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<document id="9720BC89184751A90F2B4518DFE23FFB" ID-DOI="10.5281/zenodo.196132" ID-GBIF-Dataset="d3d9679d-811d-4860-92e6-c9127f71e9f4" ID-ISSN="1175-5326" ID-Zenodo-Dep="196132" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460239415135" checkinUser="plazi" docAuthor="Mantilleri, Antoine" docDate="2010" docId="6E4F87A8FF947868A4B9FC503E98BC84" docLanguage="en" docName="zt02516p026.pdf" docOrigin="Zootaxa 2516" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hoplopisthius (Pseudotaphroderes) maximus Mantilleri, 2010, n. sp." docType="treatment" docVersion="7" lastPageNumber="15" masterDocId="9276FFD0FF9A7866A42EFFD83A64BB17" masterDocTitle="A taxonomic review and phylogeny of the genera Hoplopisthius Senna, 1892 and Carcinopisthius Kolbe, 1892 (Coleoptera: Brentidae, Hoplopisthiini)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="15" updateTime="1698240316480" updateUser="plazi">
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<mods:title id="DC3F065E7EE2B249077F2017BF18CB25">A taxonomic review and phylogeny of the genera Hoplopisthius Senna, 1892 and Carcinopisthius Kolbe, 1892 (Coleoptera: Brentidae, Hoplopisthiini)</mods:title>
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<taxonomicName id="21E64D3DFF947868A4B9FC5038DCB8B5" ID-CoL="3MMQP" box="[151,696,904,930]" class="Insecta" family="Brentidae" genus="Hoplopisthius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maximus" status="sp. nov." subGenus="Pseudotaphroderes">
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(
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) 
<emphasis id="D492EAACFF947868A667FC5038DCB8B5" bold="true" box="[585,696,904,930]" italics="true" pageId="14" pageNumber="15">maximus</emphasis>
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<typeStatus id="395D881CFF947868A4B9FC083B69B8FD" box="[151,269,976,1002]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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Ψ, &quot;Mailu, Brit. N. G., 
<date id="9258107EFF947868A603FC0838C1B8FD" box="[557,677,976,1002]" pageId="14" pageNumber="15" value="1895-07">July 1895</date>
, Anthony, Muséum Paris, 1952, coll. R. Oberthür, Prép. micro. n°AM00076, 
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<emphasis id="D492EAACFF947868A516FC2E3BB8BF18" box="[312,476,1014,1039]" italics="true" pageId="14" pageNumber="15">Hoplopisthius</emphasis>
(
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) 
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, 
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, 
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1151&quot;, 
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1 Ψ, Nouv. 
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, Baie de Geelvink, Raffray &amp; Maindron 1878, 
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1152 (
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); 1 Ψ, New 
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Pr., N. 
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, Lelet Plat., 
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SE Kamiraba, 
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, 
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– 
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, 
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, leg A. Weigel (
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); 1 ɗ and 1 Ψ, New 
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, Mt Sinewit, 
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, 27.6– 
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, W. W. Brandt (
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).
</paragraph>
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<paragraph id="E65936BEFF947868A4E3FB68389DBFE7" blockId="14.[151,1437,976,1939]" pageId="14" pageNumber="15">
<emphasis id="D492EAACFF947868A4E3FB683B09BFDD" bold="true" box="[205,365,1200,1226]" pageId="14" pageNumber="15">Description.</emphasis>
Total length: 9.1–12.0 mm. Width across humeral calli: 
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. Body red-brown without darker elytral area. Habitus of 
<typeStatus id="395D881CFF947868A670FB0E38F1BFE7" box="[606,661,1238,1264]" pageId="14" pageNumber="15">type</typeStatus>
: 
<figureCitation id="7EDD2A3BFF947868A6B1FB0E3892BFE7" box="[671,758,1238,1264]" captionStart="FIGURES 17 – 19" captionStartId="11.[151,270,1625,1649]" captionTargetBox="[168,1422,211,1584]" captionTargetId="figure@11.[151,1436,194,1603]" captionTargetPageId="11" captionText="FIGURES 17 – 19. Hoplopisthius spp., types. 17, Hoplopisthius maximus n. sp., Ψ holotype; 18, Carcinopisthius loriae Senna &amp; Calabresi, 1919, Ψ lectotype; 19, Hoplopisthius doriae Senna, 1893, ♂ holotype." httpUri="https://zenodo.org/record/196136/files/figure.png" pageId="14" pageNumber="15">Fig. 17</figureCitation>
.
