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<document id="472AD5AD971257B58036BCB0DC81BAF6" ID-DOI="10.5281/zenodo.10415212" ID-ISSN="0253-116X" ID-Zenodo-Dep="10415212" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703112367482" checkinUser="felipe" docAuthor="Wood, Thomas" docDate="2023" docId="165887FC0356ED4232EBFF2DC958FC69" docLanguage="en" docName="LinzerbiolBeitr.55.1.355-371.pdf" docOrigin="Linzer biologische Beiträge 55 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.10:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="10" docTitle="Andrena fumea Wood 2023, nov.sp." docType="treatment" docVersion="2" lastPageNumber="366" masterDocId="EA61FF84035FED493259FFF7CA32FF9D" masterDocTitle="New Andrena species from Central Asia (Hymenoptera, Andrenidae)" masterLastPageNumber="371" masterPageNumber="355" pageNumber="364" updateTime="1703114625571" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F2C2B110F2A70D68851507094995016C">New Andrena species from Central Asia (Hymenoptera, Andrenidae)</mods:title>
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<mods:namePart id="E5F2EBE644511F72637E57CB9C29FB85">Wood, Thomas</mods:namePart>
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<mods:date id="44ED7F4E1855640486D602E9B30898CE">2023</mods:date>
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<paragraph id="9E4E36EA0356ED4032EBFF2DC8E4FF6D" blockId="9.[178,1086,218,322]" box="[178,726,218,240]" pageId="9" pageNumber="364">
<heading id="C50681860356ED4032EBFF2DC8E4FF6D" box="[178,726,218,240]" fontSize="8" level="2" pageId="9" pageNumber="364" reason="2">
<emphasis id="AC85EAF80356ED4032EBFF2DC8E4FF6D" bold="true" box="[178,726,218,240]" pageId="9" pageNumber="364">
<taxonomicName id="59F14D690356ED4032EBFF2DCBCCFF6D" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[178,510,218,240]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="fumea" status="sp. nov.">
<emphasis id="AC85EAF80356ED4032EBFF2DCB3FFF6D" bold="true" box="[178,269,218,240]" italics="true" pageId="9" pageNumber="364">Andrena</emphasis>
(incertae sedis)
<emphasis id="AC85EAF80356ED4033E5FF2DCBCCFF6D" bold="true" box="[444,510,218,240]" italics="true" pageId="9" pageNumber="364">fumea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B657830356ED40305CFF2CC863FF6D" box="[517,593,219,240]" pageId="9" pageNumber="364" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="06CA2A6F0356ED403007FF2DC8FDFF6D" box="[606,719,218,240]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figs 21-26</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="9E4E36EA0356ED4032EBFF0BCB3CFEDF" blockId="9.[178,1086,218,322]" pageId="9" pageNumber="364">
M a t e r i a l e x a m i n e d:
<typeStatus id="414A88480356ED40305EFF0BC85AFE8F" box="[519,616,252,274]" pageId="9" pageNumber="364" type="holotype">Holotypes</typeStatus>
:
<collectingCountry id="E6E6767A0356ED403028FF0BC8EBFE8C" box="[625,729,252,273]" name="Kyrgyzstan" pageId="9" pageNumber="364">Kyrgyzstan</collectingCountry>
: sept. Čon Aryk [Chong-Aryk] (Frunze) [
<collectingRegion id="5C35F8080356ED4032BDFEE3CB00FEB7" box="[228,306,276,298]" country="Kyrgyzstan" name="Bishkek Shaary" pageId="9" pageNumber="364">Bishkek</collectingRegion>
], cca
<quantity id="59099B0F0356ED403306FEE3CB90FEB4" box="[351,418,276,298]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="364" unit="m" value="1050.0">1050 m</quantity>
, 42.8119
<superScript id="69849BA20356ED4033ABFEE6C838FEB4" attach="right" box="[498,522,273,297]" fontSize="6" pageId="9" pageNumber="364">oN</superScript>
, 74.5844
<superScript id="69849BA20356ED403000FEE6C85DFEB4" attach="right" box="[601,623,273,297]" fontSize="6" pageId="9" pageNumber="364">oE</superScript>
,
<date id="EA4F102A0356ED40302EFEE3C8E3FEB7" box="[631,721,276,298]" pageId="9" pageNumber="364" value="1983-07-05">5.vii.1983</date>
,
<specimenCount id="88F7FD630356ED403080FEE3C8C7FEB4" box="[729,757,276,297]" count="1" pageId="9" pageNumber="364" type="female">1♀</specimenCount>
, leg. B. Tkalců, coll. Biologiezentrum Linz.
