150 lines
20 KiB
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150 lines
20 KiB
XML
<document id="B2B943F538FFDDBA5B0B730FEEB8B1DD" ID-CLB-Dataset="28504" ID-DOI="10.5281/zenodo.2579565" ID-GBIF-Dataset="2ffd1299-7a53-4ce9-868c-64e82f6fc053" ID-ISSN="1942-1354" ID-Zenodo-Dep="2579565" ID-ZooBank="urn:lsid:zoobank.org:pub:600D70CF-9674-44C6-87FF-DE1B0D2CAA4D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1551440540580" checkinUser="PlaziZenodoSync" docAuthor="Paulsen, M. J." docDate="2019" docId="BC3287C6FFC3FFC1E4AAFCE0FA54FA35" docLanguage="en" docName="insectaMundi.684.pdf" docOrigin="Insecta Mundi 2019 (684)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ochodaeus Dejean 1821" docType="key" docVersion="9" lastPageNumber="11" masterDocId="400BFFBEFFCFFFCDE46AFF9BFFF4FFF0" masterDocTitle="Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae)" masterLastPageNumber="14" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1551337755633" originalUpdateUser="PlaziZenodoSync" pageNumber="11" updateTime="1698700379637" updateUser="ExternalLinkService">
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<mods:mods id="853DCE64799D0607C6282E037E5B3D45" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9FB65BD913706CA529361227796FE126">
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<mods:title id="DABDC1B309AEB5820F17D2645A71401B">Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae)</mods:title>
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<mods:name id="7355CEC91C24DFB923FC933E5C7B775A" type="personal">
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<mods:roleTerm id="B154A1608DFFE112B1B676AA15C2ADBD">Author</mods:roleTerm>
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<mods:namePart id="54B9E1BF897DDD370698D467FB5CAE30">Paulsen, M. J.</mods:namePart>
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<mods:affiliation id="C4998E202B2DB9BA6612409BA9FE2864">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
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<mods:typeOfResource id="14CD32309EDEDBA771C92F38402B5B72">text</mods:typeOfResource>
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<mods:title id="9ECEAEDBBF072B52AD1F886E03A7B825">Insecta Mundi</mods:title>
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<mods:part id="1D234A23B7C7235D799AB7EFE1961EE8">
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<mods:date id="152A1778EA252CDC89F544BE9C0483CF">2019</mods:date>
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<mods:number id="77ED07082A57862D9006E63844260D34">2019-01-18</mods:number>
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<mods:number id="7E7E843CCE75B6A0AACA4E727125FDD0">2019</mods:number>
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<mods:detail id="EE25747CF53E3278FC7FA75B4387D872" type="issue">
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<mods:number id="184B5E4197313244E6047035B3CC56EF">684</mods:number>
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<mods:identifier id="AB759D1F5505FDFED8B124CDC7258981" type="CLB-Dataset">28504</mods:identifier>
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<mods:identifier id="3EC0D2C1DE529F9D74F9E1B7C684AADC" type="DOI">10.5281/zenodo.2579565</mods:identifier>
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<mods:identifier id="32DB014CC8652DAD44B8D1705510E8A0" type="ISSN">1942-1354</mods:identifier>
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<treatment id="BC3287C6FFC3FFC1E4AAFCE0FA54FA35" ID-DOI="http://doi.org/10.5281/zenodo.5315612" ID-GBIF-Taxon="154006306" ID-Zenodo-Dep="5315612" LSID="urn:lsid:plazi:treatment:BC3287C6FFC3FFC1E4AAFCE0FA54FA35" httpUri="http://treatment.plazi.org/id/BC3287C6FFC3FFC1E4AAFCE0FA54FA35" lastPageNumber="11" pageId="12" pageNumber="11">
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<subSubSection id="7C81655BFFC3FFC1E4AAFCE0FCE1FC63" box="[192,789,891,915]" pageId="12" pageNumber="11" type="nomenclature">
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<paragraph id="342436D0FFC3FFC1E4AAFCE0FCE1FC63" blockId="12.[192,789,891,915]" box="[192,789,891,915]" pageId="12" pageNumber="11">
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<emphasis id="06EFEAC2FFC3FFC1E4AAFCE0FCE1FC63" bold="true" box="[192,789,891,915]" pageId="12" pageNumber="11">
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Key to species of
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<taxonomicName id="F39B4D53FFC3FFC1E5C7FCE0FDB7FC63" ID-CoL="66FL" authorityName="Dejean" authorityYear="1821" box="[429,579,891,915]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="06EFEAC2FFC3FFC1E5C7FCE0FDB7FC63" bold="true" box="[429,579,891,915]" italics="true" pageId="12" pageNumber="11">Ochodaeus</emphasis>
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</taxonomicName>
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in Madagascar
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="7C81655BFFC3FFC1E4AAFC37FA54FA35" pageId="12" pageNumber="11" type="key">
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<paragraph id="342436D0FFC3FFC1E4AAFC37FA54FC33" blockId="12.[192,1440,939,1477]" box="[192,1440,939,964]" pageId="12" pageNumber="11">
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1. Pronotum everywhere strongly, densely granulate with elevated tubercles (
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<figureCitation id="ACA02A55FFC3FFC1E0C0FC37FB0AFC34" box="[1194,1278,940,964]" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="12" pageNumber="11">Fig. 13</figureCitation>
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)........
