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<document id="408E4FCF20EDEE88756C49174AB4A79B" ID-DOI="10.5281/zenodo.212354" ID-GBIF-Dataset="a56fd6bf-13cc-4858-becb-0b6665a74cc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="212354" ID-ZooBank="D25AC77D-2E5B-4B99-8449-812558DB22E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460530626989" checkinUser="plazi" docAuthor="Frederiksen, Sara B. &amp; Enghoff, Henrik" docDate="2012" docId="42445940FF91BF6FFF2110E1FEE4FD82" docLanguage="en" docName="zt03485p082.pdf" docOrigin="Zootaxa 3485" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Xystopyge pelecys Frederiksen &amp; Enghoff, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="75" masterDocId="BE7D2138FF95BF69FFB61508FFE4FFBE" masterDocTitle="East African odontopygid millipedes 1: Five new species of Xystopyge (Attems, 1909) and a proposal for a new gonopod terminology (Diplopoda; Spirostreptida; Odontopygidae)" masterLastPageNumber="82" masterPageNumber="69" pageNumber="73" updateTime="1698315754598" updateUser="plazi">
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<mods:title id="3B4B5D123F9B3D88F19CDA5A6351F484">East African odontopygid millipedes 1: Five new species of Xystopyge (Attems, 1909) and a proposal for a new gonopod terminology (Diplopoda; Spirostreptida; Odontopygidae)</mods:title>
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<mods:namePart id="BFDF9F620686D15DC29EDDC703364F60">Frederiksen, Sara B.</mods:namePart>
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<mods:name id="21CE7F27486EFD21AF6C8ECE76089F17" type="personal">
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<mods:namePart id="043AEB6BAF613871153A97C7F10618F2">Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:typeOfResource id="B853CF193A9B70107346BB92E09A0B9F">text</mods:typeOfResource>
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<mods:title id="082378382476C3D28132B4B5ADDBB9E9">Zootaxa</mods:title>
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<mods:part id="7F0A8A73C9231F712A3A832AEA0AEFE1">
<mods:date id="9E3F414D453E92DDB3EB89D06F3239F8">2012</mods:date>
<mods:detail id="603D68BD63E2746C325E8CD1F89D2492" type="volume">
<mods:number id="B4DF5D6567893E3E6B6B2660A5910D7C">3485</mods:number>
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<mods:identifier id="7F5A00846099AA54AD58D849F471C926" type="DOI">10.5281/zenodo.212354</mods:identifier>
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</mods:mods>
<treatment id="42445940FF91BF6FFF2110E1FEE4FD82" ID-DOI="http://doi.org/10.5281/zenodo.6172389" ID-GBIF-Taxon="119527117" ID-Zenodo-Dep="6172389" LSID="urn:lsid:plazi:treatment:42445940FF91BF6FFF2110E1FEE4FD82" httpUri="http://treatment.plazi.org/id/42445940FF91BF6FFF2110E1FEE4FD82" lastPageId="6" lastPageNumber="75" pageId="4" pageNumber="73">
<subSubSection id="82F7BBDDFF91BF6DFF2110E1FEE9F99A" pageId="4" pageNumber="73" type="nomenclature">
<paragraph id="CA52E856FF91BF6DFF2110E1FE5CF9BC" blockId="4.[151,440,1512,1572]" box="[151,440,1512,1539]" pageId="4" pageNumber="73">
<heading id="911A5F3AFF91BF6DFF2110E1FE5CF9BC" bold="true" box="[151,440,1512,1539]" fontSize="11" level="1" pageId="4" pageNumber="73" reason="1">
<emphasis id="F8993444FF91BF6DFF2110E1FE5CF9BC" bold="true" box="[151,440,1512,1539]" pageId="4" pageNumber="73">
<taxonomicName id="0DED93D5FF91BF6DFF2110E1FE8AF9BD" ID-CoL="5CQHV" box="[151,366,1513,1539]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="pelecys" status="sp. nov.">
<emphasis id="F8993444FF91BF6DFF2110E1FE8AF9BD" bold="true" box="[151,366,1513,1539]" italics="true" pageId="4" pageNumber="73">Xystopyge pelecys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E3AA893FFF91BF6DFEC010E0FE5CF9BC" box="[374,440,1512,1538]" pageId="4" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CA52E856FF91BF6DFF211304FEE9F99A" blockId="4.[151,440,1512,1572]" box="[151,269,1548,1572]" pageId="4" pageNumber="73">