</paragraph>
<paragraph id="E65936BEFF947868A4EBFB233E24BE9F" blockId="14.[151,1437,976,1939]" pageId="14" pageNumber="15">
Head (
<figureCitation id="7EDD2A3BFF947868A536FB233B17BE02" box="[280,371,1275,1301]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 83</figureCitation>
) as long as broad, shiny, hardly punctate. Vertex and frons with a superficial groove. Metarostrum hardly grooved. Prorostrum 0.30–0.35 as long as head + metarostrum + mesorostrum. Antennomere 3 hardly conical, slightly longer than broad; antennomeres 4–8 subcylindrical, as long as broad; 11th 1.75–2.00 longer than 10th. Venter of head with a very weak median groove.
</paragraph>
<paragraph id="E65936BEFF947868A4EBFA4B381ABD54" blockId="14.[151,1437,976,1939]" pageId="14" pageNumber="15">
Pronotum shiny, smooth. Prosternellum distinct. Underside of first protarsal segment denticulate (
<figureCitation id="7EDD2A3BFF947868A119FA4B3FEBBEBA" box="[1335,1423,1427,1453]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 86</figureCitation>
). Metasternum smooth, convex, shiny, not deeply grooved. Interstria 3 abruptly stopped before apical declivity but not forming a sharp tooth (
<figureCitation id="7EDD2A3BFF947868A62DFA063838BEEF" box="[515,604,1502,1528]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 84</figureCitation>
); interstria 8 only present on the rear part of elytra. External border of apical &quot;forceps&quot; without small tooth at base; inner side of apical &quot;forceps&quot; with a short median tooth in female (
<figureCitation id="7EDD2A3BFF947868A4B1F9F13A92BD54" box="[159,246,1577,1603]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 84</figureCitation>
), the tooth much weaker in male.
</paragraph>
<paragraph id="E65936BEFF947868A4EBF9963E51BDA4" blockId="14.[151,1437,976,1939]" pageId="14" pageNumber="15">
Sternites III–IV of female (
<figureCitation id="7EDD2A3BFF947868A629F9963838BD7F" box="[519,604,1614,1640]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 85</figureCitation>
) very convex, shiny, not grooved nor depressed. Tergite VII of female (
<figureCitation id="7EDD2A3BFF947868A48EF9AB3A9ABD9A" box="[160,254,1651,1677]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 88</figureCitation>
) with long hair. Apical teeth of tergite VIII almost missing (
<figureCitation id="7EDD2A3BFF947868A02BF9AB3E02BD9A" box="[1029,1126,1651,1677]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 89</figureCitation>
). Gonopods IX: 
<figureCitation id="7EDD2A3BFF947868A114F9AB3FFCBD9A" box="[1338,1432,1651,1677]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 87</figureCitation>
. Spermatheca: 
<figureCitation id="7EDD2A3BFF947868A513F9413BF0BDA4" box="[317,404,1689,1715]" captionStart="FIGURES 83 – 90" captionStartId="20.[151,270,1566,1590]" captionTargetBox="[171,1390,219,1488]" captionTargetId="figure@20.[151,1436,194,1545]" captionTargetPageId="20" captionText="FIGURES 83 – 90. Hoplopisthius (Pseudotaphroderes) maximus n. sp. 83, Head (Ψ), dorsal; 84, Apex of elytra (Ψ); 85, Abdomen (Ψ), venter; 86, Foreleg; 87, Apex of gonopods IX; 88, Tergite VII (Ψ); 89, Tergite VIII (Ψ); 90, Spermatheca. Scale: 1 mm, except fig. 87: 0.17 mm." httpUri="https://zenodo.org/record/196142/files/figure.png" pageId="14" pageNumber="15">Fig. 90</figureCitation>
. Sternites III–IV of male convex, longitudinally grooved.