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<emphasis id="06EFEAC2FFC3FFC1E1FBFC30FA54FC33" bold="true" box="[1425,1440,939,963]" pageId="12" pageNumber="11">2</emphasis>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFC57FA54FBF4" blockId="12.[192,1440,939,1477]" pageId="12" pageNumber="11">
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— Pronotum either entirely smooth and punctate or with weak granulation confined to disc (
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<figureCitation id="ACA02A55FFC3FFC1E11AFC57FEA2FBF4" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="12" pageNumber="11">Fig. 14</figureCitation>
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)........................................................................
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<emphasis id="06EFEAC2FFC3FFC1E1FBFC77FA54FBF4" bold="true" box="[1425,1440,1004,1028]" pageId="12" pageNumber="11">4</emphasis>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFB87FA54FBC4" blockId="12.[192,1440,939,1477]" box="[192,1440,1052,1076]" pageId="12" pageNumber="11">
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2(1). Elytral tubercles on intervals sparse; body length less than 4 mm....
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<taxonomicName id="F39B4D53FFC3FFC1E0EAFB87FA54FBC4" ID-CoL="6SF2R" authority="Paulian" authorityName="Paulian" authorityYear="1976" box="[1152,1440,1052,1076]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="pygmaeus">
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<emphasis id="06EFEAC2FFC3FFC1E0EAFB87FA54FBC4" bold="true" box="[1152,1440,1052,1076]" pageId="12" pageNumber="11">
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<emphasis id="06EFEAC2FFC3FFC1E0EAFB87FAD9FBC3" bold="true" box="[1152,1325,1052,1076]" italics="true" pageId="12" pageNumber="11">O. pygmaeus</emphasis>
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Paulian
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFBA6FA54FBA4" blockId="12.[192,1440,939,1477]" box="[192,1440,1084,1109]" pageId="12" pageNumber="11">
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— Elytral tubercles dense; body length greater than 4 mm...............................
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<emphasis id="06EFEAC2FFC3FFC1E1FBFBA7FA54FBA4" bold="true" box="[1425,1440,1084,1108]" pageId="12" pageNumber="11">3</emphasis>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFBF7FA54FB54" blockId="12.[192,1440,939,1477]" pageId="12" pageNumber="11">
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3(2). Median thickening of clypeal bead variable, lateral thickening always evident (
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<figureCitation id="ACA02A55FFC3FFC1E09FFBF7FAB9FB74" box="[1269,1357,1132,1156]" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="12" pageNumber="11">Fig. 15</figureCitation>
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); body usually> 9.0 mm.......................................
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<taxonomicName id="F39B4D53FFC3FFC1E037FB17FA54FB54" ID-CoL="74F35" authority="Paulian" authorityName="Paulian" box="[1117,1440,1164,1188]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="isoanalensis">
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<emphasis id="06EFEAC2FFC3FFC1E037FB17FA54FB54" bold="true" box="[1117,1440,1164,1188]" pageId="12" pageNumber="11">
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<emphasis id="06EFEAC2FFC3FFC1E037FB17FAD8FB54" bold="true" box="[1117,1324,1164,1188]" italics="true" pageId="12" pageNumber="11">O. isoanalensis</emphasis>
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Paulian
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFB36FA54FB14" blockId="12.[192,1440,939,1477]" pageId="12" pageNumber="11">
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— Median thickening of clypeal bead simply convex, bead thin laterally (
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<figureCitation id="ACA02A55FFC3FFC1E024FB37FB55FB34" box="[1102,1185,1196,1220]" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="12" pageNumber="11">Fig. 16</figureCitation>
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); body length usually 4.5–8.0 mm...................................................
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<taxonomicName id="F39B4D53FFC3FFC1E0A9FB57FA54FB14" ID-CoL="48DDD" authority="Klug" authorityName="Klug" authorityYear="1832" box="[1219,1440,1228,1252]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="miliaris">
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<emphasis id="06EFEAC2FFC3FFC1E0A9FB57FA54FB14" bold="true" box="[1219,1440,1228,1252]" pageId="12" pageNumber="11">
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<emphasis id="06EFEAC2FFC3FFC1E0A9FB57FAA2FB14" bold="true" box="[1219,1366,1228,1252]" italics="true" pageId="12" pageNumber="11">O. miliaris</emphasis>
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Klug
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFB66FA54FAE4" blockId="12.[192,1440,939,1477]" box="[192,1440,1276,1301]" pageId="12" pageNumber="11">
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4(1). Pronotal surface simply punctate.............