<figureCitation id="52D6F4D3FF91BF6DFF211304FEE9F99A" box="[151,269,1548,1572]" captionStart-0="FIGURE 1 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9 10" captionStartId-0="3.[151,253,1616,1639]" captionStartId-1="5.[151,250,610,633]" captionStartId-2="6.[151,250,1442,1465]" captionTargetBox-0="[170,1410,217,1566]" captionTargetBox-1="[155,1431,201,580]" captionTargetBox-2="[165,1377,629,1406]" captionTargetId-0="figure@3.[151,1436,195,1592]" captionTargetId-1="figure@5.[151,1436,193,589]" captionTargetId-2="figure@6.[151,1436,625,1419]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1 7. Xystopyge pelecys n. sp. 1: Limbus, external view. 2: Limbus, internal view. 3: Posterior sternum. 4: Gonopod coxa, anterior view. 5: Gonopod telepodite. 6 7: Telomere apex. Scales 10 μm (1 2), 100 μm (3 7). — al 1, al 2: apical lamellae of telomere." captionText-1="FIGURE 8. Xystopyge spp., solenomere apex (X. proplicatus: complete solenomere). — asl: apical solenomere lamella, ass: apical solenomere spine, bss: basal solenomere spine, msl: mesal solenomere lamella." captionText-2="FIGURE 9 10. Xystopyge pelecys n. sp., 9: left gonopod, anterior view. 10: left gonopod, apical view. Scales 0.5 mm. — al 1, al 2: apical lamellae of telomere, amp: apical metaplical process, btl: basal lamella on telomere, h: hook on telomere, lmp: lateral metaplical process, lms: lateral metaplical spine, mf: metaplical flange, mmf: mesal metaplical flange, mmp, median metaplical process, pl: proplical lobe, st: anterior sternum tp: axehead-like process on telomere." httpUri-0="https://zenodo.org/record/212355/files/figure.png" httpUri-1="https://zenodo.org/record/212356/files/figure.png" httpUri-2="https://zenodo.org/record/212357/files/figure.png" pageId="4" pageNumber="73">Figs. 110</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="82F7BBDDFF91BF6DFF21135BFC63F8FA" pageId="4" pageNumber="73" type="materials_examined">
<paragraph id="CA52E856FF91BF6DFF21135BFC63F8FA" blockId="4.[151,1436,1619,1968]" pageId="4" pageNumber="73">
<emphasis id="F8993444FF91BF6DFF21135BFEEDF9D2" bold="true" box="[151,265,1619,1644]" pageId="4" pageNumber="73">Material:</emphasis>
<emphasis id="F8993444FF91BF6DFEAD135CFE66F9D2" box="[283,386,1620,1644]" italics="true" pageId="4" pageNumber="73">
<typeStatus id="155656F4FF91BF6DFEAD135CFE66F9D2" box="[283,386,1620,1644]" pageId="4" pageNumber="73" type="holotype">Holotype</typeStatus>
</emphasis>
male:
<collectingCountry id="B2FAA8C6FF91BF6DFE50135CFD9CF9D2" box="[486,632,1620,1644]" name="Tanzania" pageId="4" pageNumber="73">TANZANIA</collectingCountry>
, Tanga region, Muheza district, Nilo forest reserve
<geoCoordinate id="AFD98E91FF91BF6DFAAE135CFA78F9D2" box="[1304,1436,1620,1644]" direction="south" orientation="latitude" pageId="4" pageNumber="73" precision="15" value="-4.9044447">04°54`16”S</geoCoordinate>
<geoCoordinate id="AFD98E91FF91BF6DFF21137FFEFBF931" box="[151,287,1655,1679]" direction="east" orientation="longitude" pageId="4" pageNumber="73" precision="15" value="38.662502">38°39`45”E</geoCoordinate>
,
<date id="BE53CE96FF91BF6DFE87137FFE51F931" box="[305,437,1655,1679]" pageId="4" pageNumber="73" value="2000-06-09">09/VI/2000</date>
, Frontier
<collectingCountry id="B2FAA8C6FF91BF6DFD98137FFD71F931" box="[558,661,1655,1679]" name="Tanzania" pageId="4" pageNumber="73">Tanzania</collectingCountry>
, (
<collectionCode id="ACFC7093FF91BF6DFD18137FFCE5F931" box="[686,769,1655,1679]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
00020501).
<emphasis id="F8993444FF91BF6DFC2A1371FBEAF92E" box="[924,1038,1657,1680]" italics="true" pageId="4" pageNumber="73">
<typeStatus id="155656F4FF91BF6DFC2A1371FBEAF92E" box="[924,1038,1657,1680]" pageId="4" pageNumber="73" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="DCEB23DFFF91BF6DFBAC137FFB9FF931" box="[1050,1147,1655,1679]" pageId="4" pageNumber="73" type="male">2 males</specimenCount>
: same data as
<typeStatus id="155656F4FF91BF6DFA85137FFA7CF931" box="[1331,1432,1655,1679]" pageId="4" pageNumber="73" type="holotype">holotype</typeStatus>
, (
<collectionCode id="ACFC7093FF91BF6DFF281394FF15F90A" box="[158,241,1692,1716]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
0 0 0 20521,
<collectionCode id="ACFC7093FF91BF6DFECB1394FE2AF90A" box="[381,462,1692,1716]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
00101494).
<specimenCount id="DCEB23DFFF91BF6DFDD21394FD53F90A" box="[612,695,1692,1716]" pageId="4" pageNumber="73" type="male">1 male</specimenCount>
:
<collectingCountry id="B2FAA8C6FF91BF6DFD731394FCB3F90A" box="[709,855,1692,1716]" name="Tanzania" pageId="4" pageNumber="73">TANZANIA</collectingCountry>
, Tanga region, Muheza + Korogwe districts, Nilo Forest reserve,
<geoCoordinate id="AFD98E91FF91BF6DFEEC13B7FE12F969" box="[346,502,1727,1751]" direction="south" orientation="latitude" pageId="4" pageNumber="73" precision="1" value="-4.959194">04°57`33,1”S</geoCoordinate>
,
<geoCoordinate id="AFD98E91FF91BF6DFDB813B7FD48F969" box="[526,684,1727,1751]" direction="east" orientation="longitude" pageId="4" pageNumber="73" precision="1" value="38.64128">38°38`28,6”E</geoCoordinate>
.
