159,252,992,1018]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Figs 23</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A524FC383B4FB8ED" box="[266,299,992,1018]" captionStart="FIGURES 35 47" captionStartId="16.[151,273,1905,1929]" captionTargetBox="[205,1382,227,1867]" captionTargetId="figure@16.[189,1404,194,1887]" captionTargetPageId="16" captionText="FIGURES 35 47. Hoplopisthius (Carcinopisthius) fruhstorferi (Kolbe, 1892). 35, Head (Ψ), dorsal; 36, Head (Ψ), venter; 37, Antenna; 38, Metatarsi and apex of metatibia; 39, Apex of elytra (Ψ); 40, Apex of elytra (♂); 41, Abdomen (♂), venter; 42, Tergite VII (Ψ); 43, Spiculum gastrale; 44, Tegmen; 45, Apex of gonopods IX; 46, Spermatheca; 47, Apex of tergite VIII (Ψ). Scale: 1 mm except fig. 45: 0.17 mm and fig. 46 47: 0.33 mm." httpUri="https://zenodo.org/record/196138/files/figure.png" pageId="3" pageNumber="4">37</figureCitation>
), 11 acute apically. Pronotum pyriform, much broader at base than at apex, with a longitudinal median groove, glabrous. Prothorax without median fovea in front of procoxae. Femora flattened; protibia very short, squat (
<figureCitation id="7EDD2A3BFF997865A544FBF23BA2BF53" box="[362,454,1066,1092]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Figs 24</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A5FCFBF23B96BF53" box="[466,498,1066,1092]" captionStart="FIGURES 48 56" captionStartId="17.[151,270,1576,1600]" captionTargetBox="[178,1412,210,1496]" captionTargetId="figure@17.[151,1436,194,1554]" captionTargetPageId="17" captionText="FIGURES 48 56. Hoplopisthius (Carcinopisthius) oberthueri Senna, 1893. 48, Apex of elytra (Ψ); 49, Apex of elytra (♂); 50, Foreleg; 51, Metatarsi and apex of metatibia; 52, Tergite VII (Ψ); 53, Abdomen (♂), venter; 54, Spermatheca; 55, Apex of epipleurites VIII (Ψ); 56, Apex of tergite VIII (Ψ). Scale: 1 mm except fig. 54 56: 0.33 mm." httpUri="https://zenodo.org/record/196139/files/figure.png" pageId="3" pageNumber="4">50</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A5D1FBF2387BBF53" box="[511,543,1066,1092]" captionStart="FIGURES 57 62" captionStartId="18.[151,272,1633,1657]" captionTargetBox="[182,1426,239,1598]" captionTargetId="figure@18.[151,1436,231,1611]" captionTargetPageId="18" captionText="FIGURES 57 62. Hoplopisthius (Pseudotaphroderes) doriae Senna, 1893. 57, Head (♂), dorsal; 58, Abdomen (♂), venter; 59, Apex of elytra (♂); 60, Foreleg; 61, Spiculum gastrale; 62, Tegmen. Figures 63 71. Hoplopisthius (Pseudotaphroderes) forcipitiger (Damoiseau, 1987). 63, Head (♂), dorsal; 64, Lateral view of pronotum (♂); 65, Abdomen (♂), venter; 66, Apex of elytra (Ψ); 67 68, Apex of elytra (♂); 69, Tergite VIII (Ψ); 70, Spermatheca; 71, Apex of gonopods IX. Scale: 1 mm, except fig. 69 70: 0.33 mm and fig. 71: 0.17 mm." httpUri="https://zenodo.org/record/196140/files/figure.png" pageId="3" pageNumber="4">60</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A605FBF2382FBF53" box="[555,587,1066,1092]" captionStart="FIGURES 72 78" captionStartId="19.[151,269,1899,1923]" captionTargetBox="[181,1417,204,1875]" captionTargetId="figure@19.[165,1431,194,1890]" captionTargetPageId="19" captionText="FIGURES 72 78. Hoplopisthius (Pseudotaphroderes) kolbei Senna, 1893. 72, Head (Ψ), dorsal; 73, Apex of elytra (Ψ); 74, Apex of elytra (♂); 75, Abdomen (♂), venter; 76, Foreleg; 77, Apex of gonopods IX; 78, Spermatheca. Figures 79 82. Hoplopisthius (Pseudotaphroderes) ocularis (Damoiseau, 1987). 79, Head (Ψ), dorsal; 80, Spermatheca; 81, Apex of elytra (Ψ); 82, Abdomen (Ψ), venter. Scale: 1 mm, except fig. 77: 0.17 mm, fig. 78: 0.33 mm and fig. 80: 0.5 mm." httpUri="https://zenodo.org/record/196141/files/figure.png" pageId="3" pageNumber="4">76</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A676FBF2381CBF53" box="[600,632,1066,1092]" captionStart="FIGURES 83 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="3" pageNumber="4">86</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A6AAFBF238C1BF53" box="[644,677,1066,1092]" captionStart="FIGURES 91 97" captionStartId="21.[151,269,931,955]" captionTargetBox="[163,1415,209,888]" captionTargetId="figure@21.[151,1436,197,907]" captionTargetPageId="21" captionText="FIGURES 91 97. Hoplopisthius (Pseudotaphroderes) loriae (Senna &amp; Calabresi, 1919). 91, Head (Ψ), dorsal; 92, Apex of elytra (Ψ); 93, Apex of elytra (♂); 94, Abdomen (Ψ), venter; 95, Foreleg; 96, Tergite VIII (Ψ); 97, Spermatheca. Scale: 1 mm." httpUri="https://zenodo.org/record/196143/files/figure.png" pageId="3" pageNumber="4">95</figureCitation>
). Elytra glabrous. Elytral striae well marked; apex of elytra with caudal processes, never regularly rounded.