</paragraph>
</subSubSection>
<subSubSection id="D6EB65610356ED4232E8FEA7C958FC69" lastPageId="11" lastPageNumber="366" pageId="9" pageNumber="364" type="description">
<paragraph id="9E4E36EA0356ED4032E8FEA7CBE8FD34" blockId="9.[177,1085,336,681]" pageId="9" pageNumber="364">
D i a g n o s i s.
<taxonomicName id="59F14D690356ED403300FEA7CBC7FEF8" authorityName="Wood" authorityYear="2023" box="[345,501,336,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="AC85EAF80356ED403300FEA7CBC7FEF8" box="[345,501,336,357]" italics="true" pageId="9" pageNumber="364">Andrena fumea</emphasis>
</taxonomicName>
is part of an undescribed subgenus best referred to as the
<emphasis id="AC85EAF80356ED4032EBFE9ACADCFE1F" box="[178,238,365,386]" italics="true" pageId="9" pageNumber="364">relata</emphasis>
-group (see
<bibRefCitation id="FA604B1B0356ED403331FE99C870FE1E" author="PISANTY G &amp; RICHTER R &amp; MARTIN T &amp; DETTMAN J" box="[360,578,365,387]" pageId="9" pageNumber="364" pagination="107151" refId="ref9195" refString="PISANTY G., RICHTER R., MARTIN T., DETTMAN J. &amp; S. CARDINAL (2022 b): Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). - Mol. Phylogenet. Evol. 170: 107151." type="journal article" year="2022">PISANTY et al. 2022b</bibRefCitation>
;
<bibRefCitation id="FA604B1B0356ED403014FE99C8EAFE1E" author="WOOD T" box="[589,728,366,387]" pageId="9" pageNumber="364" pagination="241 - 484" refId="ref9587" refString="WOOD T. J. (2023 a): The genus Andrena FABRICIUS, 1775 in the Iberian Peninsula. - J. Hymenopt. Res. 96: 241 - 484." type="journal article" year="2023">WOOD 2023a</bibRefCitation>
;
<bibRefCitation id="FA604B1B0356ED4030BAFE9AC916FE1E" author="WOOD T" box="[739,804,365,387]" pageId="9" pageNumber="364" pagination="1 - 72" refId="ref9621" refString="WOOD T. J. (2023 b): New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). - Zootaxa 5266: 1 - 72." type="journal article" year="2023">2023b</bibRefCitation>
) that does not display clear defining or distinctive characters, leading to its lack of historical recognition. This group is incompletely characterised, but can predominantly be recognised by the relatively short and compact head with the fore margin of the clypeus slightly raised and forming a ] shape (
<figureCitation id="06CA2A6F0356ED4032A3FE14CBBCFE65" box="[250,398,483,505]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figures 22-23</figureCitation>
), by the often weak and poorly defined propodeal triangle (
<figureCitation id="06CA2A6F0356ED4031A0FE14CAFFFD8B" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 25</figureCitation>
, though this is less applicable in the case of
<taxonomicName id="59F14D690356ED4030C9FDF7C8D8FD88" authorityName="Wood" authorityYear="2023" box="[656,746,512,533]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="AC85EAF80356ED4030C9FDF7C8D8FD88" box="[656,746,512,533]" italics="true" pageId="9" pageNumber="364">A. fumea</emphasis>
</taxonomicName>
), by the ovoid metasoma (
<figureCitation id="06CA2A6F0356ED4031A0FDF7CAFCFDAE" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 26</figureCitation>
), and the lack of any distinctive derived characters. It is best diagnosed against the most morphologically similar species which is
<taxonomicName id="59F14D690356ED403003FDCCC945FDCC" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[602,887,571,593]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="melanota">
<emphasis id="AC85EAF80356ED403003FDCCC8E6FDCD" box="[602,724,571,592]" italics="true" pageId="9" pageNumber="364">A. melanota</emphasis>
Warncke, 1975
</taxonomicName>
(
<collectingCountry id="E6E6767A0356ED4031DCFDCCC9E3FDCC" box="[901,977,571,593]" name="Turkey" pageId="9" pageNumber="364">Turkey</collectingCountry>
) due to its similar structure, size, and pubescence (pubescence including scopae predominantly dark with white hairs on the face, mesepisternum, and forming apical white patches laterally on the terga,
<figureCitation id="06CA2A6F0356ED40334FFD64CBFCFD34" box="[278,462,659,681]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figures 21-22; 26</figureCitation>
).