<date id="BE53CE96FF91BF6DFD7413B7FCD8F969" box="[706,828,1727,1751]" pageId="4" pageNumber="73" value="2001-02-13">13/II/2001</date>
, Frontier
<collectingCountry id="B2FAA8C6FF91BF6DFC0913B7FBC3F969" box="[959,1063,1727,1751]" name="Tanzania" pageId="4" pageNumber="73">Tanzania</collectingCountry>
, (
<collectionCode id="ACFC7093FF91BF6DFBF313B7FB7CF969" box="[1093,1176,1727,1751]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
00020511).
<specimenCount id="DCEB23DFFF91BF6DFA8B13B7FA7DF969" box="[1341,1433,1727,1751]" pageId="4" pageNumber="73" type="male">1 male</specimenCount>
:
<collectingCountry id="B2FAA8C6FF91BF6DFF2113ECFECDF942" box="[151,297,1764,1788]" name="Tanzania" pageId="4" pageNumber="73">TANZANIA</collectingCountry>
, Tanga region, Muheza district, Nilo Forest reserve,
<geoCoordinate id="AFD98E91FF91BF6DFCC313ECFC2BF942" box="[885,975,1764,1788]" direction="south" orientation="latitude" pageId="4" pageNumber="73" precision="925" value="-4.85">04°51'S</geoCoordinate>
,
<geoCoordinate id="AFD98E91FF91BF6DFC6C13ECFBD2F942" box="[986,1078,1764,1788]" direction="east" orientation="longitude" pageId="4" pageNumber="73" precision="925" value="38.65">38°39'E</geoCoordinate>
,
<date id="BE53CE96FF91BF6DFBF413ECFB20F942" box="[1090,1220,1764,1788]" pageId="4" pageNumber="73" value="2000-09-24">24.IX.2000</date>
, Frontier
<collectingCountry id="B2FAA8C6FF91BF6DFA8613ECFA73F942" box="[1328,1431,1764,1788]" name="Tanzania" pageId="4" pageNumber="73">Tanzania</collectingCountry>
, (
<collectionCode id="ACFC7093FF91BF6DFF28120FFF15F8A1" box="[158,241,1799,1823]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
00020516).
<specimenCount id="DCEB23DFFF91BF6DFE33120FFE07F8A1" box="[389,483,1799,1823]" pageId="4" pageNumber="73" type="male">2 males</specimenCount>
:
<collectingCountry id="B2FAA8C6FF91BF6DFE46120FFDB3F8A1" box="[496,599,1799,1823]" name="Tanzania" pageId="4" pageNumber="73">Tanzania</collectingCountry>
, East Usambara mts, Amani,
<quantity id="0D1545B3FF91BF6DFC04120FFBE8F89E" box="[946,1036,1799,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="73" unit="m" value="1000.0">1000 m</quantity>
.
<date id="BE53CE96FF91BF6DFBAC120FFB41F8A1" box="[1050,1189,1799,1823]" pageId="4" pageNumber="73" value="1980-07-10">10/VII/1980</date>
and
<date id="BE53CE96FF91BF6DFB57120FFA8AF8A1" box="[1249,1390,1799,1823]" pageId="4" pageNumber="73" value="1980-07-20">20/VII/1980</date>
, M. Stoltze and N. Scharff leg. (
<collectionCode id="ACFC7093FF91BF6DFE7B1224FDC4F8FA" box="[461,544,1836,1860]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
00020506),
<collectionCode id="ACFC7093FF91BF6DFD181224FD1BF8FA" box="[686,767,1836,1860]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="73">ZMUC</collectionCode>
00020505).
</paragraph>
</subSubSection>
<subSubSection id="82F7BBDDFF91BF6DFF711247FE5BF80E" pageId="4" pageNumber="73" type="diagnosis">
<paragraph id="CA52E856FF91BF6DFF711247FE5BF80E" blockId="4.[151,1436,1619,1968]" pageId="4" pageNumber="73">
<emphasis id="F8993444FF91BF6DFF711247FEA7F8D6" bold="true" box="[199,323,1871,1896]" pageId="4" pageNumber="73">Diagnosis:</emphasis>
Metaplica large, sickle-shaped, proplica very small in comparison. Differs from congeneric species by the broad flat rounded lateral metaplical process and the axe-head-shaped process on the telomere, the shape of both processes are unique.
