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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZBK="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel &amp; Njunjic, Iva" docDate="2018" docId="1674009F540A6D2E2BF7516D116EFF89" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaminus hanskii Schilthuizen &amp; Perreau, sp. n." docType="treatment" docUuid="375C731A-B3B0-49DF-A150-51FA4F0D7A41" docUuidSource="ZooBank" docVersion="5" lastPageNumber="89" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="88" updateTime="1668166102793" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Njunjic, Iva</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>777</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>108</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.777.23212</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.777.23212</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-777-57</mods:identifier>
<mods:identifier type="ZBK">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
<mods:identifier type="ZooBank">D9F353643DCD4BA6B70D62FB275DEB1B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="146746187" LSID="urn:lsid:zoobank.org:act:375C731A-B3B0-49DF-A150-51FA4F0D7A41" httpUri="http://treatment.plazi.org/id/1674009F540A6D2E2BF7516D116EFF89" lastPageId="32" lastPageNumber="89" pageId="31" pageNumber="88">
<subSubSection pageId="31" pageNumber="88" type="nomenclature">
<paragraph pageId="31" pageNumber="88">
<taxonomicName LSID="http://zoobank.org/375C731A-B3B0-49DF-A150-51FA4F0D7A41" authority="Schilthuizen &amp; Perreau" class="Insecta" family="Leiodidae" genus="Ptomaphaminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaminus hanskii" order="Coleoptera" pageId="31" pageNumber="88" phylum="Arthropoda" rank="species" species="hanskii">
<pageBreakToken pageId="31" pageNumber="88" start="start">Ptomaphaminus</pageBreakToken>
hanskii Schilthuizen &amp; Perreau
</taxonomicName>
<taxonomicNameLabel pageId="31" pageNumber="88">sp. n.</taxonomicNameLabel>
Figure 11a, 12k, l, 13i, j
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="89" type="materials_examined">
<paragraph pageId="32" pageNumber="89">
<pageBreakToken pageId="32" pageNumber="89" start="start">Material</pageBreakToken>
.
</paragraph>
<paragraph pageId="32" pageNumber="89">
Holotype: Malaysia, Sarawak, 4th Division, Gunung Mulu National Park, Slope, 4.5.78, 9.1, 9.3,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, in NHMUK, B.M.1978-49), 1 male. Paratypes: Sarawak. Gunung Mulu National Park, Slope, 4.5.78, 9.1, 9.3,
<normalizedToken originalValue="vviii">v-viii</normalizedToken>
.1978 (leg. P.M. Hammond &amp; J.E. Marshall, NHMUK, B.M.1978-49), 5 males, 8 females.
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="89" type="description">
<paragraph pageId="32" pageNumber="89">Description.</paragraph>
<paragraph pageId="32" pageNumber="89">Length: 1.6-2.1 mm. General colour brown; the tarsi and two first antennomeres yellowish. Winged. Eyes well developed. Pronotum 1.65 times as wide as long. Elytra 1.30 times as long as wide. Sutural angle of female elytra angular. Apex of the spiculum gastrale of the male genital segment dilated into a diamond-like form (Figure 13i). Aedeagus elongated, approximately 3.5 times shorter than the body length. Median lobe regularly narrowed from base to the apex in dorsal view (Figure 12l), thick and strongly arcuate, with a long ventrally deflexed but not retroverted apical hook in lateral view (Figure 12k). Six lateroventral preapical setae and one lateroapical seta on each side. Internal stylus of the endophallus moderately sinuate. Female genital segment with reduced gonocoxites. Spermiduct membranous, vaguely helical at least near the base of the spermatheca. Apex of the spermatheca rounded, without apical sclerotised plate (Figure 13j).</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="89" type="differential diagnosis">
<paragraph pageId="32" pageNumber="89">Differential diagnosis.</paragraph>
<paragraph pageId="32" pageNumber="89">
Species with normally developed eyes and a long aedeagus. Very similar to
<taxonomicName lsidName="P. sarawacensis" pageId="32" pageNumber="89" rank="species" species="sarawacensis">P. sarawacensis</taxonomicName>
, from which it cannot be confidently distinguished in the female sex. Males of
<taxonomicName lsidName="P. hanskii" pageId="32" pageNumber="89" rank="species" species="hanskii">P. hanskii</taxonomicName>
have a thicker median lobe of the aedeagus than
<taxonomicName lsidName="P. sarawacensis" pageId="32" pageNumber="89" rank="species" species="sarawacensis">P. sarawacensis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="89" type="habitat">
<paragraph pageId="32" pageNumber="89">Habitat and distribution.</paragraph>
<paragraph pageId="32" pageNumber="89">Known from lowland forests of Gunung Mulu, Sarawak, Malaysia.</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="89" type="remarks">
<paragraph pageId="32" pageNumber="89">Remarks.</paragraph>
<paragraph pageId="32" pageNumber="89">
The external morphology and the female genital morphology are extremely similar to
<taxonomicName lsidName="P. sarawacensis" pageId="32" pageNumber="89" rank="species" species="sarawacensis">P. sarawacensis</taxonomicName>
, so that females are nearly impossible to distinguish. However, the male aedeagi of these species are very different.
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="89" type="etymology">
<paragraph pageId="32" pageNumber="89">Etymology.</paragraph>
<paragraph pageId="32" pageNumber="89">Named in honour of Ilkka Hanski, the Finnish ecologist who played an important role in the Royal Geographical Society expedition to Mulu of 1978, and who passed away in 2016.</paragraph>
</subSubSection>
</treatment>
</document>