<collectingCountry id="9EF1762EFF997865A6BAFB8838ABBF7D" box="[660,719,1104,1130]" name="India" pageId="3" pageNumber="4">Hind</collectingCountry>
wings (
<figureCitation id="7EDD2A3BFF997865A703FB8839E0BF7D" box="[813,900,1104,1130]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Fig. 34</figureCitation>
) with a well developed mediocubital cell and only one primary anal vein; anal lobe large. Metasternum with a longitudinal median groove and usually not laterally carinate.
<collectingCountry id="9EF1762EFF997865A547FB423BC1BFA3" box="[361,421,1178,1204]" name="Malta" pageId="3" pageNumber="4">Male</collectingCountry>
genitalia with parameres reduced, filiform, not fused together (
<figureCitation id="7EDD2A3BFF997865A0A5FB423E86BFA3" box="[1163,1250,1178,1204]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Fig. 33</figureCitation>
,
<figureCitation id="7EDD2A3BFF997865A0C3FB423F6BBFA3" box="[1261,1295,1178,1204]" captionStart="FIGURES 35 47" captionStartId="16.[151,273,1905,1929]" captionTargetBox="[205,1382,227,1867]" captionTargetId="figure@16.[189,1404,194,1887]" captionTargetPageId="16" captionText="FIGURES 35 47. Hoplopisthius (Carcinopisthius) fruhstorferi (Kolbe, 1892). 35, Head (Ψ), dorsal; 36, Head (Ψ), venter; 37, Antenna; 38, Metatarsi and apex of metatibia; 39, Apex of elytra (Ψ); 40, Apex of elytra (♂); 41, Abdomen (♂), venter; 42, Tergite VII (Ψ); 43, Spiculum gastrale; 44, Tegmen; 45, Apex of gonopods IX; 46, Spermatheca; 47, Apex of tergite VIII (Ψ). Scale: 1 mm except fig. 45: 0.17 mm and fig. 46 47: 0.33 mm." httpUri="https://zenodo.org/record/196138/files/figure.png" pageId="3" pageNumber="4">44</figureCitation>
). Gonopods IX of female genitalia with lateral and much reduced styli (
<figureCitation id="7EDD2A3BFF997865A773FB1839D2BFCD" box="[861,950,1216,1242]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Fig. 32</figureCitation>
); coxites IX very long and fused in their basal part (
<figureCitation id="7EDD2A3BFF997865A536FB3D3B0BBFE8" box="[280,367,1253,1279]" captionStart="FIGURES 20 34" captionStartId="15.[151,270,1879,1903]" captionTargetBox="[190,1413,219,1850]" captionTargetId="figure@15.[151,1436,194,1858]" captionTargetPageId="15" captionText="FIGURES 20 34. Hoplopisthius (Hoplopisthius) trichemerus Senna, 1892. 20, Head (♂), dorsal; 21, Head (♂), venter; 22, Head (Ψ), dorsal; 23, Antenna; 24, Foreleg; 25, Metatarsi and apex of metatibia; 26, Apex of elytra (♂); 27, Abdomen (♂), venter; 28, Tergite VIII (Ψ); 29, Epipleurites VIII (Ψ); 30, Spermatheca; 31, Gonopods IX; 32, Apex of gonopods IX; 33, Tegmen; 34, Hind wing (1 A: primary anal vein; mcu: mediocubital cell). Scale: fig. 20 27, 31: 1 mm; fig. 28 30: 0.33 mm; fig. 32: 0.17 mm; fig. 33: 0.5 mm; fig. 34: 2 mm." httpUri="https://zenodo.org/record/196137/files/figure.png" pageId="3" pageNumber="4">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AEFC6535FF997865A4EBFAD238F6BE5D" pageId="3" pageNumber="4" type="discussion">
<paragraph id="E65936BEFF997865A4EBFAD238F6BE5D" blockId="3.[151,1437,879,1354]" pageId="3" pageNumber="4">
<emphasis id="D492EAACFF997865A4EBFAD23B59BE33" bold="true" box="[197,317,1290,1316]" pageId="3" pageNumber="4">Remarks</emphasis>
. All characters given in the diagnostic parts of the genus
<taxonomicName id="21E64D3DFF997865A032FAD33EA8BE33" box="[1052,1228,1291,1316]" class="Insecta" family="Brentidae" genus="Hoplopisthius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="D492EAACFF997865A032FAD33EA8BE33" box="[1052,1228,1291,1316]" italics="true" pageId="3" pageNumber="4">Hoplopisthius</emphasis>
</taxonomicName>
and of the three subgenera are not restated for each species.
</paragraph>
</subSubSection>
</treatment>
</document>