</paragraph>
</subSubSection>
<caption id="9E92B8DEFF90BF6CFF21176AFC1FFD29" httpUri="https://zenodo.org/record/212356/files/figure.png" pageId="5" pageNumber="74" targetBox="[155,1431,201,580]" targetPageId="5">
<paragraph id="CA52E856FF90BF6CFF21176AFC1FFD29" blockId="5.[151,1436,610,663]" pageId="5" pageNumber="74">
<emphasis id="F8993444FF90BF6CFF21176AFEF1FDC6" bold="true" box="[151,277,610,633]" pageId="5" pageNumber="74">FIGURE 8.</emphasis>
<taxonomicName id="0DED93D5FF90BF6CFEA8176CFE62FDC7" box="[286,390,612,633]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="F8993444FF90BF6CFEA8176CFE62FDC7" box="[286,390,612,633]" italics="true" pageId="5" pageNumber="74">Xystopyge</emphasis>
</taxonomicName>
spp., solenomere apex (
<taxonomicName id="0DED93D5FF90BF6CFD30176CFCF3FDC7" box="[646,791,611,633]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="proplicatus">
<emphasis id="F8993444FF90BF6CFD30176CFCF3FDC7" box="[646,791,611,633]" italics="true" pageId="5" pageNumber="74">X. proplicatus</emphasis>
</taxonomicName>
: complete solenomere).—
<emphasis id="F8993444FF90BF6CFB9C176BFBA3FDC7" box="[1066,1095,611,633]" italics="true" pageId="5" pageNumber="74">asl</emphasis>
: apical solenomere
<taxonomicName id="0DED93D5FF90BF6CFAAD176AFA8CFDC6" box="[1307,1384,610,632]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lamella">lamella</taxonomicName>
,
<emphasis id="F8993444FF90BF6CFAC2176CFA70FDC7" box="[1396,1428,612,633]" italics="true" pageId="5" pageNumber="74">ass</emphasis>
: apical solenomere spine,
<emphasis id="F8993444FF90BF6CFE211789FE53FD29" box="[407,439,641,663]" italics="true" pageId="5" pageNumber="74">bss</emphasis>
: basal solenomere spine,
<emphasis id="F8993444FF90BF6CFD0A1789FD3BFD29" box="[700,735,641,663]" italics="true" pageId="5" pageNumber="74">msl</emphasis>
: mesal solenomere
<taxonomicName id="0DED93D5FF90BF6CFC1F1789FC13FD29" box="[937,1015,641,663]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lamella">lamella</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="82F7BBDDFF90BF6CFF7117B7FF12FD44" pageId="5" pageNumber="74" type="etymology">
<paragraph id="CA52E856FF90BF6CFF7117B7FF12FD44" blockId="5.[151,1436,703,2023]" pageId="5" pageNumber="74">
<emphasis id="F8993444FF90BF6CFF7117B7FEB5FD69" bold="true" box="[199,337,703,727]" pageId="5" pageNumber="74">Etymology:</emphasis>
The name is a noun in apposition and refers to the axe-shaped telomeric process (
<taxonomicName id="0DED93D5FF90BF6CFB4217B7FA78FD69" authority="Greek" authorityName="Greek" box="[1268,1436,703,727]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="pelecys">pelecys, Greek</taxonomicName>
for axe).
</paragraph>
</subSubSection>
<subSubSection id="82F7BBDDFF90BF6CFF71160EFD28F8C4" pageId="5" pageNumber="74" type="materials_examined">
<paragraph id="CA52E856FF90BF6CFF71160EFD66FCA1" blockId="5.[151,1436,703,2023]" box="[199,642,774,799]" pageId="5" pageNumber="74">
<emphasis id="F8993444FF90BF6CFF71160EFD66FCA1" bold="true" box="[199,642,774,799]" pageId="5" pageNumber="74">Description (based on all specimens):</emphasis>
</paragraph>
<paragraph id="CA52E856FF90BF6CFF711622FA6BFCFC" blockId="5.[151,1436,703,2023]" box="[199,1423,810,834]" pageId="5" pageNumber="74">
Body length
<quantity id="0D1545B3FF90BF6CFEE11622FE36FCFC" box="[343,466,810,834]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" metricValueMax="6.4" metricValueMin="5.9" pageId="5" pageNumber="74" unit="mm" value="61.5" valueMax="64.0" valueMin="59.0">5964 mm</quantity>
, diameter 3.54.0 mm. 6265 podous body rings, no apodous rings in front of telson.
</paragraph>
<paragraph id="CA52E856FF90BF6CFF711647FB50FBDC" blockId="5.[151,1436,703,2023]" pageId="5" pageNumber="74">
Coloration (after 11 years in alcohol): Head and antennae mostly brown, but with two lighter brown/yellow spots between the antennae and a darker band between the eyes. Supralabral region yellow, labrum brown. Collum brown with yellow edges. The rest of the body rings mostly brown (although much is faded to grey in alcohol, original colour still visible some places), but posterior part of metazona yellow, resulting in a banded look. Legs yellow. Preanal ring dark brown, anal valves yellowish brown, lighter towards the edges. Some individuals darker brown ranging to almost black (with greyish areas probably due to preservation). Yellow areas and colour pattern the same in all specimens. One specimen (
<collectionCode id="ACFC7093FF90BF6CFDCE112FFD2FFB81" box="[632,715,1063,1087]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="5" pageNumber="74">ZMUC</collectionCode>
00020511) with a very light yellow colour instead of the brown colour; the yellow colour shared by all specimens being the darker of the two yellow nuances.