</paragraph>
<paragraph id="9E4E36EA0356ED4032EBFD4FCBCBFB1B" blockId="9.[178,1085,696,1158]" pageId="9" pageNumber="364">
The two species are very close, but
<taxonomicName id="59F14D690356ED40306AFD4FC8A2FD50" authorityName="Wood" authorityYear="2023" box="[563,656,696,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="fumea">
<emphasis id="AC85EAF80356ED40306AFD4FC8A2FD50" box="[563,656,696,717]" italics="true" pageId="9" pageNumber="364">A. fumea</emphasis>
</taxonomicName>
can be separated by the structure of the clypeus which is evenly punctate, without a slightly raised impunctate mid-line (in
<taxonomicName id="59F14D690356ED40367EFD21CB22FC95" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="melanota">
<emphasis id="AC85EAF80356ED40367EFD21CB22FC95" italics="true" pageId="9" pageNumber="364">A. melanota</emphasis>
</taxonomicName>
with a clear and slightly raised impunctate mid-line), by the scutum which is more finely and densely punctate, punctures separated by 0.5-1 puncture diameters, underlying surface finely shagreened and weakly shining (
<figureCitation id="06CA2A6F0356ED40315AFCDAC95BFCDE" box="[771,873,813,835]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 24</figureCitation>
; in
<taxonomicName id="59F14D690356ED4031D7FCD9CE3BFCDF" authorityName="Warncke" authorityYear="1975" box="[910,1033,813,835]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="melanota">
<emphasis id="AC85EAF80356ED4031D7FCD9CE3BFCDF" box="[910,1033,813,835]" italics="true" pageId="9" pageNumber="364">A. melanota</emphasis>
</taxonomicName>
with the scutum more strongly punctate, punctures separated by 1-2 puncture diameters, underlying surface polished and shining), by the structure of the propodeum which is covered with coarse raised rugae over its basal 1/3
<superScript id="69849BA20356ED4030EAFC75C8F2FC0D" attach="left" box="[691,704,898,912]" fontSize="6" pageId="9" pageNumber="364">rd</superScript>
and with fine granular shagreen over the remaining area, thus strongly contrasting both the sculpture of the basal half and the dorsolateral parts of the propodeum (
<figureCitation id="06CA2A6F0356ED40301AFC37C89DFC4B" box="[579,687,960,982]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 25</figureCitation>
; in
<taxonomicName id="59F14D690356ED403087FC37C96CFC48" authorityName="Warncke" authorityYear="1975" box="[734,862,960,981]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="melanota">
<emphasis id="AC85EAF80356ED403087FC37C96CFC48" box="[734,862,960,981]" italics="true" pageId="9" pageNumber="364">A. melanota</emphasis>
</taxonomicName>
the sculpture of the propodeum is more consistent, with fine raised rugae in its basal half, these becoming weaker but the remaining area finely reticulate, not strongly contrasting the dorsolateral parts of the propodeum), and by the pubescence of the terga, with T2-3 only forming weak and sparse apical hair fringes of white hairs that do not obscure the underlying surface (
<figureCitation id="06CA2A6F0356ED4032E0FBA4CB2CFBF4" box="[185,286,1107,1129]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 26</figureCitation>
; in
<taxonomicName id="59F14D690356ED403318FBA4CB88FBF5" authorityName="Warncke" authorityYear="1975" box="[321,442,1107,1128]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="melanota">
<emphasis id="AC85EAF80356ED403318FBA4CB88FBF5" box="[321,442,1107,1128]" italics="true" pageId="9" pageNumber="364">A. melanota</emphasis>
</taxonomicName>
with T2-4 laterally with strong and thick white hair patches that obscure the underlying surface).
