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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF9AFFBBB49C1F1D7DCB2262" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax annexus" docType="treatment" docVersion="4" lastPageNumber="97" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="94" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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<mods:title>PeerJ</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
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<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102465" ID-GBIF-Taxon="183542589" ID-Zenodo-Dep="5102465" LSID="urn:lsid:plazi:treatment:054FDB70FF9AFFBBB49C1F1D7DCB2262" httpUri="http://treatment.plazi.org/id/054FDB70FF9AFFBBB49C1F1D7DCB2262" lastPageId="97" lastPageNumber="97" pageId="93" pageNumber="94">
<subSubSection box="[498,1047,1280,1307]" pageId="93" pageNumber="94" type="nomenclature">
<paragraph blockId="93.[498,1047,1280,1347]" box="[498,1047,1280,1307]" pageId="93" pageNumber="94">
<heading box="[498,1047,1280,1307]" level="2" pageId="93" pageNumber="94">
<taxonomicName authority="(Baroni Urbani, 1978)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,1047,1280,1307]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="annexus">
<emphasis bold="true" box="[498,1047,1280,1307]" pageId="93" pageNumber="94">
<emphasis bold="true" box="[498,768,1280,1307]" italics="true" pageId="93" pageNumber="94">Temnothorax annexus</emphasis>
(
<bibRefCitation author="Baroni Urbani C." box="[787,1037,1280,1307]" journalOrPublisher="Entomologica Basiliensia" pageId="93" pageNumber="94" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis bold="true" box="[787,1037,1280,1307]" italics="true" pageId="93" pageNumber="94">Baroni Urbani, 1978</emphasis>
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,954,1320,1347]" pageId="93" pageNumber="94" type="distribution">
<paragraph blockId="93.[498,1047,1280,1347]" box="[498,954,1320,1347]" pageId="93" pageNumber="94">
Distribution:
<figureCitation box="[657,756,1320,1347]" captionStart="Figure 95" captionStartId="92.[116,181,1124,1146]" captionTargetBox="[90,1541,234,1111]" captionTargetId="figure-210@92.[90,1524,234,1096]" captionTargetPageId="92" captionText="Figure 95 Geographical and elevational distribution of the annexus group. (A) Temnothorax annexus (B) T. arbustus sp. nov. (C) T. balnearius sp. nov. (D) T. obtusigaster sp. nov. (E) T. quercicola sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-95" figureDoi="http://doi.org/10.5281/zenodo.5102275" httpUri="https://zenodo.org/record/5102275/files/figure.png" pageId="93" pageNumber="94">Fig. 95A</figureCitation>
; worker:
<figureCitation box="[869,948,1320,1346]" captionStart="Figure 96" captionStartId="93.[524,589,1174,1196]" captionTargetBox="[498,1541,234,1161]" captionTargetId="figure-131@93.[498,1295,234,1146]" captionTargetPageId="93" captionText="Figure 96 Temnothorax annexus lectotype worker (MCZENT00561747). (A) Profile view. (B) Fullface view. (C) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-96" figureDoi="http://doi.org/10.5281/zenodo.5102277" httpUri="https://zenodo.org/record/5102277/files/figure.png" pageId="93" pageNumber="94">Fig. 96</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="93" pageNumber="94" type="reference_group">
<paragraph blockId="93.[498,1486,1383,1649]" pageId="93" pageNumber="94">
<treatmentCitationGroup pageId="93" pageNumber="94">
<treatmentCitation author="Wheeler WM" box="[498,995,1383,1410]" page="11" pageId="93" pageNumber="94" year="1931">
<taxonomicName authority="Wheeler, 1931: 11" authorityName="Wheeler" authorityPageNumber="11" authorityYear="1931" box="[498,995,1383,1410]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="annectens">
<emphasis box="[498,950,1383,1409]" italics="true" pageId="93" pageNumber="94">
Macromischa annectens
<bibRefCitation author="Wheeler WM" box="[785,995,1383,1410]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="93" pageNumber="94" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">Wheeler, 1931: 11</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[1008,1100,1383,1409]" pageId="93" pageNumber="94">Syntype</typeStatus>
workers. Cuernavaca,
<collectingCountry box="[1369,1458,1383,1409]" name="Mexico" pageId="93" pageNumber="94">Mexico</collectingCountry>
. One worker here designated
<emphasis bold="true" box="[839,948,1423,1449]" pageId="93" pageNumber="94">
<typeStatus box="[839,948,1423,1449]" pageId="93" pageNumber="94">lectotype</typeStatus>
</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="93.[498,1486,1383,1649]" pageId="93" pageNumber="94">
<treatmentCitationGroup pageId="93" pageNumber="94">
<treatmentCitation author="Baroni Urbani C." box="[498,1052,1463,1489]" page="421" pageId="93" pageNumber="94" year="1978">
<taxonomicName authority="Baroni Urbani, 1978: 421" authorityName="Baroni Urbani" authorityPageNumber="421" authorityYear="1978" box="[498,1052,1463,1489]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[498,992,1463,1489]" italics="true" pageId="93" pageNumber="94">
Leptothorax annexus
<bibRefCitation author="Baroni Urbani C." box="[751,1052,1463,1489]" journalOrPublisher="Entomologica Basiliensia" pageId="93" pageNumber="94" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<author box="[751,924,1463,1489]" case="TC" fns="*" inm="true" npo="*" pageId="93" pageNumber="94">Baroni Urbani</author>
,
<year box="[937,992,1463,1489]" pageId="93" pageNumber="94">1978</year>
: 421
</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
. Transferred to
<taxonomicName authority="Mayr." authorityName="Mayr." box="[1242,1382,1463,1489]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis box="[1242,1382,1463,1489]" italics="true" pageId="93" pageNumber="94">Leptothorax</emphasis>
</taxonomicName>
. Nomen novum for
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[632,1024,1502,1529]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="annectens">
<emphasis box="[632,894,1502,1528]" italics="true" pageId="93" pageNumber="94">Leptothorax annectens</emphasis>
(Wheeler)
</taxonomicName>
: 11 (nec
<treatmentCitation author="Wheeler WM" page="242" pageId="93" pageNumber="94" year="1903">
<taxonomicName authority="Wheeler, 1903 b: 242" authorityName="Wheeler" authorityPageNumber="242" authorityYear="1903" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="curvispinosus" subSpecies="annectens">
<emphasis box="[1132,1434,1502,1528]" italics="true" pageId="93" pageNumber="94">Leptothorax curvispinosus</emphasis>
ssp.