</paragraph>
<paragraph id="CA52E856FF90BF6CFF711167FDE3FA3C" blockId="5.[151,1436,703,2023]" pageId="5" pageNumber="74">
Head and body: 1+4+1 supralabral setae and a row of labral setae. Antennae reaching back to body ring 7. Ocelli in 6 horizontal rows. Length of eye (along upper row of ocelli) 0.53 × interocular distance. Distal margin of mandibular stipes shallowly bilobed. Lateral lobes of collum square, with three furrows, upper two furrows distinct, the lower marginal furrow barely visible. Ozopores from body ring 6, the exact location of the ozopores in relation to suture varying amongst individuals. Metazonites with furrows ventrally. Anal valves smooth, posteriorly perfectly rounded, each with a small dorsal spine. Anal valves apparently with at least one, possibly two posterior setae each. On many specimens the setae appear to be broken off. Limbus (
<figureCitation id="52D6F4D3FF90BF6CFBEB104FFB51FAE1" box="[1117,1205,1351,1375]" captionStart="FIGURE 1 7" captionStartId="3.[151,253,1616,1639]" captionTargetBox="[170,1410,217,1566]" captionTargetId="figure@3.[151,1436,195,1592]" captionTargetPageId="3" captionText="FIGURE 1 7. Xystopyge pelecys n. sp. 1: Limbus, external view. 2: Limbus, internal view. 3: Posterior sternum. 4: Gonopod coxa, anterior view. 5: Gonopod telepodite. 6 7: Telomere apex. Scales 10 μm (1 2), 100 μm (3 7). — al 1, al 2: apical lamellae of telomere." httpUri="https://zenodo.org/record/212355/files/figure.png" pageId="5" pageNumber="74">Fig.12</figureCitation>
) serrated. Male legs with postfemoral and tibial pads.
</paragraph>
<paragraph id="CA52E856FF90BF6CFF711087FD28F8C4" blockId="5.[151,1436,703,2023]" pageId="5" pageNumber="74">
Gonopods (
<figureCitation id="52D6F4D3FF90BF6CFEFF1087FE21FA19" box="[329,453,1423,1447]" captionStart-0="FIGURE 1 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9 10" captionStartId-0="3.[151,253,1616,1639]" captionStartId-1="5.[151,250,610,633]" captionStartId-2="6.[151,250,1442,1465]" captionTargetBox-0="[170,1410,217,1566]" captionTargetBox-1="[155,1431,201,580]" captionTargetBox-2="[165,1377,629,1406]" captionTargetId-0="figure@3.[151,1436,195,1592]" captionTargetId-1="figure@5.[151,1436,193,589]" captionTargetId-2="figure@6.[151,1436,625,1419]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1 7. Xystopyge pelecys n. sp. 1: Limbus, external view. 2: Limbus, internal view. 3: Posterior sternum. 4: Gonopod coxa, anterior view. 5: Gonopod telepodite. 6 7: Telomere apex. Scales 10 μm (1 2), 100 μm (3 7). — al 1, al 2: apical lamellae of telomere." captionText-1="FIGURE 8. Xystopyge spp., solenomere apex (X. proplicatus: complete solenomere). — asl: apical solenomere lamella, ass: apical solenomere spine, bss: basal solenomere spine, msl: mesal solenomere lamella." captionText-2="FIGURE 9 10. Xystopyge pelecys n. sp., 9: left gonopod, anterior view. 10: left gonopod, apical view. Scales 0.5 mm. — al 1, al 2: apical lamellae of telomere, amp: apical metaplical process, btl: basal lamella on telomere, h: hook on telomere, lmp: lateral metaplical process, lms: lateral metaplical spine, mf: metaplical flange, mmf: mesal metaplical flange, mmp, median metaplical process, pl: proplical lobe, st: anterior sternum tp: axehead-like process on telomere." httpUri-0="https://zenodo.org/record/212355/files/figure.png" httpUri-1="https://zenodo.org/record/212356/files/figure.png" httpUri-2="https://zenodo.org/record/212357/files/figure.png" pageId="5" pageNumber="74">Figs. 310</figureCitation>
): Anterior sternum as in
<emphasis id="F8993444FF90BF6CFD6A1098FCB2FA19" box="[732,854,1423,1447]" italics="true" pageId="5" pageNumber="74">
<taxonomicName id="0DED93D5FF90BF6CFD6A1098FCB6FA19" box="[732,850,1423,1447]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="robusta">X. robusta</taxonomicName>
,
</emphasis>
a simple narrow triangle.—Coxa: Mesal metaplical flange (
<emphasis id="F8993444FF90BF6CFF5810BBFEFAFA75" box="[238,286,1459,1483]" italics="true" pageId="5" pageNumber="74">mmf</emphasis>
) small as in all
<taxonomicName id="0DED93D5FF90BF6CFE6010BCFDADFA75" box="[470,585,1460,1483]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="F8993444FF90BF6CFE6010BCFDADFA75" box="[470,585,1460,1483]" italics="true" pageId="5" pageNumber="74">Xystopyge</emphasis>
</taxonomicName>
species. Proplica somewhat larger than mesal metaplical flange, but not as big as in other
<taxonomicName id="0DED93D5FF90BF6CFE8B10D0FE54FA51" box="[317,432,1496,1519]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="F8993444FF90BF6CFE8B10D0FE54FA51" box="[317,432,1496,1519]" italics="true" pageId="5" pageNumber="74">Xystopyge</emphasis>
</taxonomicName>