</paragraph>
<paragraph id="9E4E36EA0356ED4032EBFB61CB5BFB36" blockId="9.[178,361,1173,1195]" box="[178,361,1173,1195]" pageId="9" pageNumber="364">D e s c r i p t i o n.</paragraph>
<paragraph id="9E4E36EA0356ED4032EBFB4CC9D6F98E" blockId="9.[178,1085,1211,1555]" pageId="9" pageNumber="364">
Female. Body length:
<quantity id="59099B0F0356ED4033CCFB4CCBEEFB4D" box="[405,476,1211,1232]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="364" unit="mm" value="13.0">13 mm</quantity>
(
<figureCitation id="06CA2A6F0356ED4033B3FB4CC863FB4D" box="[490,593,1211,1233]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 21</figureCitation>
). Head: Dark, 1.3 times wider than long (
<figureCitation id="06CA2A6F0356ED4031A0FB4CCAFDFB70" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 22</figureCitation>
). Clypeus very weakly domed, surface very densely punctate, punctures separated by 0.5 puncture diameters, interspaces weakly shining. Fore margin of clypeus distinctly upturned, forming shallowly rounded semicircle (
<figureCitation id="06CA2A6F0356ED4030B9FAE4C960FAB5" box="[736,850,1299,1321]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="9" pageNumber="364">Figure 23</figureCitation>
). Process of labrum trapezoidal, twice as broad as long, apical margin with angular emargination. Gena slightly exceeding diameter of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Foveae occupying slightly more than ½ space between compound eye and lateral ocellus; foveae filled with dark brown hairs. Head with intermixed black and white hairs, black hairs covering majority of gena, vertex, dorsal part of frons, and fore margin of clypeus narrowly; white hairs covering majority of lower part of frons, antennal insertions, paraocular areas, and clypeus. Antennae dark, A5 ventrally weakly lightened by presence of grey-brown scales. A3 slightly exceeding A4+5, clearly shorter than A4+5+6.
</paragraph>
<caption id="CA8E66620355ED4332EBFB15CBC9FAB5" ID-DOI="http://doi.org/10.5281/zenodo.10415223" ID-Zenodo-Dep="10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="10" pageNumber="365" startId="10.[178,218,1250,1272]" targetBox="[181,1090,219,1239]" targetPageId="10" targetType="figure">
<paragraph id="9E4E36EA0355ED4332EBFB15CBC9FAB5" blockId="10.[178,1085,1250,1320]" pageId="10" pageNumber="365">
<emphasis id="AC85EAF80355ED4332EBFB15CB2EFB6A" bold="true" box="[178,284,1250,1272]" pageId="10" pageNumber="365">Figs 21-26:</emphasis>
<taxonomicName id="59F14D690355ED43337BFB15C80EFB6A" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[290,572,1250,1272]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="fumea" status="sp. nov.">
<emphasis id="AC85EAF80355ED43337BFB15CB42FB6A" box="[290,368,1250,1271]" italics="true" pageId="10" pageNumber="365">Andrena</emphasis>
(incertae sedis)
<emphasis id="AC85EAF80355ED43305CFB15C80EFB6A" box="[517,572,1250,1271]" italics="true" pageId="10" pageNumber="365">fumea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B657830355ED43301BFB14C8B7FB65" box="[578,645,1251,1272]" pageId="10" pageNumber="365" rank="species">nov.sp.</taxonomicNameLabel>
female (
<emphasis id="AC85EAF80355ED43308DFB15C8D9FB6A" bold="true" box="[724,747,1250,1271]" pageId="10" pageNumber="365">21</emphasis>
) profile; (
<emphasis id="AC85EAF80355ED433111FB15C96DFB6A" bold="true" box="[840,863,1250,1271]" pageId="10" pageNumber="365">22</emphasis>
) face, frontal view; (
<emphasis id="AC85EAF80355ED433646FB15CE04FB6A" bold="true" box="[1055,1078,1250,1271]" pageId="10" pageNumber="365">23</emphasis>
) process of the labrum, fronto-ventral view; (
<emphasis id="AC85EAF80355ED433011FB0DC86DFA92" bold="true" box="[584,607,1274,1295]" pageId="10" pageNumber="365">24</emphasis>
) scutum, dorsolateral view; (
<emphasis id="AC85EAF80355ED433135FB0DC9B1FA92" bold="true" box="[876,899,1274,1295]" pageId="10" pageNumber="365">25</emphasis>
) propodeal triangle, dorsal view; (
<emphasis id="AC85EAF80355ED433376FAE5CB74FABA" bold="true" box="[303,326,1298,1319]" pageId="10" pageNumber="365">26</emphasis>
) terga, dorsal view.
</paragraph>
</caption>
<paragraph id="9E4E36EA0355ED4232EBFAB8C98FFE46" blockId="10.[178,1085,1359,1586]" lastBlockId="11.[178,1086,219,475]" lastPageId="11" lastPageNumber="366" pageId="10" pageNumber="365">
Mesosoma: Scutum and scutellum finely and densely punctate, punctures separated by 0.5-1 puncture diameters, underlying surface finely shagreened and weakly shining (
<figureCitation id="06CA2A6F0355ED4332E0FA7ECB1DFA02" box="[185,303,1417,1439]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="10" pageNumber="365">Figure 24</figureCitation>
). Pronotum with strong humeral angle. Mesepisternum with fine microreticulation, surface with shallow punctation, punctures separated by 0.5-1 puncture diameters. Dorsolateral parts of propodeum reticulate, forming dense network of raised reticulation. Propodeal triangle broad, in basal 1/3
<superScript id="69849BA20355ED4330ECFA29C8F0FA71" attach="left" box="[693,706,1502,1516]" fontSize="6" pageId="10" pageNumber="365">rd</superScript>
covered with coarse raised rugae, in apical 2/3rds with fine granular shagreen, strongly contrasting dorsolateral parts of propodeum (
<figureCitation id="06CA2A6F0355ED433361F9EBCB96F9AF" box="[312,420,1564,1586]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="10" pageNumber="365">Figure 25</figureCitation>
). Mesepisternum with predominantly white hairs, scutum and scutellum with black hairs; metanotum and propodeum with predominantly white hairs. Propodeal corbicula incomplete, but with small and weak apical fringe composed of black plumose hairs; dorsal fringe strong, composed of long white plumose hairs, internal surface with scattered long white hairs. Legs dark, apical tarsal segments slightly lightened brownish, pubescence predominantly dark brown to black, ventral margin of fore femur with long fringe of white hairs. Flocculus moderately weak, composed of white and black plumose hairs; femoral scopae composed of long white simple hairs, tibial scopae composed of shorter black simple hairs. Hind tibial claws with strong inner tooth. Wings weakly brownish-infumate, stigma and venation orange, nervulus antefurcal.
</paragraph>
<paragraph id="9E4E36EA0354ED4232EBFE1CC888FD2D" blockId="11.[178,1085,491,689]" pageId="11" pageNumber="366">
Metasoma: Tergal discs dark, tergal margins narrowly lightened brownish (
<figureCitation id="06CA2A6F0354ED42319EFE1CCE03FD9C" box="[967,1073,491,513]" captionStart="Figs 21-26" captionStartId="10.[178,218,1250,1272]" captionTargetBox="[181,1090,219,1239]" captionTargetId="figure-138@10.[181,1090,219,1239]" captionTargetPageId="10" captionText="Figs 21-26: Andrena (incertae sedis) fumea nov.sp. female (21) profile; (22) face, frontal view; (23) process of the labrum, fronto-ventral view; (24) scutum, dorsolateral view; (25) propodeal triangle, dorsal view; (26) terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10415223" httpUri="https://zenodo.org/record/10415223/files/figure.png" pageId="11" pageNumber="366">Figure 26</figureCitation>
). Terga with weak sculpture, more or less polished and shining; terga punctate, punctures separated by 1-2 puncture diameters. Discs of T1-2 with sparse white hairs, discs of T3-5 with dense short black hairs; T1-3 laterally with obscure patches of whitish hairs. Apical fringe of T5 medially and hairs flanking pygidial plate dark brown; apical fringe of T5 laterally with white hairs. Pygidial plate triangular with rounded apex, lateral margins weakly raised, medially with raised triangular area.
</paragraph>
<paragraph id="9E4E36EA0354ED4232EBFD37CB6CFD4B" blockId="11.[178,350,704,726]" box="[178,350,704,726]" pageId="11" pageNumber="366">Male. Unknown.</paragraph>
<paragraph id="9E4E36EA0354ED4232EBFD11C850FD66" blockId="11.[178,610,741,763]" box="[178,610,741,763]" pageId="11" pageNumber="366">
D i s t r i b u t i o n: Northern
<collectingCountry id="E6E6767A0354ED4233BFFD11C86CFD66" box="[486,606,742,763]" name="Kyrgyzstan" pageId="11" pageNumber="366">Kyrgyzstan</collectingCountry>
.
</paragraph>
<paragraph id="9E4E36EA0354ED4232EBFCFCCB64FCC6" blockId="11.[178,1086,779,1012]" pageId="11" pageNumber="366">
D e r i v a t i o n o m i n i s: Feminine singular nominative form of the Latin adjective
<emphasis id="AC85EAF80354ED4232EBFCDFCAC8FCA0" box="[178,250,808,829]" italics="true" pageId="11" pageNumber="366">fumeus</emphasis>
meaning smoky, in reference to its dark pubescence, integument, and slightly infumate wings.
</paragraph>
<paragraph id="9E4E36EA0354ED4232EBFC90C958FC69" blockId="11.[178,1086,779,1012]" pageId="11" pageNumber="366">
<materialsCitation id="2E993CB70354ED4232EBFC90C91CFC09" collectingDate="1970-05-28" collectorName="H. Ozbek &amp; Biologiezentrum Linz" country="Turkey" location="Erzurum" pageId="11" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Erzurum" typeStatus="holotype">
O t h e r m a t e r i a l e x a m i n e d: (
<taxonomicName id="59F14D690354ED42306EFC91C9BAFCE1" authority="WARNCKE, 1975" authorityName="Warncke" authorityYear="1975" box="[567,904,870,892]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="366" phylum="Arthropoda" rank="species" species="melanota">
<emphasis id="AC85EAF80354ED42306EFC91C8D3FCE6" box="[567,737,870,891]" italics="true" pageId="11" pageNumber="366">Andrena melanota</emphasis>
<smallCapsWord id="98A8A0360354ED4230B1FC90C978FCE6" baselines="887,887" box="[744,842,871,892]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Warncke" pageId="11" pageNumber="366">WARNCKE</smallCapsWord>
, 1975
</taxonomicName>
):
<collectingCountry id="E6E6767A0354ED4231C0FC91C9ECFCE1" box="[921,990,870,892]" name="Turkey" pageId="11" pageNumber="366">Turkey</collectingCountry>
:
<collectingRegion id="5C35F8080354ED4231B1FC90CE0BFCE1" box="[1000,1081,871,892]" country="Turkey" name="Erzurum" pageId="11" pageNumber="366">Erzurum</collectingRegion>
,
<date id="EA4F102A0354ED423286FC88CB0AFC09" box="[223,312,895,916]" pageId="11" pageNumber="366" value="1970-05-28">
<collectingDate id="FA0BE9C20354ED423286FC88CB0AFC09" box="[223,312,895,916]" pageId="11" pageNumber="366" value="1970-05-28">28.v.1970</collectingDate>
</date>
,
<specimenCount id="88F7FD630354ED423366FC88CB69FC09" box="[319,347,895,916]" count="1" pageId="11" pageNumber="366" type="female">1♀</specimenCount>
, leg.
<collectorName id="3304533C0354ED4233D1FC88CBEFFC09" box="[392,477,894,916]" pageId="11" pageNumber="366">H. Ozbek</collectorName>
, coll.
