161 lines
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161 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.727.20222" ID-GBIF-Dataset="13853cb6-522e-4f94-b633-c4a51ba63624" ID-PMC="PMC5799789" ID-Pensoft-Pub="1313-2970-727-1" ID-PubMed="29416388" ID-ZBK="12B663F7190040788E1EEF8BAC4DF81B" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-727-1" ModsDocOrigin="ZooKeys 727" ModsDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" checkinTime="1515696595036" checkinUser="pensoft" docAuthor="Wood, Hannah M. & Scharff, Nikolaj" docDate="2017" docId="630F0DA193D8E55AC03067423AA9AC07" docLanguage="en" docName="ZooKeys 727: 1-96" docOrigin="ZooKeys 727" docSource="http://dx.doi.org/10.3897/zookeys.727.20222" docTitle="Madagascarchaea rabesahala Wood & Scharff, 2017, sp. n." docType="treatment" docUuid="90FD4DB2-9E20-448F-BB80-1B614B975321" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFEBD959ED2FFFAA9450FF8B1957B348" masterDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="1" updateTime="1668165276717" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wood, Hannah M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Scharff, Nikolaj</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>727</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>96</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.727.20222</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.727.20222</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-727-1</mods:identifier>
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<mods:identifier type="ZBK">12B663F7190040788E1EEF8BAC4DF81B</mods:identifier>
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<mods:identifier type="ZooBank">12B663F7190040788E1EEF8BAC4DF81B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="140245906" LSID="urn:lsid:zoobank.org:act:90FD4DB2-9E20-448F-BB80-1B614B975321" httpUri="http://treatment.plazi.org/id/630F0DA193D8E55AC03067423AA9AC07" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/90FD4DB2-9E20-448F-BB80-1B614B975321" class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea rabesahala" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rabesahala">Madagascarchaea rabesahala</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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Figs 25, 33
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Male holotype: MADAGASCAR, Antananarivo, 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-18.473333">18°28'24"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="47.96">47°57'36"E</geoCoordinate>
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, 1300 m, 5-13 Dec 2000, montane rainforest, beating and sweeping, Fisher, Griswold et al. (deposited in CAS: CASENT9004011).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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MADAGASCAR: Female paratype, same data as holotype except beating low vegetation (CASENT9003843); 1M, together with the holotype (CASENT9004011); 3M,3F,1juv, same data as holotype except general collecting (CASENT9004086); 1F,1juv,1 eggcase, Toamasina, Station Forestier Analamazaotra, administered by Mitsinjo, 0.75 km N Andasibe,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-18.929716">18°55.783'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.4116">48°24.696'E</geoCoordinate>
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, 964m, 2 Feb 2009, primary montane rainforest, sifting litter around logs, dead fern fronds, and at base of
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<normalizedToken originalValue="traveler’s">traveler's</normalizedToken>
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palm, H. Wood (USNMENT01377200).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specific name is a noun in apposition and commemorates
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<normalizedToken originalValue="Gisèle">Gisele</normalizedToken>
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Rabesahala, a Malagasy activist and politician.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Distinguished from all other archaeids, except
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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,
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<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
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,
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<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
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sp. n., and
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<taxonomicName lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
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(presumably, because the male is unknown) by having a retrolateral apophysis on the distal side of the male pedipalpal femur and patella (see fig. 12B in
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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), and because the FSGP is highly reduced and lacks
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<normalizedToken originalValue="“wings”">"wings"</normalizedToken>
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(Fig. 25B).