species, ending in a distad proplical lobe (
<emphasis id="F8993444FF90BF6CFC3910DFFC41FA51" box="[911,933,1495,1519]" italics="true" pageId="5" pageNumber="74">pl</emphasis>
). Metaplica with a broad flat rounded lateral metaplical process (
<emphasis id="F8993444FF90BF6CFEC310F3FE7BF9AD" box="[373,415,1531,1555]" italics="true" pageId="5" pageNumber="74">lmp</emphasis>
) with an anteriad lateral metaplical spine (
<emphasis id="F8993444FF90BF6CFCCD10F3FC46F9AD" box="[891,930,1531,1555]" italics="true" pageId="5" pageNumber="74">lms</emphasis>
) situated just below it, a pointed mediodistad apical metaplical process (
<emphasis id="F8993444FF90BF6CFE731328FE11F989" box="[453,501,1568,1591]" italics="true" pageId="5" pageNumber="74">amp</emphasis>
), a triangular median metaplical process (
<emphasis id="F8993444FF90BF6CFC611328FBE9F989" box="[983,1037,1568,1591]" italics="true" pageId="5" pageNumber="74">mmp</emphasis>
) projecting anteriad and forming a roof over the emerging telopodite basomere, and a triangular distal metaplical flange (
<emphasis id="F8993444FF90BF6CFB12134BFB24F9E5" box="[1188,1216,1603,1627]" italics="true" pageId="5" pageNumber="74">mf</emphasis>
), situated between apical metaplical process and median metaplical process.—Telopodite: Basomere with an extended torsotope, without spines, distally slightly thickened. Solenomere whip-like with a spine apically (
<emphasis id="F8993444FF90BF6CFBCD1384FB7AF91D" box="[1147,1182,1676,1699]" italics="true" pageId="5" pageNumber="74">ass</emphasis>
), closely appressed to telomere; solenomere and telomere forming a complete horizontal loop. Telomere ribbon-shaped, with a basal
<taxonomicName id="0DED93D5FF90BF6CFF2113DAFF0EF954" box="[151,234,1746,1770]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lamella">lamella</taxonomicName>
(
<emphasis id="F8993444FF90BF6CFF4813DBFEFFF955" box="[254,283,1747,1771]" italics="true" pageId="5" pageNumber="74">btl</emphasis>
) ending distally in a blunt hook (
<emphasis id="F8993444FF90BF6CFD0413DBFD24F955" box="[690,704,1747,1771]" italics="true" pageId="5" pageNumber="74">h</emphasis>
); c. 1/3 along its length a flattened, axehead-like process (
<emphasis id="F8993444FF90BF6CFACB13DCFA77F955" box="[1405,1427,1748,1771]" italics="true" pageId="5" pageNumber="74">tp</emphasis>
) projecting into the telopodital loop. Telomere apically broadened, ending in two rounded lamellae (
<emphasis id="F8993444FF90BF6CFABE13FFFA83F8B1" box="[1288,1383,1783,1807]" italics="true" pageId="5" pageNumber="74">al1, al2.</emphasis>
<figureCitation id="52D6F4D3FF90BF6CFAD913FFFF20F88C" captionStart="FIGURE 1 7" captionStartId="3.[151,253,1616,1639]" captionTargetBox="[170,1410,217,1566]" captionTargetId="figure@3.[151,1436,195,1592]" captionTargetPageId="3" captionText="FIGURE 1 7. Xystopyge pelecys n. sp. 1: Limbus, external view. 2: Limbus, internal view. 3: Posterior sternum. 4: Gonopod coxa, anterior view. 5: Gonopod telepodite. 6 7: Telomere apex. Scales 10 μm (1 2), 100 μm (3 7). — al 1, al 2: apical lamellae of telomere." httpUri="https://zenodo.org/record/212355/files/figure.png" pageId="5" pageNumber="74">Fig. 67</figureCitation>
);
<emphasis id="F8993444FF90BF6CFF6D1213FF1BF88D" box="[219,255,1819,1843]" italics="true" pageId="5" pageNumber="74">al1</emphasis>
with a spine-like process. Posterior sternum (
<figureCitation id="52D6F4D3FF90BF6CFCBB1212FCB6F88C" box="[781,850,1818,1842]" captionStart="FIGURE 1 7" captionStartId="3.[151,253,1616,1639]" captionTargetBox="[170,1410,217,1566]" captionTargetId="figure@3.[151,1436,195,1592]" captionTargetPageId="3" captionText="FIGURE 1 7. Xystopyge pelecys n. sp. 1: Limbus, external view. 2: Limbus, internal view. 3: Posterior sternum. 4: Gonopod coxa, anterior view. 5: Gonopod telepodite. 6 7: Telomere apex. Scales 10 μm (1 2), 100 μm (3 7). — al 1, al 2: apical lamellae of telomere." httpUri="https://zenodo.org/record/212355/files/figure.png" pageId="5" pageNumber="74">Fig. 3</figureCitation>
) as in
<emphasis id="F8993444FF90BF6CFC121214FB99F88D" box="[932,1149,1819,1843]" italics="true" pageId="5" pageNumber="74">
<taxonomicName id="0DED93D5FF90BF6CFC121214FB93F88D" box="[932,1143,1819,1843]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="robusta">Xystopyge robusta</taxonomicName>
:
</emphasis>
slightly elongate square, indented with a narrowing towards one end. There are minute variations in the exact shapes of the basal
<taxonomicName id="0DED93D5FF90BF6CFAF41237FA73F8E9" box="[1346,1431,1855,1879]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lamella">lamella</taxonomicName>
, blunt hook and axehead-shaped telomeric process.