</paragraph>
</subSubSection>
<subSubSection id="AEFC6535FF947868A4EBF9653802BDEA" pageId="14" pageNumber="15" type="distribution">
<paragraph id="E65936BEFF947868A4EBF9653802BDEA" blockId="14.[151,1437,976,1939]" pageId="14" pageNumber="15">
<emphasis id="D492EAACFF947868A4EBF9653B0CBDC0" bold="true" box="[197,360,1725,1751]" pageId="14" pageNumber="15">Distribution</emphasis>
. This species is presently known only from New 
<collectingCountry id="9EF1762EFF947868A7F2F9663E52BDCF" box="[988,1078,1726,1752]" name="Guinea-Bissau" pageId="14" pageNumber="15">Guinea</collectingCountry>
(
<collectingCountry id="9EF1762EFF947868A069F9663EA7BDCF" box="[1095,1219,1726,1752]" name="Indonesia" pageId="14" pageNumber="15">Indonesia</collectingCountry>
and Papua-New- 
<collectingCountry id="9EF1762EFF947868A4B9F93B3A94BDEA" box="[151,240,1763,1789]" name="Guinea-Bissau" pageId="14" pageNumber="15">Guinea</collectingCountry>
), New 
<collectingCountry id="9EF1762EFF947868A56DF93B3BF2BDEA" box="[323,406,1763,1789]" name="Ireland" pageId="14" pageNumber="15">Ireland</collectingCountry>
and New 
<collectingCountry id="9EF1762EFF947868A623F93B3806BDEA" box="[525,610,1763,1789]" name="United Kingdom" pageId="14" pageNumber="15">Britain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="AEFC6535FF947868A4EBF8D03EA8BC34" box="[197,1228,1800,1827]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="E65936BEFF947868A4EBF8D03EA8BC34" blockId="14.[151,1437,976,1939]" box="[197,1228,1800,1827]" pageId="14" pageNumber="15">
<emphasis id="D492EAACFF947868A4EBF8D03B2EBC35" bold="true" box="[197,330,1800,1826]" pageId="14" pageNumber="15">Etymology</emphasis>
. The epithet &quot; 
<taxonomicName id="21E64D3DFF947868A5DFF8D1383DBC35" box="[497,601,1801,1826]" class="Insecta" family="Brentidae" genus="Hoplopisthius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="D492EAACFF947868A5DFF8D1383DBC35" box="[497,601,1801,1826]" italics="true" pageId="14" pageNumber="15">maximus</emphasis>
</taxonomicName>
&quot; refers to the large size of individuals of this species.
</paragraph>
</subSubSection>
<subSubSection id="AEFC6535FF947868A4EBF8F53E98BC84" pageId="14" pageNumber="15" type="discussion">
<paragraph id="E65936BEFF947868A4EBF8F53E98BC84" blockId="14.[151,1437,976,1939]" pageId="14" pageNumber="15">
<emphasis id="D492EAACFF947868A4EBF8F53B5BBC50" bold="true" box="[197,319,1837,1863]" pageId="14" pageNumber="15">Remarks</emphasis>
. Both the 
<typeStatus id="395D881CFF947868A5E9F8F63852BC5F" box="[455,566,1838,1864]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
and the female 
<typeStatus id="395D881CFF947868A729F8F63911BC5F" box="[775,885,1838,1864]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
in the MNHN collection were erroneously identified. Damoiseau thought the 
<typeStatus id="395D881CFF947868A604F88B38F4BC7A" box="[554,656,1875,1901]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
belonged to 
<taxonomicName id="21E64D3DFF947868A709F88B3F76BC7A" authority="Senna &amp; Calabresi" authorityName="Senna &amp; Calabresi" box="[807,1298,1875,1901]" class="Insecta" family="Brentidae" genus="Carcinopisthius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="loriae">
<emphasis id="D492EAACFF947868A709F88B3E4ABC7B" box="[807,1070,1875,1900]" italics="true" pageId="14" pageNumber="15">Carcinopisthius loriae</emphasis>
Senna &amp; Calabresi
</taxonomicName>
, and Kleine identified the female 
<typeStatus id="395D881CFF947868A5BFF8A13B92BC84" box="[401,502,1913,1939]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
as 
<taxonomicName id="21E64D3DFF947868A632F8A139DBBC84" authority="Senna" authorityName="Senna" box="[540,959,1913,1939]" class="Insecta" family="Brentidae" genus="Carcinopisthius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="oberthueri">
<emphasis id="D492EAACFF947868A632F8A1393DBC85" box="[540,857,1913,1938]" italics="true" pageId="14" pageNumber="15">Carcinopisthius oberthueri</emphasis>
(Senna)
</taxonomicName>
. Distribution map: 
<figureCitation id="7EDD2A3BFF947868A08FF8A13E9CBC84" box="[1185,1272,1913,1939]" captionStart="FIGURE 99" captionStartId="23.[151,255,1402,1426]" captionTargetBox="[151,1424,530,1375]" captionTargetId="figure@23.[151,1436,530,1381]" captionTargetPageId="23" captionText="FIGURE 99. Distribution map of Hoplopisthius spp." httpUri="https://zenodo.org/record/196144/files/figure.png" pageId="14" pageNumber="15">Fig. 99</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>