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<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
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is distinguished from
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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by having the cephalic spines not on protrusions (Fig. 24A), rather than the protrusions seen in
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<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
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(see fig. 18C in
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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); from
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<taxonomicName lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
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by having less than 6 teeth on the cheliceral retromargin, by having a rounded bulge that the AME rests upon, and by lacking a bulge on the posterio-basal side of the chelicerae (see fig. 18D in
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<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
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); from
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<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
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sp. n. by lacking a heavily sclerotized rod-shaped sclerite on the palpal bulb (Fig. 23H,J, arrow), and by lacking a retrolateral cymbial protrusion (Fig. 25C, arrow); and from
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<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
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by having a cephalon that is not as perfectly rounded (compare cephalon shape in Figs 24A, 25A), and by having a larger and more sclerotized triangular basal portion of the conductor (compare basal
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<normalizedToken originalValue="“c”">"c"</normalizedToken>
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in Figs 24G, 25G).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Male holotype (CASENT9004011, from forest close to Andranomay, Madagascar). Total length 1.66, carapace 0.73 long, 0.61 wide. Abdomen 0.90 long, 1.07 high. Carapace tilt angle 62.4°, tilt height (CtH) 1.42, constriction 0.39, head length 0.63, neck length 0.71. CtH divided by carapace length 1.95. Cephalon with AME on large bulge, and with 6 short post-occular spines at the crown, not on protrusions, and lacking spine between the LE and AME. Chelicerae 1.44 long, and with a small spine 0.33 from base of chelicerae and projecting downward (Fig. 25A). Femur I 2.01 long. Sternum 0.46 long, 0.30 wide. Carapace, chelicerae, sternum and legs reddish brown with white setae. Legs with darker annulations on tibiae and metatarsi. Abdomen dark brown with lighter circular patches throughout, with white setae (Fig. 25A). Pedipalpal bulb of the "
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<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
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group" form (Fig. 25
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<normalizedToken originalValue="D–I">D-I</normalizedToken>
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): pedipalpal bulbs elongated with the conductor swirling around a mostly membraneous embolus; conductor base triangular (Fig. 25D), with remainder of conductor elongate and cradling embolus, and with a curved tip; MA translucent and fans out (Fig. 25
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<normalizedToken originalValue="E–F,H–I">E-F,H-I</normalizedToken>
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). Cymbium lacks retrolateral protrusion (Fig. 25C, arrow).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Female paratype (CASENT903843). Total length 1.91, carapace 0.80 long, 0.67 wide. Abdomen 1.11 long, 1.55 high. Carapace tilt angle 62.3°, tilt height (CtH) 1.55, constriction 0.42, head length 0.78, neck length 0.82. CtH divided by carapace length 1.94. Cephalon as in male, except spine between LE and AME present. Chelicerae 1.61 long, and with a small spine 0.34 from base of chelicerae and projecting downward (Fig. 25A). Femur I 1.97 long. Sternum 0.49 long, 0.32 wide. Colors as in male. Genitalia of the "
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<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
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-group" form (Fig. 25B): with one group of poreplates on each side of the bursa anterior; with FSGP highly reduced, lacking
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<normalizedToken originalValue="“wings.”">"wings."</normalizedToken>
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 25.
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<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea rabesahala" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rabesahala">Madagascarchaea rabesahala</taxonomicName>
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sp. n. A male (holotype, CASENT9004011) habitus, lateral view, image reversed B female (CASENT9004086) internal genitalia, dorsal view
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<normalizedToken originalValue="C–I">C-I</normalizedToken>
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male pedipalpal bulbs
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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right bulb (CASENT9004086), image reversed C,
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<normalizedToken originalValue="G–I">G-I</normalizedToken>
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left bulb (CASENT9004086): C cymbium close-up, dorsal view, arrow showing the absence of a retrolateral cymbium protrusion D, G prolateral view E, H ventral view F, I retrolateral view. Scale bars: 1 mm (A); 0.25 mm (B, D).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">Total length 1.56-1.66 (males; n=5), 1.91-2.07 (females; n=4); Carapace length 0.72-0.75 (males; n=5), 0.74-0.81 (females; n=4); Femur I 2.69-2.86 times the length of carapace in males (n=5), 2.38-2.69 in females (n=4); CtH divided by carapace length 1.93-2.12 in males (n=5), 1.94-2.15 in females (n=4); Average femur I length 2.03 in males (n=5), 1.96 in females (n= 4). The spine between the LE and AME is present is 3 males out of 5, and 4 females out of 4.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="natural history">
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<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
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<paragraph pageId="0" pageNumber="1">Specimens were collected in montane rainforest from 960-1300 m in elevation by beating low vegetation, by sifting litter, by beating and sweeping, and by general collecting.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known only from central eastern Madagascar (Fig. 33).</paragraph>
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</subSubSection>
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</treatment>
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</document> |