</paragraph>
</subSubSection>
<subSubSection id="82F7BBDDFF90BF6FFF71128FFEE4FD82" lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74" type="discussion">
<paragraph id="CA52E856FF90BF6FFF71128FFB48FE81" blockId="5.[151,1436,703,2023]" lastBlockId="6.[151,1436,151,572]" lastPageId="6" lastPageNumber="75" pageId="5" pageNumber="74">
<emphasis id="F8993444FF90BF6CFF71128FFED0F821" bold="true" box="[199,308,1927,1951]" pageId="5" pageNumber="74">Remarks</emphasis>
: The new species very much resembles the taxon described as
<taxonomicName id="0DED93D5FF90BF6CFC441280FB34F821" box="[1010,1232,1927,1951]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="subSpecies" species="robusta" subSpecies="hoffmani">
<emphasis id="F8993444FF90BF6CFC441280FB34F821" box="[1010,1232,1927,1951]" italics="true" pageId="5" pageNumber="74">X. robusta hoffmani</emphasis>
</taxonomicName>
by
<bibRefCitation id="AE7C95A7FF90BF6CFB4F128FFE86F87C" author="Vandenspiegel" pageId="5" pageNumber="74" refString="Vandenspiegel, D. &amp; Pierrard, G. (2004) Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Annales du Musee d'Afrique Centrale (Zoology), 288, 31 - 46." type="journal article" year="2004">Vandenspiegel &amp; Pierrard (2004)</bibRefCitation>
. At the same place where
<taxonomicName id="0DED93D5FF90BF6CFD3212A4FD12F87D" box="[644,758,1963,1987]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="pelecys">
<emphasis id="F8993444FF90BF6CFD3212A4FD12F87D" box="[644,758,1963,1987]" italics="true" pageId="5" pageNumber="74">X. pelecys</emphasis>
</taxonomicName>
has an axehead-shaped process,
<taxonomicName id="0DED93D5FF90BF6CFBD212A3FB2CF87D" box="[1124,1224,1963,1987]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="hoffmani">
<emphasis id="F8993444FF90BF6CFBD212A3FB2CF87D" box="[1124,1224,1963,1987]" italics="true" pageId="5" pageNumber="74">hoffmani</emphasis>
</taxonomicName>
has a hook-shaped process. The basal telomeric
<taxonomicName id="0DED93D5FF90BF6CFE6B12C7FDD4F859" box="[477,560,1999,2023]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lamella">lamella</taxonomicName>
seen in
<taxonomicName id="0DED93D5FF90BF6CFD3A12D8FD1BF859" box="[652,767,1999,2023]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="pelecys">
<emphasis id="F8993444FF90BF6CFD3A12D8FD1BF859" box="[652,767,1999,2023]" italics="true" pageId="5" pageNumber="74">X. pelecys</emphasis>
</taxonomicName>
is lacking in
<taxonomicName id="0DED93D5FF90BF6CFC2012C7FC1DF859" box="[918,1017,1999,2023]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="hoffmani">
<emphasis id="F8993444FF90BF6CFC2012C7FC1DF859" box="[918,1017,1999,2023]" italics="true" pageId="5" pageNumber="74">hoffmani</emphasis>
</taxonomicName>
, which instead has a serrated
<taxonomicName id="0DED93D5FF90BF6CFAFC12C7FA78F859" box="[1354,1436,1999,2023]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="lamella">lamella</taxonomicName>
situated medially on the lower side (inside the loop) running the length of the telomere. The metaplica of
<taxonomicName id="0DED93D5FF93BF6FFA9D1591FA78FF0E" box="[1323,1436,152,176]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="pelecys">
<emphasis id="F8993444FF93BF6FFA9D1591FA78FF0E" box="[1323,1436,152,176]" italics="true" pageId="6" pageNumber="75">X. pelecys</emphasis>
</taxonomicName>
is a bit simpler than that of X.
<taxonomicName id="0DED93D5FF93BF6FFDBA15B4FD8BFF6A" box="[524,623,188,212]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="hoffmani">
<emphasis id="F8993444FF93BF6FFDBA15B4FD8BFF6A" box="[524,623,188,212]" italics="true" pageId="6" pageNumber="75">hoffmani</emphasis>
</taxonomicName>
, having no triangular distolateral metaplical flange (
<bibRefCitation id="AE7C95A7FF93BF6FFB6015B4FDB6FF49" author="Vandenspiegel" pageId="6" pageNumber="75" refString="Vandenspiegel, D. &amp; Pierrard, G. (2004) Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Annales du Musee d'Afrique Centrale (Zoology), 288, 31 - 46." type="journal article" year="2004">Vandenspiegel &amp; Pierrard 2004: “projection dentiforme”</bibRefCitation>
) by the lateral metaplical process nor any rounded mesolateral metaplical flange (
<bibRefCitation id="AE7C95A7FF93BF6FFF58140CFC8BFEA2" author="Vandenspiegel" box="[238,879,260,284]" pageId="6" pageNumber="75" refString="Vandenspiegel, D. &amp; Pierrard, G. (2004) Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Annales du Musee d'Afrique Centrale (Zoology), 288, 31 - 46." type="journal article" year="2004">Vandenspiegel &amp; Pierrard 2004: “projection lamellaire”</bibRefCitation>
) by the proplical lobe. Instead,
<taxonomicName id="0DED93D5FF93BF6FFB54140DFAB3FEA2" box="[1250,1367,260,284]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="pelecys">
<emphasis id="F8993444FF93BF6FFB54140DFAB3FEA2" box="[1250,1367,260,284]" italics="true" pageId="6" pageNumber="75">X. pelecys</emphasis>
</taxonomicName>
has a lateral metaplical spine situated between the proplical lobe and the lateral metaplical process.
</paragraph>
<paragraph id="CA52E856FF93BF6FFF711444FEE4FD82" blockId="6.[151,1436,151,572]" pageId="6" pageNumber="75">
Instead of regarding
<taxonomicName id="0DED93D5FF93BF6FFE021445FDC3FEDA" box="[436,551,332,356]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="pelecys">
<emphasis id="F8993444FF93BF6FFE021445FDC3FEDA" box="[436,551,332,356]" italics="true" pageId="6" pageNumber="75">X. pelecys</emphasis>
</taxonomicName>
as another subspecies of
<emphasis id="F8993444FF93BF6FFCFB1445FC2DFEDA" box="[845,969,332,356]" italics="true" pageId="6" pageNumber="75">
<taxonomicName id="0DED93D5FF93BF6FFCFB1445FC21FEDA" box="[845,965,332,356]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="robusta">X. robusta</taxonomicName>
,
</emphasis>
due to its similarity especially with
<emphasis id="F8993444FF93BF6FFADA1445FEE5FE36" italics="true" pageId="6" pageNumber="75">
<taxonomicName id="0DED93D5FF93BF6FFADA1445FF1AFE36" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="robusta" subSpecies="hoffmani">X. r. hoffmani</taxonomicName>
,
</emphasis>
we treat it as a full species. At the same time, we formally elevate
<taxonomicName id="0DED93D5FF93BF6FFC431479FEA4FE12" authority="Vandenspiegel &amp; Pierrard, 2004" authorityName="Vandenspiegel &amp; Pierrard" authorityYear="2004" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="subSpecies" species="robusta" subSpecies="hoffmani">
<emphasis id="F8993444FF93BF6FFC431479FB31FE36" box="[1013,1237,368,392]" italics="true" pageId="6" pageNumber="75">X. robusta hoffmani</emphasis>
<bibRefCitation id="AE7C95A7FF93BF6FFB6A1467FEA4FE12" author="Vandenspiegel" pageId="6" pageNumber="75" refString="Vandenspiegel, D. &amp; Pierrard, G. (2004) Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Annales du Musee d'Afrique Centrale (Zoology), 288, 31 - 46." type="journal article" year="2004">Vandenspiegel &amp; Pierrard, 2004</bibRefCitation>
</taxonomicName>
, to a full species:
<taxonomicName id="0DED93D5FF93BF6FFD93149DFBD2FE12" authority="Vandenspiegel &amp; Pierrard, 2004" authorityName="Vandenspiegel &amp; Pierrard" authorityYear="2004" box="[549,1078,404,428]" class="Diplopoda" family="Odontopygidae" genus="Xystopyge" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" species="hoffmani" status="stat. nov.">
<emphasis id="F8993444FF93BF6FFD93149DFD49FE12" box="[549,685,404,428]" italics="true" pageId="6" pageNumber="75">X. hoffmani</emphasis>
<bibRefCitation id="AE7C95A7FF93BF6FFD0D149CFBD2FE12" author="Vandenspiegel" box="[699,1078,404,428]" pageId="6" pageNumber="75" refString="Vandenspiegel, D. &amp; Pierrard, G. (2004) Revision du genre Xystopyge (Odontopygidae, Diplopoda) et description de nouvelles especes et sous-especes d'Afrique de l'Est. Annales du Musee d'Afrique Centrale (Zoology), 288, 31 - 46." type="journal article" year="2004">Vandenspiegel &amp; Pierrard, 2004</bibRefCitation>
</taxonomicName>
,
<emphasis id="F8993444FF93BF6FFBF1149CFB74FE12" bold="true" box="[1095,1168,404,428]" pageId="6" pageNumber="75">
<taxonomicNameLabel id="E3AA893FFF93BF6FFBF1149CFB74FE12" box="[1095,1168,404,428]" pageId="6" pageNumber="75">n.stat.</taxonomicNameLabel>
</emphasis>
We adopt this splitter approach because 1) we do not recommend use of the subspecies category in a group as poorly known as
<taxonomicName id="0DED93D5FF93BF6FFF2114D4FEA3FE4A" box="[151,327,476,500]" class="Diplopoda" family="Odontopygidae" kingdom="Animalia" order="Spirostreptida" pageId="6" pageNumber="75" phylum="Arthropoda" rank="family">Odontopygidae</taxonomicName>
, 2) recent studies have shown that even very subtle morphological differences between millipede populations may be linked to considerable genetic distances (e.g.,
<bibRefCitation id="AE7C95A7FF93BF6FFC0114F7FB2AFDA9" author="Frederiksen" box="[951,1230,511,535]" pageId="6" pageNumber="75" refString="Frederiksen, S., Petersen, G. &amp; Enghoff, H. (2012) How many species are there of Pachyiulus? A contribution to the taxonomy of Europe's largest millipedes (Diplopoda, Julida, Julidae). Journal of Natural History, 46, 599 - 611." type="journal article" year="2012">Frederiksen et al. 2012</bibRefCitation>
, Pimvichai et al. 2011a,b).
</paragraph>
</subSubSection>
</treatment>
</document>