<collectorName id="3304533C0354ED423057FC89C8FFFC09" box="[526,717,894,916]" pageId="11" pageNumber="366">Biologiezentrum Linz</collectorName>
(
<typeStatus id="414A88480354ED42308EFC89C91AFC09" box="[727,808,894,916]" pageId="11" pageNumber="366" type="holotype">holotype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2E993CB70354ED42316CFC89C8E5FC31" collectingDate="1966-07-26" collectorName="H. Ozbek &amp; Biologiezentrum Linz" country="Turkey" location="Erzurum" pageId="11" pageNumber="366" specimenCount="1" stateProvince="Erzurum" typeStatus="paratype">
<collectingCountry id="E6E6767A0354ED42316CFC89C94AFC09" box="[821,888,894,916]" name="Turkey" pageId="11" pageNumber="366">Turkey</collectingCountry>
:
<collectingRegion id="5C35F8080354ED423126FC88C9FCFC09" box="[895,974,895,916]" country="Turkey" name="Erzurum" pageId="11" pageNumber="366">Erzurum</collectingRegion>
,
<date id="EA4F102A0354ED42318CFC89CE08FC09" box="[981,1082,894,916]" pageId="11" pageNumber="366" value="1966-07-26">
<collectingDate id="FA0BE9C20354ED42318CFC89CE08FC09" box="[981,1082,894,916]" pageId="11" pageNumber="366" value="1966-07-26">26.vii.1966</collectingDate>
</date>
, 1♁, leg.
<collectorName id="3304533C0354ED423372FC60CBB0FC31" box="[299,386,918,940]" pageId="11" pageNumber="366">H. Ozbek</collectorName>
, coll.
<collectorName id="3304533C0354ED4233ECFC61C847FC31" box="[437,629,918,940]" pageId="11" pageNumber="366">Biologiezentrum Linz</collectorName>
(
<typeStatus id="414A88480354ED4230D8FC60C8E3FC31" box="[641,721,919,940]" pageId="11" pageNumber="366" type="paratype">paratype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2E993CB70354ED4230B9FC61C8A0FC59" collectingDate="1937-08-31" collectorName="O. Ecevit &amp; Biologiezentrum Linz" country="Turkey" location="Erzurum" pageId="11" pageNumber="366" specimenCount="1" stateProvince="Erzurum" typeStatus="paratype">
<collectingCountry id="E6E6767A0354ED4230B9FC61C911FC31" box="[736,803,918,940]" name="Turkey" pageId="11" pageNumber="366">Turkey</collectingCountry>
:
<collectingRegion id="5C35F8080354ED423172FC60C948FC31" box="[811,890,919,940]" country="Turkey" name="Erzurum" pageId="11" pageNumber="366">Erzurum</collectingRegion>
,
<date id="EA4F102A0354ED4231DBFC61C9DFFC31" box="[898,1005,918,940]" pageId="11" pageNumber="366" value="1937-08-31">
<collectingDate id="FA0BE9C20354ED4231DBFC61C9DFFC31" box="[898,1005,918,940]" pageId="11" pageNumber="366" value="1937-08-31">31.viii.1937</collectingDate>
</date>
, 1♁, leg.
<collectorName id="3304533C0354ED423286FC58CB04FC59" box="[223,310,942,964]" pageId="11" pageNumber="366">O. Ecevit</collectorName>
, coll.
<collectorName id="3304533C0354ED423334FC59C81DFC59" box="[365,559,942,964]" pageId="11" pageNumber="366">Biologiezentrum Linz</collectorName>
(
<typeStatus id="414A88480354ED423065FC58C8BEFC59" box="[572,652,943,964]" pageId="11" pageNumber="366" type="paratype">paratype</typeStatus>
)
</materialsCitation>
;
<materialsCitation id="2E993CB70354ED4230C4FC59C93DFC41" collectingDate="1984-06-18" collectorName="S. Varegos &amp; K. Warncke &amp; Biologiezentrum Linz" country="Turkey" elevation="2000" location="Mt. Sat" pageId="11" pageNumber="366" specimenCount="2" specimenCount-female="2" stateProvince="Hakkari" typeStatus="paratype">
<collectingCountry id="E6E6767A0354ED4230C4FC59C8D2FC59" box="[669,736,942,964]" name="Turkey" pageId="11" pageNumber="366">Turkey</collectingCountry>
:
<collectingRegion id="5C35F8080354ED4230B0FC59C900FC59" box="[745,818,942,964]" country="Turkey" name="Hakkari" pageId="11" pageNumber="366">Hakkari</collectingRegion>
,
<collectorName id="3304533C0354ED423165FC58C9ADFC59" box="[828,927,943,964]" pageId="11" pageNumber="366">S. Varegös</collectorName>
/
<location id="9B2E60310354ED4231FFFC58C9DAFC59" LSID="urn:lsid:plazi:treatment:165887FC0356ED4232EBFF2DC958FC69:9B2E60310354ED4231FFFC58C9DAFC59" box="[934,1000,943,964]" country="Turkey" name="Mt. Sat" pageId="11" pageNumber="366" stateProvince="Hakkari">Mt. Sat</location>
,
<quantity id="59099B0F0354ED4231AAFC58CE08FC59" box="[1011,1082,943,964]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="366" unit="m" value="2000.0">
<elevation id="15DCD1D90354ED4231AAFC58CE08FC59" box="[1011,1082,943,964]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="366" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="EA4F102A0354ED423286FC31CB0CFC41" box="[223,318,966,988]" pageId="11" pageNumber="366" value="1984-06-18">
<collectingDate id="FA0BE9C20354ED423286FC31CB0CFC41" box="[223,318,966,988]" pageId="11" pageNumber="366" value="1984-06-18">18.vi.1984</collectingDate>
</date>
,
<specimenCount id="88F7FD630354ED423311FC30CB44FC41" box="[328,374,967,988]" count="2" pageId="11" pageNumber="366" type="female">2♀♀</specimenCount>
, leg.
<collectorName id="3304533C0354ED4233FEFC30C827FC41" box="[423,533,966,988]" pageId="11" pageNumber="366">K. Warncke</collectorName>
, coll.
<collectorName id="3304533C0354ED423013FC31C93DFC41" box="[586,783,966,988]" pageId="11" pageNumber="366">Biologiezentrum Linz</collectorName>
</materialsCitation>
;
<materialsCitation id="2E993CB70354ED423141FC31C958FC69" collectingDate="1984-06-14" collectorName="S. Varegos &amp; K. Warncke &amp; Biologiezentrum Linz." country="Turkey" elevation="2100" location="Mt. Sat" pageId="11" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Hakkari" typeStatus="paratype">
<collectingCountry id="E6E6767A0354ED423141FC31C969FC41" box="[792,859,966,988]" name="Turkey" pageId="11" pageNumber="366">Turkey</collectingCountry>
:
<collectingRegion id="5C35F8080354ED42313DFC31C99FFC41" box="[868,941,966,988]" country="Turkey" name="Hakkari" pageId="11" pageNumber="366">Hakkari</collectingRegion>
,
<collectorName id="3304533C0354ED4231EEFC30CE2BFC41" box="[951,1049,967,988]" pageId="11" pageNumber="366">S. Varegös</collectorName>
/
<location id="9B2E60310354ED423679FC30CACFFC69" LSID="urn:lsid:plazi:treatment:165887FC0356ED4232EBFF2DC958FC69:9B2E60310354ED423679FC30CACFFC69" country="Turkey" name="Mt. Sat" pageId="11" pageNumber="366" stateProvince="Hakkari">Mt. Sat</location>
,
<quantity id="59099B0F0354ED42335CFC28CB7BFC69" box="[261,329,991,1012]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="366" unit="m" value="2100.0">
<elevation id="15DCD1D90354ED42335CFC28CB7BFC69" box="[261,329,991,1012]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="366" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="EA4F102A0354ED42330BFC29CB83FC69" box="[338,433,990,1012]" pageId="11" pageNumber="366" value="1984-06-14">
<collectingDate id="FA0BE9C20354ED42330BFC29CB83FC69" box="[338,433,990,1012]" pageId="11" pageNumber="366" value="1984-06-14">14.vi.1984</collectingDate>
</date>
,
<specimenCount id="88F7FD630354ED4233E0FC28CBE7FC69" box="[441,469,991,1012]" count="1" pageId="11" pageNumber="366" type="female">1♀</specimenCount>
, leg.
<collectorName id="3304533C0354ED42305CFC28C843FC69" box="[517,625,990,1012]" pageId="11" pageNumber="366">K. Warncke</collectorName>
, coll.
<collectorName id="3304533C0354ED4230FAFC29C958FC69" box="[675,874,990,1012]" pageId="11" pageNumber="366">Biologiezentrum Linz.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>