<emphasis box="[498,799,1542,1568]" italics="true" pageId="93" pageNumber="94">
annectens
<bibRefCitation author="Wheeler WM" box="[621,862,1542,1569]" journalOrPublisher="Proceedings of the Academy of Natural Sciences of: Philadelphia" pageId="93" pageNumber="94" pagination="215 - 260" refId="ref273272" refString="Wheeler WM. 1903 b. A revision of the North American ants of the genus Leptothorax Mayr. In: Proceedings of the Academy of Natural Sciences of: Philadelphia. 215 - 260." title="A revision of the North American ants of the genus Leptothorax Mayr" type="book chapter" year="1903">
<author box="[621,718,1542,1568]" fns="*" inm="true" npo="*" pageId="93" pageNumber="94">Wheeler</author>
,
<year box="[731,787,1542,1568]" pageId="93" pageNumber="94">1903</year>
b: 242
</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="93.[498,1486,1383,1649]" pageId="93" pageNumber="94">
<treatmentCitationGroup pageId="93" pageNumber="94">
<taxonomicName authority="(Baroni Urbani)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,961,1582,1609]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[498,760,1582,1608]" italics="true" pageId="93" pageNumber="94">Temnothorax annexus</emphasis>
(Baroni Urbani)
</taxonomicName>
:
<treatmentCitation author="Bolton B." box="[974,1179,1582,1609]" page="271" pageId="93" pageNumber="94" year="2003">
<bibRefCitation author="Bolton B." box="[974,1179,1582,1609]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="93" pageNumber="94" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
<emphasis box="[974,1119,1582,1608]" italics="true" pageId="93" pageNumber="94">
<collectingRegion box="[974,1052,1582,1608]" country="United Kingdom" name="Bolton" pageId="93" pageNumber="94">
<author box="[974,1052,1582,1608]" fns="*" inm="true" npo="*" pageId="93" pageNumber="94">Bolton</author>
</collectingRegion>
,
<year box="[1064,1119,1582,1608]" pageId="93" pageNumber="94">2003</year>
</emphasis>
: 271
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1622,1648]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis box="[498,655,1622,1648]" italics="true" pageId="93" pageNumber="94">Temnothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="93" pageNumber="94" type="materials_examined">
<paragraph blockId="93.[498,1542,1684,1871]" pageId="93" pageNumber="94">
<emphasis bold="true" box="[530,834,1684,1711]" pageId="93" pageNumber="94">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861491" collectingDate="1929-06-26" collectionCode="MCZC" collectorName="E. Skwarra " country="Mexico" location="Cuernavaca" pageId="93" pageNumber="94" specimenCount="1" specimenCount-worker="1" stateProvince="Morelos" typeStatus="lectotype">
<emphasis box="[843,1045,1685,1711]" italics="true" pageId="93" pageNumber="94">
<typeStatus box="[843,951,1685,1710]" pageId="93" pageNumber="94">Lectotype</typeStatus>
worker:
</emphasis>
<collectingCountry box="[1054,1169,1685,1711]" name="Mexico" pageId="93" pageNumber="94">MEXICO</collectingCountry>
:
<collectingRegion box="[1181,1280,1685,1711]" country="Mexico" name="Morelos" pageId="93" pageNumber="94">Morelos</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:054FDB70FF9AFFBBB49C1F1D7DCB2262:88393CBDFF9AFF87B0621C887CA521F8" box="[1292,1431,1685,1711]" country="Mexico" name="Cuernavaca" pageId="93" pageNumber="94" stateProvince="Morelos">Cuernavaca</location>
,
<date pageId="93" pageNumber="94" value="1929-06-26">
<collectingDate pageId="93" pageNumber="94" value="1929-06-26">26 June 1929</collectingDate>
</date>
,
<collectorName box="[569,696,1724,1751]" pageId="93" pageNumber="94">E. Skwarra</collectorName>
#Z867/Sk., ex
<taxonomicName box="[859,1099,1724,1750]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="species" species="circinnata">
<emphasis box="[859,1099,1724,1750]" italics="true" pageId="93" pageNumber="94">Tillandsia circinnata</emphasis>
</taxonomicName>
,
<specimenCount box="[1113,1217,1724,1751]" pageId="93" pageNumber="94" type="worker">1 worker</specimenCount>
(M.C.Z. co-type 2-3 16359, MCZENT00561747, bottom specimen on pin) [
<collectionCode box="[1059,1150,1764,1791]" pageId="93" pageNumber="94">MCZC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="93.[498,1542,1684,1871]" pageId="93" pageNumber="94">
<materialsCitation ID-GBIF-Occurrence="3321861522" box="[530,1421,1804,1831]" collectingDate="1929-06-26" collectionCode="MCZC" collectorName="Menozzi C" country="Mexico" location="Cuernavaca" pageId="93" pageNumber="94" specimenCount="1" specimenCount-worker="1" stateProvince="Morelos" typeStatus="paralectotype">
<emphasis box="[530,787,1804,1830]" italics="true" pageId="93" pageNumber="94">
<typeStatus box="[530,684,1804,1830]" pageId="93" pageNumber="94">Paralectotype</typeStatus>
workers:
</emphasis>
same pin as
<typeStatus box="[933,1040,1804,1830]" pageId="93" pageNumber="94">lectotype</typeStatus>
,
<specimenCount box="[1050,1152,1804,1830]" pageId="93" pageNumber="94" type="worker">1 worker</specimenCount>
(top specimen on pin);
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861496" collectingDate="1929-06-26" collectionCode="MCZC" collectorName="Menozzi C" country="Mexico" location="Cuernavaca" pageId="93" pageNumber="94" specimenCount="1" specimenCount-worker="1" stateProvince="Morelos" typeStatus="paralectotype">
same data as lectotype, 2 teneral workers (
<collectorName box="[869,927,1844,1870]" name="Menozzi C" pageId="93" pageNumber="94">M.C.</collectorName>
Z. co-type 4-5 16359, MCZENT00561746) [
<collectionCode box="[1441,1532,1843,1870]" pageId="93" pageNumber="94">MCZC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="94" pageNumber="95" type="distribution">
<paragraph blockId="94.[498,1542,234,1935]" pageId="94" pageNumber="95">
<emphasis bold="true" box="[530,753,234,260]" pageId="94" pageNumber="95">Geographic range:</emphasis>
South-Central
<collectingCountry box="[931,1019,234,260]" name="Mexico" pageId="94" pageNumber="95">Mexico</collectingCountry>
, mid elevations of the Southern Sierra Madre (
<figureCitation box="[508,608,274,301]" captionStart="Figure 95" captionStartId="92.[116,181,1124,1146]" captionTargetBox="[90,1541,234,1111]" captionTargetId="figure-210@92.[90,1524,234,1096]" captionTargetPageId="92" captionText="Figure 95 Geographical and elevational distribution of the annexus group. (A) Temnothorax annexus (B) T. arbustus sp. nov. (C) T. balnearius sp. nov. (D) T. obtusigaster sp. nov. (E) T. quercicola sp. nov. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-95" figureDoi="http://doi.org/10.5281/zenodo.5102275" httpUri="https://zenodo.org/record/5102275/files/figure.png" pageId="94" pageNumber="95">Fig. 95A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="94" pageNumber="95" type="diagnosis">
<paragraph blockId="94.[498,1542,234,1935]" pageId="94" pageNumber="95">
<emphasis bold="true" box="[530,752,314,340]" pageId="94" pageNumber="95">Worker diagnosis:</emphasis>
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[759,1018,314,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[759,1018,314,340]" italics="true" pageId="94" pageNumber="95">Temnothorax annexus</emphasis>
</taxonomicName>
can be separated from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[498,573,354,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[498,573,354,380]" italics="true" pageId="94" pageNumber="95">salvini</emphasis>
</taxonomicName>
clade by the following character combination: subdecumbent setae present on gula; metanotal groove absent; propodeum bearing erect setae; propodeum not depressed; propodeal spines short, slightly shorter than the propodeal declivity (PSI 2829); hind femora strongly incrassate (FI 338349); petiole with a moderately long peduncle: peduncle comprising about half of the total length of the petiole; petiolar node erect and subquadrate, not overhanging the caudal cylinder; postpetiole narrow (PWI 169173); first gastral tergite smooth and shining; integument testaceous; dorsum of head, mesosoma, waist segments and gaster with erect, short, blunt-tipped setae; legs without erect setae.
</paragraph>
<paragraph blockId="94.[498,1542,234,1935]" pageId="94" pageNumber="95">
<emphasis bold="true" box="[530,719,672,698]" pageId="94" pageNumber="95">Similar species:</emphasis>
<taxonomicName authority="(Mackay)" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[727,1128,673,700]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="andersoni">
<emphasis box="[727,1006,673,699]" italics="true" pageId="94" pageNumber="95">Temnothorax andersoni</emphasis>
(Mackay)
</taxonomicName>
,
<taxonomicName authority="(Emery)" baseAuthorityName="Emery" box="[1141,1356,673,699]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="andrei">
<emphasis box="[1141,1248,673,699]" italics="true" pageId="94" pageNumber="95">T. andrei</emphasis>
(Emery)
</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1369,1479,673,698]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">
<emphasis box="[1369,1479,673,698]" italics="true" pageId="94" pageNumber="95">T. aureus</emphasis>
</taxonomicName>
<taxonomicNameLabel pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[564,711,712,738]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecoides" status="sp. nov.">
<emphasis box="[564,711,712,738]" italics="true" pageId="94" pageNumber="95">T. aztecoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[720,814,713,739]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis box="[827,1099,713,738]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[827,945,713,738]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecus">T. aztecus</taxonomicName>
,
<taxonomicName authority="(Cole)" baseAuthorityName="Cole" box="[958,1182,712,739]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="carinatus">T. carinatus (Cole)</taxonomicName>
</emphasis>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1196,1341,713,738]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="casanovai" status="sp. nov.">
<emphasis box="[1196,1341,713,738]" italics="true" pageId="94" pageNumber="95">T. casanovai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1351,1444,713,739]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="(Mackay)" baseAuthorityName="Mackay" box="[498,813,752,779]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="cokendolpheri">
<emphasis box="[498,691,752,778]" italics="true" pageId="94" pageNumber="95">T. cokendolpheri</emphasis>
(Mackay)
</taxonomicName>
,
<emphasis box="[826,1063,753,778]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[826,948,753,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="goniops">T. goniops</taxonomicName>
,
<taxonomicName authority="(Emery)" baseAuthorityName="Emery" box="[961,1170,752,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="nitens">T. nitens (Emery)</taxonomicName>
</emphasis>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1183,1338,753,778]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paraztecus" status="sp. nov.">
<emphasis box="[1183,1338,753,778]" italics="true" pageId="94" pageNumber="95">T. paraztecus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1346,1440,753,779]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="(Mackay)" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[498,795,792,819]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="punctaticeps">
<emphasis box="[498,673,793,818]" italics="true" pageId="94" pageNumber="95">T. punctaticeps</emphasis>
(Mackay)
</taxonomicName>
,
<emphasis box="[808,1092,792,818]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[808,930,793,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="rugosus">T. rugosus</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[944,1092,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
</emphasis>
(light form),
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1256,1438,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tenuisculptus">
<emphasis box="[1256,1438,792,818]" italics="true" pageId="94" pageNumber="95">T. tenuisculptus</emphasis>
</taxonomicName>
,
<emphasis box="[498,749,832,858]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[498,636,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="terrigena">T. terrigena</taxonomicName>
,
<taxonomicName authority="Snelling et al." authorityName="Snelling" box="[650,917,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="wardi">T. wardi Snelling et al.</taxonomicName>
</emphasis>
, and species of the
<emphasis box="[1150,1353,832,858]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1150,1248,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">annexus</taxonomicName>
,
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1261,1353,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="fuscatus">fuscatus</taxonomicName>
</emphasis>
, and
<taxonomicName baseAuthorityName="Santschi" baseAuthorityYear="1908" box="[1422,1516,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="silvestrii">
<emphasis box="[1422,1516,832,858]" italics="true" pageId="94" pageNumber="95">silvestrii</emphasis>
</taxonomicName>
groups.
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[591,851,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[591,851,872,898]" italics="true" pageId="94" pageNumber="95">Temnothorax annexus</emphasis>
</taxonomicName>
can be separated from
<emphasis box="[1119,1541,872,898]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1119,1266,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="andersoni">T. andersoni</taxonomicName>
,
<taxonomicName box="[1277,1385,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1396,1541,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecoides" status="sp. nov.">T. aztecoides</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[498,592,912,938]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis box="[605,1527,912,938]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[605,723,912,937]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecus">T. aztecus</taxonomicName>
,
<taxonomicName box="[736,879,912,937]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="carinatus">T. carinatus</taxonomicName>
,
<taxonomicName box="[892,1087,912,938]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="cokendolpheri">T. cokendolpheri</taxonomicName>
,
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1100,1221,912,937]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="goniops">T. goniops</taxonomicName>
,
<taxonomicName box="[1235,1338,912,937]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="nitens">T. nitens</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1352,1490,912,937]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="terrigena">T. terrigena</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="wardi">
T.
<emphasis box="[498,565,951,977]" italics="true" pageId="94" pageNumber="95">wardi</emphasis>
</taxonomicName>
</emphasis>
, and species of the
<taxonomicName baseAuthorityName="Santschi" baseAuthorityYear="1908" box="[790,884,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="silvestrii">
<emphasis box="[790,884,951,977]" italics="true" pageId="94" pageNumber="95">silvestrii</emphasis>
</taxonomicName>
group by the strongly incrassate hind femora (FI&gt; 330). Furthermore, it can be distinguished from
<emphasis box="[1003,1435,991,1017]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1003,1152,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="andersoni">T. andersoni</taxonomicName>
,
<taxonomicName box="[1165,1275,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="andrei">T. andrei</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1287,1435,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecoides" status="sp. nov.">T. aztecoides</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[1444,1537,992,1018]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis box="[498,1381,1031,1057]" italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[498,616,1032,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecus">T. aztecus</taxonomicName>
,
<taxonomicName box="[630,772,1032,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="carinatus">T. carinatus</taxonomicName>
,
<taxonomicName box="[786,980,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="cokendolpheri">T. cokendolpheri</taxonomicName>
,
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[993,1115,1032,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="goniops">T. goniops</taxonomicName>
,
<taxonomicName box="[1128,1232,1032,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="nitens">T. nitens</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1245,1381,1032,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="terrigena">T. terrigena</taxonomicName>
</emphasis>
, and the
<taxonomicName baseAuthorityName="Santschi" baseAuthorityYear="1908" box="[498,592,1071,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="silvestrii">
<emphasis box="[498,592,1071,1097]" italics="true" pageId="94" pageNumber="95">silvestrii</emphasis>
</taxonomicName>
group by the anterior clypeal margin, which is emarginate in
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1328,1457,1072,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1328,1457,1072,1097]" italics="true" pageId="94" pageNumber="95">T. annexus</emphasis>
</taxonomicName>
. The absence of erect, long, tapering setae on the body will separate
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1297,1426,1112,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1297,1426,1112,1137]" italics="true" pageId="94" pageNumber="95">T. annexus</emphasis>
</taxonomicName>
from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,646,1151,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecoides" status="sp. nov.">
<emphasis box="[498,646,1151,1177]" italics="true" pageId="94" pageNumber="95">T. aztecoides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[655,748,1151,1177]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1931" box="[760,876,1152,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="aztecus">
<emphasis box="[760,876,1152,1177]" italics="true" pageId="94" pageNumber="95">T. aztecus</emphasis>
</taxonomicName>
, and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[943,1098,1152,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="paraztecus" status="sp. nov.">
<emphasis box="[943,1098,1152,1177]" italics="true" pageId="94" pageNumber="95">T. paraztecus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1106,1199,1151,1177]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
, while the erect, subquadrate petiolar node differentiates
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[815,941,1191,1216]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[815,941,1191,1216]" italics="true" pageId="94" pageNumber="95">T. annexus</emphasis>
</taxonomicName>
from the
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1053,1145,1191,1217]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[1053,1145,1191,1217]" italics="true" pageId="94" pageNumber="95">fuscatus</emphasis>
</taxonomicName>
group,
<emphasis italics="true" pageId="94" pageNumber="95">
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1233,1408,1191,1216]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="punctaticeps">T. punctaticeps</taxonomicName>
,
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1418,1537,1191,1216]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="rugosus">T. rugosus</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[498,647,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
</emphasis>
, and nearly all other species listed above. To separate
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1290,1419,1231,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1290,1419,1231,1256]" italics="true" pageId="94" pageNumber="95">T. annexus</emphasis>
</taxonomicName>
from other members of the
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[762,858,1271,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[762,858,1271,1296]" italics="true" pageId="94" pageNumber="95">annexus</emphasis>
</taxonomicName>
group, look for the following character combination: a continuous dorsal margin of the mesosoma in profile (propodeum not depressed as in
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[498,681,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tenuisculptus">
<emphasis box="[498,681,1350,1376]" italics="true" pageId="94" pageNumber="95">T. tenuisculptus</emphasis>
</taxonomicName>
); decumbent setae on the gular region (
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1148,1278,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="arbustus" status="sp. nov.">
<emphasis box="[1148,1278,1350,1376]" italics="true" pageId="94" pageNumber="95">T. arbustus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1285,1377,1351,1377]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1390,1541,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="balnearius" status="sp. nov.">
<emphasis box="[1390,1541,1350,1376]" italics="true" pageId="94" pageNumber="95">T. balnearius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[498,592,1390,1416]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[657,827,1390,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="obtusigaster" status="sp. nov.">
<emphasis box="[657,827,1390,1416]" italics="true" pageId="94" pageNumber="95">T. obtusigaster</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[836,927,1390,1416]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
have erect setae); propodeal spines slightly shorter than the propodeal declivity (PSI ~ 29), pedunculate petiole, with the peduncle comprising about half the total petiole length, and heavily sculptured, predominantly areolate-rugose head and mesosoma will all distinguish
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[971,1099,1510,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[971,1099,1510,1535]" italics="true" pageId="94" pageNumber="95">T. annexus</emphasis>
</taxonomicName>
from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1174,1323,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="quercicola" status="sp. nov.">
<emphasis box="[1174,1323,1509,1535]" italics="true" pageId="94" pageNumber="95">T. quercicola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1332,1424,1510,1536]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="96" lastPageNumber="97" pageId="94" pageNumber="95" type="description">
<paragraph blockId="94.[498,1542,234,1935]" lastBlockId="95.[498,1542,234,1934]" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95">
<emphasis bold="true" box="[530,1018,1549,1576]" pageId="94" pageNumber="95">
Worker measurements &amp; indices (
<emphasis bold="true" box="[941,957,1550,1576]" italics="true" pageId="94" pageNumber="95">n</emphasis>
= 3):
</emphasis>
SL =
<geoCoordinate box="[1084,1146,1549,1576]" degrees="0.759" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.759">0.759</geoCoordinate>
<geoCoordinate box="[1146,1223,1549,1575]" degrees="0.766" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.766">0.766</geoCoordinate>
(0.762); FRS =
<geoCoordinate box="[1402,1464,1549,1575]" degrees="0.261" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.261">0.261</geoCoordinate>
<geoCoordinate box="[1464,1541,1549,1575]" degrees="0.302" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.302">0.302</geoCoordinate>
(0.284); CW =
<geoCoordinate box="[678,740,1589,1615]" degrees="0.891" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.891">0.891</geoCoordinate>
<geoCoordinate box="[740,817,1589,1615]" degrees="0.917" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.917">0.917</geoCoordinate>
(0.904); CWb =
<geoCoordinate box="[1020,1082,1589,1615]" degrees="0.778" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.778">0.778</geoCoordinate>
<geoCoordinate box="[1082,1159,1589,1615]" degrees="0.826" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.826">0.826</geoCoordinate>
(0.807); PoOC =
<geoCoordinate box="[1372,1434,1589,1616]" degrees="0.350" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.35">0.350</geoCoordinate>
<geoCoordinate box="[1434,1511,1589,1615]" degrees="0.378" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.378">0.378</geoCoordinate>
(0.367); CL =
<geoCoordinate box="[665,727,1629,1655]" degrees="0.947" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.947">0.947</geoCoordinate>
<geoCoordinate box="[727,804,1629,1655]" degrees="0.993" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.993">0.993</geoCoordinate>
(0.973); EL =
<geoCoordinate box="[977,1039,1629,1655]" degrees="0.218" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.218">0.218</geoCoordinate>
<geoCoordinate box="[1039,1116,1629,1655]" degrees="0.238" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.238">0.238</geoCoordinate>
(0.226); EW =
<geoCoordinate box="[1302,1364,1629,1655]" degrees="0.163" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.163">0.163</geoCoordinate>
<geoCoordinate box="[1364,1441,1629,1655]" degrees="0.182" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.182">0.182</geoCoordinate>
(0.173); MD =
<geoCoordinate box="[580,642,1669,1695]" degrees="0.236" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.236">0.236</geoCoordinate>
<geoCoordinate box="[642,719,1669,1695]" degrees="0.262" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.262">0.262</geoCoordinate>
(0.245); WL =
<geoCoordinate box="[904,966,1669,1695]" degrees="1.304" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="1.304">1.304</geoCoordinate>
<geoCoordinate box="[966,1043,1669,1696]" degrees="1.395" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-1.395">1.395</geoCoordinate>
(1.345); SPST =
<geoCoordinate box="[1245,1307,1669,1695]" degrees="0.361" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.361">0.361</geoCoordinate>
<geoCoordinate box="[1308,1385,1669,1695]" degrees="0.408" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.408">0.408</geoCoordinate>
(0.387); MPST =
<geoCoordinate box="[607,669,1709,1735]" degrees="0.433" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.433">0.433</geoCoordinate>
<geoCoordinate box="[669,746,1708,1735]" degrees="0.435" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.435">0.435</geoCoordinate>
(0.434); PEL =
<geoCoordinate box="[935,997,1708,1735]" degrees="0.505" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.505">0.505</geoCoordinate>
<geoCoordinate box="[997,1074,1708,1735]" degrees="0.529" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.529">0.529</geoCoordinate>
(0.514); NOL =
<geoCoordinate box="[1274,1336,1709,1735]" degrees="0.299" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.299">0.299</geoCoordinate>
<geoCoordinate box="[1336,1413,1709,1735]" degrees="0.313" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.313">0.313</geoCoordinate>
(0.304); NOH =
<geoCoordinate box="[598,660,1748,1775]" degrees="0.185" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.185">0.185</geoCoordinate>
<geoCoordinate box="[660,737,1749,1775]" degrees="0.204" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.204">0.204</geoCoordinate>
(0.198); PEH =
<geoCoordinate box="[933,995,1749,1775]" degrees="0.322" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.322">0.322</geoCoordinate>
<geoCoordinate box="[995,1072,1749,1775]" degrees="0.344" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.344">0.344</geoCoordinate>
(0.336); PPL =
<geoCoordinate box="[1262,1324,1749,1775]" degrees="0.229" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.229">0.229</geoCoordinate>
<geoCoordinate box="[1324,1401,1749,1775]" degrees="0.248" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.248">0.248</geoCoordinate>
(0.241); PW =
<geoCoordinate box="[577,639,1788,1815]" degrees="0.572" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.572">0.572</geoCoordinate>
<geoCoordinate box="[640,716,1788,1814]" degrees="0.622" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.622">0.622</geoCoordinate>
(0.590); SBPA =
<geoCoordinate box="[925,987,1788,1814]" degrees="0.243" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.243">0.243</geoCoordinate>
<geoCoordinate box="[987,1064,1788,1814]" degrees="0.290" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.29">0.290</geoCoordinate>
(0.272); SPTI =
<geoCoordinate box="[1263,1325,1788,1814]" degrees="0.329" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.329">0.329</geoCoordinate>
<geoCoordinate box="[1325,1402,1788,1814]" degrees="0.340" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.34">0.340</geoCoordinate>
(0.333); PEW =
<geoCoordinate box="[594,656,1828,1854]" degrees="0.230" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.23">0.230</geoCoordinate>
<geoCoordinate box="[656,733,1828,1854]" degrees="0.234" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.234">0.234</geoCoordinate>
(0.233); PNW =
<geoCoordinate box="[940,1002,1828,1855]" degrees="0.250" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.25">0.250</geoCoordinate>
<geoCoordinate box="[1002,1079,1828,1854]" degrees="0.273" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.273">0.273</geoCoordinate>
(0.260); PPW =
<geoCoordinate box="[1281,1343,1828,1854]" degrees="0.389" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.389">0.389</geoCoordinate>
<geoCoordinate box="[1344,1421,1828,1855]" degrees="0.405" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.405">0.405</geoCoordinate>
(0.400); HFL =
<geoCoordinate box="[586,648,1868,1895]" degrees="0.835" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.835">0.835</geoCoordinate>
<geoCoordinate box="[648,725,1868,1895]" degrees="0.850" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.85">0.850</geoCoordinate>
(0.841); HFWmax =
<geoCoordinate box="[981,1043,1868,1895]" degrees="0.251" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.251">0.251</geoCoordinate>
<geoCoordinate box="[1044,1121,1868,1894]" degrees="0.274" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.274">0.274</geoCoordinate>
(0.266); HFWmin =
<geoCoordinate box="[1374,1436,1868,1894]" degrees="0.072" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.072">0.072</geoCoordinate>
<geoCoordinate box="[1437,1514,1868,1894]" degrees="0.081" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.081">0.081</geoCoordinate>
(0.078); CS =
<geoCoordinate box="[660,722,1908,1935]" degrees="1.252" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="1.252">1.252</geoCoordinate>
<geoCoordinate box="[722,799,1908,1934]" degrees="1.323" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-1.323">1.323</geoCoordinate>
(1.294); ES =
<geoCoordinate box="[966,1028,1908,1934]" degrees="0.300" direction="north" orientation="latitude" pageId="94" pageNumber="95" precision="55" value="0.3">0.300</geoCoordinate>
<geoCoordinate box="[1029,1106,1908,1934]" degrees="0.329" direction="west" orientation="longitude" pageId="94" pageNumber="95" precision="55" value="-0.329">0.329</geoCoordinate>
(0.312); SI = 9298 (94); OI = 2325 (24); CI = 8284 (83); WLI = 163169 (167); SBI = 3037 (34); PSI = 2829 (29); PWI = 169173 (172); PLI = 206221 (213); NI = 147162 (154); PNWI = 107117 (112); NLI = 59; FI = 338349 (342).
</paragraph>
<paragraph blockId="95.[498,1542,234,1934]" pageId="95" pageNumber="96">
<emphasis bold="true" box="[530,775,353,380]" pageId="95" pageNumber="96">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 8284). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin emarginated medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 9298). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment about one and a half times as long as the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about one and a half times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin weakly emarginate medially but rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="95.[498,1542,234,1934]" pageId="95" pageNumber="96">
In profile view, compound eyes ovular and moderately large (OI 2325), with 13 ommatidia in longest row. Pronotal declivity distinct: dorsal margin of anterior face of pronotum marked by a carina; neck and anterior face of pronotum forming a ~120
<emphasis box="[1456,1466,947,965]" italics="true" pageId="95" pageNumber="96">
<superScript attach="left" box="[1456,1466,947,965]" fontSize="8" pageId="95" pageNumber="96">°</superScript>
</emphasis>
angle. Mesosoma very weakly convex from where it joins the pronotal declivity to the propodeal spines, nearly flat. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is moderately well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about four spiracle diameters. Propodeal spines moderately well developed and moderately long (PSI 2829), about as long as the propodeal declivity, flared at the base, downcurved, and blunt. Propodeal declivity weakly concave, forming a rounded ~110
<emphasis box="[1349,1359,1345,1363]" italics="true" pageId="95" pageNumber="96">
<superScript attach="left" box="[1349,1359,1345,1363]" fontSize="8" pageId="95" pageNumber="96">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion halfway to the propodeal spiracle. Petiole moderately long (PLI 206221), with weakly developed tubercles anterodorsally. Subpetiolar process in the form of a small, blunt tooth; ventral margin of petiole flat posterior to it. Petiolar peduncle moderately long: comprising about half of the total petiole length. Petiolar node robust, erect, and subquadrate: transition between peduncle and node marked by a rounded angle of ~120
<emphasis box="[1248,1258,1624,1642]" italics="true" pageId="95" pageNumber="96">
<superScript attach="none" box="[1248,1258,1624,1642]" fontSize="8" pageId="95" pageNumber="96">°</superScript>
</emphasis>
; anterior face forming a ~110
<emphasis box="[557,567,1664,1682]" italics="true" pageId="95" pageNumber="96">
<superScript attach="left" box="[557,567,1664,1682]" fontSize="8" pageId="95" pageNumber="96">°</superScript>
</emphasis>
angle with the dorsal face, which is weakly convex, nearly flat; dorsal face meeting the posterior face at a ~90
<emphasis box="[808,818,1704,1722]" italics="true" pageId="95" pageNumber="96">
<superScript attach="left" box="[808,818,1704,1722]" fontSize="8" pageId="95" pageNumber="96">°</superScript>
</emphasis>
angle, which forms a ~100
<emphasis box="[1141,1151,1704,1722]" italics="true" pageId="95" pageNumber="96">
<superScript attach="left" box="[1141,1151,1704,1722]" fontSize="8" pageId="95" pageNumber="96">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole flat anteriorly, bulging slightly anterodorsally before flattening posterodorsally; weakly lobed ventrally.
</paragraph>
<paragraph blockId="95.[498,1542,234,1934]" lastBlockId="96.[498,1542,234,1934]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">In dorsal view, humeri moderately well developed: evenly rounded and barely wider than the rest of the mesosoma; mesothoracic spiracles weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal suture represented by a disruption in the ground sculpture. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and weakly diverging apically, their apices separated from each other by slightly more than their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles not protruding past the lateral margins. Petiolar node nearly evenly ovular, but posterior margin flattened; node broader than the peduncle, and evenly grading into the caudal cylinder, which is the same width as the node. Postpetiole narrow (PWI 169173) and subquadrate. Anterior margin of the postpetiole flat and evenly rounds into the lateral margins, which diverge slightly to the rounded posterior corners; posterior margin flat. Metafemur strongly incrassate (FI 338349).</paragraph>
<paragraph blockId="96.[498,1542,234,1934]" pageId="96" pageNumber="97">Sculpture: median clypeal carina present, extending posteriorly to the frontal triangle, and flanked on either side by two equally strong carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture shining through weak areolae. Antennal scapes areolate. Cephalic dorsum areolate, with coarse rugae over the ground sculpture; concentric costulae surrounding the antennal insertions; overlying sculpture primarily costate between the frontal carinae. Lateral surfaces of head areolate, with coarse rugae over the ground sculpture. Ventral surface of head smooth and shining anteromedially, but otherwise weakly areolate-costulate. Pronotal neck areolate. Lateral surfaces mesosoma areolate with coarse rugae over the ground sculpture, but with longitudinal rugae stronger than transverse rugae. Propodeal declivity areolate-rugulose. Dorsal surface of mesosoma sculptured similarly to the lateral surface, but stronger longitudinal rugae restricted to the pronotum. Femora finely, densely areolate. Petiole uniformly areolate; a weak carina present laterally, extending longitudinally from the petiolar spiracle to the caudal cylinder; weak rugae on the dorsal and posterior faces of the node. Postpetiole uniformly areolate, with weak rugulose sculpture on the lateral faces. First gastral tergite areolate, but with sculpture becoming weaker posteriorly, without spectral iridescence. First gastral sternite smooth and shining, without spectral iridescence.</paragraph>
<paragraph blockId="96.[498,1542,234,1934]" pageId="96" pageNumber="97">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about half the width of the compound eye. The head bears ~18, mesosoma ~26, petiole 10, postpetiole ~16, and first gastral tergite ~20 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
<paragraph blockId="96.[498,1542,234,1934]" box="[530,1354,1589,1616]" pageId="96" pageNumber="97">Color: predominantly testaceous, with apex of mandibles dark brown.</paragraph>
<paragraph blockId="96.[498,1542,234,1934]" box="[530,734,1629,1655]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[530,601,1629,1655]" pageId="96" pageNumber="97">Gyne:</emphasis>
Unknown.
</paragraph>
<paragraph blockId="96.[498,1542,234,1934]" box="[530,730,1669,1695]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[530,597,1669,1695]" pageId="96" pageNumber="97">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="96" pageNumber="97" type="etymology">
<paragraph blockId="96.[498,1542,234,1934]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[530,668,1709,1735]" pageId="96" pageNumber="97">Etymology:</emphasis>
In the original description,
<bibRefCitation author="Wheeler WM" box="[989,1164,1709,1735]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="96" pageNumber="97" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[989,1164,1709,1735]" italics="true" pageId="96" pageNumber="97">Wheeler (1931)</emphasis>
</bibRefCitation>
states that this species, which he placed in
<taxonomicName authorityName="Roger" authorityYear="1863" box="[614,770,1748,1774]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis box="[614,770,1748,1774]" italics="true" pageId="96" pageNumber="97">Macromischa</emphasis>
</taxonomicName>
, nonetheless bears many features of the
<emphasis box="[1250,1383,1748,1774]" italics="true" pageId="96" pageNumber="97">rottenbergii</emphasis>
group of the Palearctic. Presumably annectens (= connected) from Greek refers to the implied link between these taxa.
</paragraph>
</subSubSection>
<subSubSection lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="discussion">
<paragraph blockId="96.[498,1542,234,1934]" lastBlockId="97.[498,1542,1147,1333]" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
<emphasis bold="true" box="[530,670,1868,1894]" pageId="96" pageNumber="97">Comments:</emphasis>
Known only from the
<typeStatus box="[945,995,1869,1895]" pageId="96" pageNumber="97">type</typeStatus>
series, which was collected from
<taxonomicName class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="96" pageNumber="97" phylum="Tracheophyta" rank="species" species="circinnata">
<emphasis italics="true" pageId="96" pageNumber="97">Tillandsia circinnata</emphasis>
</taxonomicName>
epiphytes near Cuernavaca,
<collectingRegion box="[955,1053,1908,1934]" country="Mexico" name="Morelos" pageId="96" pageNumber="97">Morelos</collectingRegion>
,
<collectingRegion box="[1066,1189,1908,1934]" country="Mexico" name="Distrito Federal" pageId="96" pageNumber="97">Mexico by</collectingRegion>
Elizabeth Skwarra.
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,758,1147,1173]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[498,758,1147,1173]" italics="true" pageId="97" pageNumber="98">Temnothorax annexus</emphasis>
</taxonomicName>
has close relatives in
<collectingRegion box="[1008,1183,1147,1173]" country="Mexico" name="Baja California" pageId="97" pageNumber="98">Baja California</collectingRegion>
, Arizona, and
<collectingRegion box="[1352,1434,1147,1173]" country="Mexico" name="Sonora" pageId="97" pageNumber="98">Sonora</collectingRegion>
that nest in dead vegetation, including live branches of oak trees. It is morphologically convergent with, but not particularly closely related to, members of the
<emphasis box="[1209,1342,1227,1253]" italics="true" pageId="97" pageNumber="98">rottenbergii</emphasis>
group of the Palearctic, or the
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[698,755,1267,1293]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="sallei">
<emphasis box="[698,755,1267,1293]" italics="true" pageId="97" pageNumber="98">sallei</emphasis>
</taxonomicName>
clade of southern North America, Central America, and the Greater Antilles (
<bibRefCitation author="Baroni Urbani C." box="[607,847,1306,1333]" journalOrPublisher="Entomologica Basiliensia" pageId="97" pageNumber="98" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis box="[607,847,1306,1333]" italics="true" pageId="97" pageNumber="98">Baroni Urbani, 1978</emphasis>
</bibRefCitation>
;
<bibRefCitation author="Fontenla Rizo JL" box="[862,1094,1306,1333]" journalOrPublisher="Avicennia" pageId="97" pageNumber="98" pagination="35 - 44" part="13" refId="ref270970" refString="Fontenla Rizo JL. 2000. Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax). Avicennia 13: 35 - 44." title="Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax)" type="journal article" year="2000">
<emphasis box="[862,1094,1306,1333]" italics="true" pageId="97" pageNumber="98">Fontenla Rizo, 2000</emphasis>
</bibRefCitation>
;
<bibRefCitation author="Prebus M." box="[1109,1257,1306,1333]" journalOrPublisher="BMC Evolutionary Biology" pageId="97" pageNumber="98" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">
<emphasis box="[1109,1257,1306,1333]" italics="true" pageId="97" pageNumber="98">Prebus, 2017</emphasis>
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>