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<emphasis id="06EFEAC2FFC3FFC1E714FB67FA54FAE4" bold="true" box="[894,1440,1276,1301]" pageId="12" pageNumber="11">
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<taxonomicName id="F39B4D53FFC3FFC1E714FB67FB1BFAE5" ID-CoL="48DDF" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[894,1263,1276,1301]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="modopunctatus" status="sp. nov.">
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<emphasis id="06EFEAC2FFC3FFC1E714FB67FB83FAE4" bold="true" box="[894,1143,1276,1300]" italics="true" pageId="12" pageNumber="11">O. modopunctatus</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="1DDC57B9FFC3FFC1E096FB66FA54FAE4" box="[1276,1440,1276,1301]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFA86FA54FAC5" blockId="12.[192,1440,939,1477]" box="[192,1440,1309,1333]" pageId="12" pageNumber="11">
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— Pronotal surface weakly granulate on disc..........................................
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<emphasis id="06EFEAC2FFC3FFC1E1FBFA86FA54FAC5" bold="true" box="[1425,1440,1309,1333]" pageId="12" pageNumber="11">5</emphasis>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFAD6FA54FA75" blockId="12.[192,1440,939,1477]" pageId="12" pageNumber="11">
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5(4). Clypeal bead evenly curved (
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<figureCitation id="ACA02A55FFC3FFC1E618FAD6FD3DFA95" box="[626,713,1357,1381]" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="12" pageNumber="11">Fig. 17</figureCitation>
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). Mentum deeply, entirely furrowed (
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<figureCitation id="ACA02A55FFC3FFC1E0FCFAD6FB29FA95" box="[1174,1245,1357,1381]" captionStart="Figures 4–12" captionStartId="11.[192,285,1819,1841]" captionTargetBox="[192,1440,213,1793]" captionTargetId="figure@11.[192,1440,213,1793]" captionTargetPageId="11" captionText="Figures 4–12. Additional characters of Madagascar Ochodaeus species. 4–6) Mentum, shading indicates foveae. 7–9) Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres. 10–12) Right stridulatory peg, dorsal view. 4, 7, 10) O. modopunctatus Paulsen, new species. 5, 8, 11) O. polypollicatus Paulsen, new species. 6, 9, 12) O. meandrus Paulsen, new species." httpUri="https://zenodo.org/record/2579571/files/figure.png" pageId="12" pageNumber="11">Fig. 5</figureCitation>
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). Males usually with 2 pollices on protibia..................
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<emphasis id="06EFEAC2FFC3FFC1E7FAFAF6FA54FA75" bold="true" box="[912,1440,1388,1413]" pageId="12" pageNumber="11">
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<taxonomicName id="F39B4D53FFC3FFC1E7FAFAF6FB1BFA75" ID-CoL="48DDY" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[912,1263,1388,1413]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="polypollicatus" status="sp. nov.">
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<emphasis id="06EFEAC2FFC3FFC1E7FAFAF6FB83FA74" bold="true" box="[912,1143,1388,1413]" italics="true" pageId="12" pageNumber="11">O. polypollicatus</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="1DDC57B9FFC3FFC1E096FAF6FA54FA75" box="[1276,1440,1389,1413]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="342436D0FFC3FFC1E4AAFA15FA54FA35" blockId="12.[192,1440,939,1477]" pageId="12" pageNumber="11">
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— Clypeal bead wavy (
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<figureCitation id="ACA02A55FFC3FFC1E66CFA16FDA9FA55" box="[518,605,1421,1445]" captionStart="Figures 13–18" captionStartId="13.[192,287,1399,1421]" captionTargetBox="[192,1440,213,1365]" captionTargetId="figure@13.[192,1440,213,1365]" captionTargetPageId="13" captionText="Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin." httpUri="https://zenodo.org/record/2579573/files/figure.png" pageId="12" pageNumber="11">Fig. 18</figureCitation>
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). Mentum with furrow not reaching past middle (
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<figureCitation id="ACA02A55FFC3FFC1E0D0FA16FAF6FA55" box="[1210,1282,1421,1445]" captionStart="Figures 4–12" captionStartId="11.[192,285,1819,1841]" captionTargetBox="[192,1440,213,1793]" captionTargetId="figure@11.[192,1440,213,1793]" captionTargetPageId="11" captionText="Figures 4–12. Additional characters of Madagascar Ochodaeus species. 4–6) Mentum, shading indicates foveae. 7–9) Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres. 10–12) Right stridulatory peg, dorsal view. 4, 7, 10) O. modopunctatus Paulsen, new species. 5, 8, 11) O. polypollicatus Paulsen, new species. 6, 9, 12) O. meandrus Paulsen, new species." httpUri="https://zenodo.org/record/2579571/files/figure.png" pageId="12" pageNumber="11">Fig. 6</figureCitation>
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). Males with a single pollex on protibia......................
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<emphasis id="06EFEAC2FFC3FFC1E7A3FA36FA54FA35" bold="true" box="[969,1440,1452,1477]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="F39B4D53FFC3FFC1E7A3FA36FB1BFA35" ID-CoL="48DDB" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[969,1263,1452,1477]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="meandrus" status="sp. nov.">
|
||
<emphasis id="06EFEAC2FFC3FFC1E7A3FA36FB83FA34" bold="true" box="[969,1143,1452,1477]" italics="true" pageId="12" pageNumber="11">O. meandrus</emphasis>
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||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="1DDC57B9FFC3FFC1E096FA36FA54FA35" box="[1276,1440,1453,1477]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |