treatments-xml/data/05/4F/DB/054FDB70FF93FF82B49C19BD7A6B253C.xml
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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF93FF82B49C19BD7A6B253C" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax acutispinosus Prebus 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="89" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="85" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PeerJ</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>462</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102442" ID-GBIF-Taxon="183542586" ID-Zenodo-Dep="5102442" LSID="urn:lsid:zoobank.org:act:05EA9F36-F838-4331-9601-58EC5393C237" httpUri="http://treatment.plazi.org/id/054FDB70FF93FF82B49C19BD7A6B253C" lastPageId="88" lastPageNumber="89" pageId="84" pageNumber="85">
<subSubSection box="[498,932,928,955]" pageId="84" pageNumber="85" type="nomenclature">
<paragraph blockId="84.[498,1117,928,1035]" box="[498,932,928,955]" pageId="84" pageNumber="85">
<heading box="[498,932,928,955]" level="2" pageId="84" pageNumber="85">
<emphasis bold="true" box="[498,932,928,955]" pageId="84" pageNumber="85">
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,828,928,955]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis bold="true" box="[498,828,928,955]" italics="true" pageId="84" pageNumber="85">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[838,932,929,955]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,862,969,995]" pageId="84" pageNumber="85" type="distribution">
<paragraph blockId="84.[498,1117,928,1035]" box="[498,862,969,995]" pageId="84" pageNumber="85">
Distribution &amp; worker:
<figureCitation box="[777,856,969,995]" captionStart="Figure 93" captionStartId="85.[524,589,964,986]" captionTargetBox="[498,1541,234,951]" captionTargetId="figure-289@85.[498,1534,234,936]" captionTargetPageId="85" captionText="Figure 93 Temnothorax acutispinosus sp. nov. holotype worker (JTLC000007447). (A) Profile view. (B) Full-face view. (C) Dorsal view. (D) Geographical and elevational distribution of specimens examined. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-93" figureDoi="http://doi.org/10.5281/zenodo.5102271" httpUri="https://zenodo.org/record/5102271/files/figure.png" pageId="84" pageNumber="85">Fig. 93</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[498,1117,1008,1035]" pageId="84" pageNumber="85" type="reference_group">
<paragraph blockId="84.[498,1117,928,1035]" box="[498,1117,1008,1035]" pageId="84" pageNumber="85">
<treatmentCitationGroup box="[498,1117,1008,1035]" pageId="84" pageNumber="85">
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1008,1034]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis box="[498,655,1008,1034]" italics="true" pageId="84" pageNumber="85">Temnothorax</emphasis>
</taxonomicName>
mmp06
<treatmentCitation author="Prebus M." box="[763,942,1008,1035]" page="7" pageId="84" pageNumber="85" year="2017">
<bibRefCitation author="Prebus M." box="[763,942,1008,1035]" journalOrPublisher="BMC Evolutionary Biology" pageId="84" pageNumber="85" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">
<emphasis box="[763,911,1008,1035]" italics="true" pageId="84" pageNumber="85">Prebus, 2017</emphasis>
: 7
</bibRefCitation>
</treatmentCitation>
. In phylogeny.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="materials_examined">
<paragraph blockId="84.[498,1542,1067,1935]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[530,834,1071,1097]" pageId="84" pageNumber="85">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861453" collectingDate="2008-05-19" collectionCode="CASC" collectorName="J. Longino" country="Mexico" elevation="1530" latitude="15.72196" location="Custepec" longLatPrecision="1" longitude="-92.95037" municipality="Custepec" pageId="84" pageNumber="85" specimenCode="6285, JTLC000007447" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis box="[842,1040,1071,1097]" italics="true" pageId="84" pageNumber="85">
<typeStatus box="[842,945,1071,1097]" pageId="84" pageNumber="85">Holotype</typeStatus>
<specimenCount box="[954,1034,1071,1097]" pageId="84" pageNumber="85" type="worker">worker</specimenCount>
:
</emphasis>
<collectingCountry box="[1049,1164,1071,1097]" name="Mexico" pageId="84" pageNumber="85">MEXICO</collectingCountry>
:
<collectingRegion box="[1175,1272,1071,1097]" country="Mexico" name="Chiapas" pageId="84" pageNumber="85">Chiapas</collectingRegion>
:
<collectingMunicipality box="[1284,1392,1071,1097]" pageId="84" pageNumber="85">Custepec</collectingMunicipality>
,
<geoCoordinate box="[1404,1542,1067,1098]" degrees="15.72196" direction="north" orientation="latitude" pageId="84" pageNumber="85" precision="1" value="15.72196">
15.72196
<emphasis box="[1508,1518,1067,1085]" italics="true" pageId="84" pageNumber="85">
<superScript attach="right" box="[1508,1518,1067,1085]" fontSize="8" pageId="84" pageNumber="85">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[498,642,1106,1138]" degrees="92.95037" direction="west" orientation="longitude" pageId="84" pageNumber="85" precision="1" value="-92.95037">
92.95037
<emphasis box="[603,613,1106,1124]" italics="true" pageId="84" pageNumber="85">
<superScript attach="right" box="[603,613,1106,1124]" fontSize="8" pageId="84" pageNumber="85">°</superScript>
</emphasis>
W
</geoCoordinate>
±
<quantity box="[677,738,1111,1138]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="84" pageNumber="85" unit="m" value="50.0">50 m</quantity>
,
<quantity box="[752,848,1111,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="84" pageNumber="85" unit="m" value="1530.0">
<elevation box="[752,848,1111,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="84" pageNumber="85" unit="m" value="1530.0">1,530 m</elevation>
</quantity>
,
<date box="[862,1018,1111,1138]" pageId="84" pageNumber="85" value="2008-05-19">
<collectingDate box="[862,1018,1111,1138]" pageId="84" pageNumber="85" value="2008-05-19">19 May 2008</collectingDate>
</date>
,
<collectorName box="[1031,1152,1111,1138]" pageId="84" pageNumber="85">J. Longino</collectorName>
<specimenCode box="[1162,1233,1111,1138]" pageId="84" pageNumber="85">#6285</specimenCode>
, mesophyll forest, under epiphytes in treefall (
<specimenCode box="[746,939,1151,1177]" collectionCode="JTLC" pageId="84" pageNumber="85">JTLC000007447</specimenCode>
) [
<collectionCode box="[962,1043,1150,1177]" pageId="84" pageNumber="85">CASC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="84.[498,1542,1067,1935]" pageId="84" pageNumber="85">
<materialsCitation ID-GBIF-Occurrence="3321861425" collectingDate="2008-05-19" collectionCode="MCZC, USNM, ECOSCE, LACM, UNAM, UCDC, FSCA, UVGC, AMNH, CASC" collectorName="J. Longino" country="Mexico" elevation="0" latitude="15.72196" location="Custepec" longLatPrecision="1" longitude="-92.95037" pageId="84" pageNumber="85" specimenCode="JTLC000007448, JTLC000007449, JTLC000007450" specimenCount="11" specimenCount-worker="11" stateProvince="Chiapas" typeStatus="paratype">
<emphasis box="[530,737,1191,1217]" italics="true" pageId="84" pageNumber="85">
<typeStatus box="[530,632,1191,1216]" pageId="84" pageNumber="85">Paratype</typeStatus>
workers:
</emphasis>
<collectingCountry box="[746,861,1191,1217]" name="Mexico" pageId="84" pageNumber="85">MEXICO</collectingCountry>
:
<collectingRegion box="[873,970,1191,1217]" country="Mexico" name="Chiapas" pageId="84" pageNumber="85">Chiapas</collectingRegion>
: same data as holotype,
<specimenCount box="[1257,1362,1191,1217]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(
<specimenCode box="[506,699,1231,1257]" collectionCode="JTLC" pageId="84" pageNumber="85">JTLC000007448</specimenCode>
) [
<collectionCode box="[722,810,1230,1257]" pageId="84" pageNumber="85">MCZC</collectionCode>
]
<specimenCount box="[825,930,1230,1257]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(
<specimenCode box="[946,1140,1231,1257]" collectionCode="JTLC" pageId="84" pageNumber="85">JTLC000007449</specimenCode>
) [
<collectionCode box="[1162,1251,1230,1257]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="84" pageNumber="85" type="Museum">USNM</collectionCode>
]
<specimenCount box="[1267,1372,1230,1257]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(
<specimenCode box="[506,699,1270,1297]" collectionCode="JTLC" pageId="84" pageNumber="85">JTLC000007450</specimenCode>
) [
<collectionCode box="[719,833,1270,1297]" pageId="84" pageNumber="85">ECOSCE</collectionCode>
]
<specimenCount box="[846,960,1270,1296]" pageId="84" pageNumber="85" type="worker">2 workers</specimenCount>
(CASENT0869149, CASENT0869156) [
<collectionCode box="[1427,1515,1270,1297]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="84" pageNumber="85" type="Museum">LACM</collectionCode>
]
<specimenCount pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0869150) [
<collectionCode box="[832,928,1310,1337]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="84" pageNumber="85">UNAM</collectionCode>
]
<specimenCount box="[942,1047,1310,1336]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0869151) [
<collectionCode box="[1298,1386,1310,1337]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="84" pageNumber="85" type="Museum">UCDC</collectionCode>
]
<specimenCount box="[1401,1506,1310,1336]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0869152) [
<collectionCode box="[740,815,1349,1376]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="84" pageNumber="85" type="Museum">FSCA</collectionCode>
]
<specimenCount box="[831,936,1350,1376]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0869153) [
<collectionCode box="[1187,1275,1349,1376]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="84" pageNumber="85">UVGC</collectionCode>
]
<specimenCount box="[1290,1395,1350,1376]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0869154) [
<collectionCode box="[741,837,1389,1416]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="84" pageNumber="85">AMNH</collectionCode>
]
<specimenCount box="[853,957,1390,1416]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0869155) [
<collectionCode box="[1208,1289,1389,1416]" pageId="84" pageNumber="85">CASC</collectionCode>
];
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3321861642" collectingDate="2008-05-19" collectionCode="MCZC, USNM, ECOSCE, LACM, UNAM, UCDC, FSCA, UVGC, AMNH, CASC" collectorName="J. Longino" country="Mexico" elevation="0" latitude="15.72196" location="Custepec" longLatPrecision="1" longitude="-92.95037" pageId="84" pageNumber="85" specimenCode="JTLC000007448, JTLC000007449, JTLC000007450" specimenCount="11" specimenCount-worker="11" stateProvince="Chiapas" typeStatus="paratype">
same data as
<typeStatus box="[498,602,1430,1456]" pageId="84" pageNumber="85">holotype</typeStatus>
, except:
<collectorName box="[704,826,1430,1456]" pageId="84" pageNumber="85">J. Longino</collectorName>
#6286, mesophyll forest, under epiphytes in treefall (CASENT0869119) [
<collectionCode box="[741,839,1469,1496]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="84" pageNumber="85">UNAM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="84.[498,1542,1067,1935]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[530,888,1509,1536]" pageId="84" pageNumber="85">Non-type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861556" collectingDate="2008-05-21" collectionCode="UCDC" collectorName="D. J. Cox" country="Mexico" elevation="0" latitude="15.72065" location="Custepec" longLatPrecision="1" longitude="-92.94008" pageId="84" pageNumber="85" specimenCount="1" specimenCount-worker="1" stateProvince="Chiapas">
<collectingCountry box="[897,1012,1510,1536]" name="Mexico" pageId="84" pageNumber="85">MEXICO</collectingCountry>
:
<collectingRegion box="[1023,1120,1509,1535]" country="Mexico" name="Chiapas" pageId="84" pageNumber="85">Chiapas</collectingRegion>
:
<quantity box="[1132,1193,1509,1536]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="84" pageNumber="85" unit="km" value="3.0">3 km</quantity>
ESE
<location LSID="urn:lsid:plazi:treatment:054FDB70FF93FF82B49C19BD7A6B253C:88393CBDFF93FF8EB1871FFB7C672157" box="[1257,1365,1510,1536]" country="Mexico" latitude="15.72065" longLatPrecision="1" longitude="-92.94008" name="Custepec" pageId="84" pageNumber="85" stateProvince="Chiapas">Custepec</location>
,
<geoCoordinate box="[1378,1516,1505,1536]" degrees="15.72065" direction="north" orientation="latitude" pageId="84" pageNumber="85" precision="1" value="15.72065">
15.72065
<emphasis box="[1482,1492,1505,1523]" italics="true" pageId="84" pageNumber="85">
<superScript attach="right" box="[1482,1492,1505,1523]" fontSize="8" pageId="84" pageNumber="85">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[498,642,1545,1576]" degrees="92.94008" direction="west" orientation="longitude" pageId="84" pageNumber="85" precision="1" value="-92.94008">
92.94008
<emphasis box="[603,613,1545,1563]" italics="true" pageId="84" pageNumber="85">
<superScript attach="right" box="[603,613,1545,1563]" fontSize="8" pageId="84" pageNumber="85">°</superScript>
</emphasis>
W
</geoCoordinate>
±
<elevation box="[677,862,1549,1576]" isEnumeration="true" pageId="84" pageNumber="85">
<quantity box="[677,752,1549,1576]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="84" pageNumber="85" unit="m" value="100.0">100 m</quantity>
,
<quantity box="[766,862,1549,1576]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="84" pageNumber="85" unit="m" value="1800.0">1,800 m</quantity>
</elevation>
,
<date box="[876,1032,1549,1576]" pageId="84" pageNumber="85" value="2008-05-21">
<collectingDate box="[876,1032,1549,1576]" pageId="84" pageNumber="85" value="2008-05-21">21 May 2008</collectingDate>
</date>
,
<collectorName box="[1045,1144,1549,1576]" pageId="84" pageNumber="85">D.J. Cox</collectorName>
#0056, mesophyll forest, canopy baiting,
<specimenCount box="[594,699,1589,1615]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0609722) [
<collectionCode box="[950,1040,1589,1616]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="84" pageNumber="85" type="Museum">UCDC</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph blockId="84.[498,1542,1067,1935]" pageId="84" pageNumber="85">
<materialsCitation ID-GBIF-Occurrence="3321861846" collectingDate="2010-05-17" collectionCode="JTLC" country="Honduras" elevation="0" latitude="14.44412" location="Comayagua" longLatPrecision="1" longitude="-87.55429" pageId="84" pageNumber="85" specimenCount="1" specimenCount-worker="1" stateProvince="Comayagua">
<collectingCountry box="[530,693,1629,1655]" name="Honduras" pageId="84" pageNumber="85">HONDURAS</collectingCountry>
:
<collectingRegion box="[705,846,1629,1655]" country="Honduras" name="Comayagua" pageId="84" pageNumber="85">Comayagua</collectingRegion>
:
<quantity box="[858,919,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="84" pageNumber="85" unit="km" value="9.0">9 km</quantity>
E
<collectingRegion box="[953,1093,1629,1655]" country="Honduras" name="Comayagua" pageId="84" pageNumber="85">Comayagua</collectingRegion>
,
<geoCoordinate box="[1105,1243,1624,1656]" degrees="14.44412" direction="north" orientation="latitude" pageId="84" pageNumber="85" precision="1" value="14.44412">
14.44412
<emphasis box="[1210,1220,1624,1642]" italics="true" pageId="84" pageNumber="85">
<superScript attach="right" box="[1210,1220,1624,1642]" fontSize="8" pageId="84" pageNumber="85">°</superScript>
</emphasis>
N
</geoCoordinate>
<geoCoordinate box="[1252,1395,1624,1656]" degrees="87.55429" direction="west" orientation="longitude" pageId="84" pageNumber="85" precision="1" value="-87.55429">
87.55429
<emphasis box="[1356,1366,1624,1642]" italics="true" pageId="84" pageNumber="85">
<superScript attach="right" box="[1356,1366,1624,1642]" fontSize="8" pageId="84" pageNumber="85">°</superScript>
</emphasis>
W
</geoCoordinate>
±
<elevation isEnumeration="true" pageId="84" pageNumber="85">
<quantity box="[1430,1506,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="84" pageNumber="85" unit="m" value="100.0">100 m</quantity>
,
<quantity box="[498,595,1669,1695]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="84" pageNumber="85" unit="m" value="1720.0">1,720 m</quantity>
</elevation>
,
<date box="[608,764,1669,1695]" pageId="84" pageNumber="85" value="2010-05-17">
<collectingDate box="[608,764,1669,1695]" pageId="84" pageNumber="85" value="2010-05-17">17 May 2010</collectingDate>
</date>
, LLAMA#Go-C-03-2-04, disturbed pine-oak forest, beating vegetation,
<specimenCount box="[633,738,1709,1735]" pageId="84" pageNumber="85" type="worker">1 worker</specimenCount>
(CASENT0615348) [
<collectionCode box="[989,1060,1708,1735]" pageId="84" pageNumber="85">JTLC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="84" pageNumber="85" type="distribution">
<paragraph blockId="84.[498,1542,1067,1935]" pageId="84" pageNumber="85">
<emphasis bold="true" box="[530,754,1748,1775]" pageId="84" pageNumber="85">Geographic range:</emphasis>
Southern Mexico to
<collectingCountry box="[1003,1122,1748,1774]" name="Honduras" pageId="84" pageNumber="85">Honduras</collectingCountry>
, mid elevations of the Sierra Madre de Chiapas and Central American Nucleus (
<figureCitation box="[1022,1122,1788,1815]" captionStart="Figure 93" captionStartId="85.[524,589,964,986]" captionTargetBox="[498,1541,234,951]" captionTargetId="figure-289@85.[498,1534,234,936]" captionTargetPageId="85" captionText="Figure 93 Temnothorax acutispinosus sp. nov. holotype worker (JTLC000007447). (A) Profile view. (B) Full-face view. (C) Dorsal view. (D) Geographical and elevational distribution of specimens examined. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-93" figureDoi="http://doi.org/10.5281/zenodo.5102271" httpUri="https://zenodo.org/record/5102271/files/figure.png" pageId="84" pageNumber="85">Fig. 93D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="diagnosis">
<paragraph blockId="84.[498,1542,1067,1935]" lastBlockId="85.[498,1542,1158,1902]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis bold="true" box="[530,753,1828,1854]" pageId="84" pageNumber="85">Worker diagnosis:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[760,1077,1828,1854]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[760,1077,1828,1854]" italics="true" pageId="84" pageNumber="85">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1084,1174,1829,1855]" pageId="84" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[663,738,1868,1894]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[663,738,1868,1894]" italics="true" pageId="84" pageNumber="85">salvini</emphasis>
</taxonomicName>
clade by the following character combination: dorsum of mesosoma very weakly sinuate in profile view; hind femora weakly incrassate (FI 258319); propodeal spines acute and directed dorsally; petiolar node rounded dorsally; postpetiole moderately broad (PWI 159175); integument dark brown; setae on head, mesosoma, waist segments and gaster erect, short, sparse and blunt (never long and tapering).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102271" ID-Zenodo-Dep="5102271" httpUri="https://zenodo.org/record/5102271/files/figure.png" pageId="85" pageNumber="86" startId="85.[524,589,964,986]" targetBox="[498,1541,234,951]" targetPageId="85">
<paragraph blockId="85.[524,1516,964,1103]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[524,1349,964,986]" pageId="85" pageNumber="86">
Figure 93
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[631,902,964,986]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis bold="true" box="[631,902,964,986]" italics="true" pageId="85" pageNumber="86">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[910,987,964,985]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
holotype worker (JTLC000007447).
</emphasis>
(A) Profile view. (B) Full-face view. (C) Dorsal view. (D) Geographical and elevational distribution of specimens examined. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-93
</paragraph>
</caption>
<paragraph blockId="85.[498,1542,1158,1902]" pageId="85" pageNumber="86">
<emphasis bold="true" box="[530,717,1277,1303]" pageId="85" pageNumber="86">Similar species:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[723,943,1278,1304]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="achii" status="sp. nov.">
<emphasis box="[723,943,1278,1304]" italics="true" pageId="85" pageNumber="86">Temnothorax achii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[950,1041,1278,1304]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1052,1190,1278,1304]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
<emphasis box="[1052,1190,1278,1304]" italics="true" pageId="85" pageNumber="86">T. altinodus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1197,1288,1278,1304]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis italics="true" pageId="85" pageNumber="86">
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1299,1376,1278,1304]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="ixili">T. ixili</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1388,1538,1278,1304]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,640,1317,1343]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">T. tuxtlanus</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[649,742,1318,1344]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[755,858,1318,1343]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="xincai" status="sp. nov.">
<emphasis box="[755,858,1318,1343]" italics="true" pageId="85" pageNumber="86">T. xincai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[867,960,1318,1344]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
, members of the
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[1167,1251,1318,1343]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="augusti">
<emphasis box="[1167,1251,1318,1343]" italics="true" pageId="85" pageNumber="86">augusti</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1266,1353,1318,1343]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="goniops">
<emphasis box="[1266,1353,1318,1343]" italics="true" pageId="85" pageNumber="86">goniops</emphasis>
</taxonomicName>
, and
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1420,1533,1317,1343]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[1420,1533,1317,1343]" italics="true" pageId="85" pageNumber="86">pulchellus</emphasis>
</taxonomicName>
groups, and species of the
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[812,869,1357,1383]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="sallei">
<emphasis box="[812,869,1357,1383]" italics="true" pageId="85" pageNumber="86">sallei</emphasis>
</taxonomicName>
clade.
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[953,1272,1357,1383]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[953,1272,1357,1383]" italics="true" pageId="85" pageNumber="86">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1281,1372,1358,1384]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from similar appearing members of the
<taxonomicName baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1852" box="[1136,1193,1397,1423]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="sallei">
<emphasis box="[1136,1193,1397,1423]" italics="true" pageId="85" pageNumber="86">sallei</emphasis>
</taxonomicName>
clade (e.g.,
<emphasis italics="true" pageId="85" pageNumber="86">
<taxonomicName box="[1334,1444,1398,1423]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="manni">T. manni</taxonomicName>
,
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[498,655,1438,1463]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="mexicanus">T. mexicanus</taxonomicName>
,
<taxonomicName authorityName="" baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[669,850,1437,1463]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="punctithorax">T. punctithorax</taxonomicName>
</emphasis>
) by the petiole, which lacks a transverse carina anterodorsally. In contrast to all of the above species (except
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1221,1361,1477,1503]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
<emphasis box="[1221,1361,1477,1503]" italics="true" pageId="85" pageNumber="86">T. altinodus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1370,1464,1477,1503]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
),
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,684,1517,1542]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[498,684,1517,1542]" italics="true" pageId="85" pageNumber="86">T. acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[693,785,1517,1543]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
has acute, dorsally directed propodeal spines.
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis italics="true" pageId="85" pageNumber="86">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[660,752,1557,1583]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1076,1215,1556,1582]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
<emphasis box="[1076,1215,1556,1582]" italics="true" pageId="85" pageNumber="86">T. altinodus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1224,1316,1557,1583]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
by the relatively compact mesosoma (WLI 131136 vs.
<quantity box="[953,1025,1596,1623]" metricMagnitude="0" metricUnit="m" metricValue="3.7592" pageId="85" pageNumber="86" unit="in" value="148.0">148 in</quantity>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1034,1173,1596,1622]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
<emphasis box="[1034,1173,1596,1622]" italics="true" pageId="85" pageNumber="86">T. altinodus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1180,1272,1597,1623]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
) and weakly incrassate hind femora, which are enlarged in
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[923,1063,1636,1662]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="altinodus" status="sp. nov.">
<emphasis box="[923,1063,1636,1662]" italics="true" pageId="85" pageNumber="86">T. altinodus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1071,1163,1637,1663]" pageId="85" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="88" lastPageNumber="89" pageId="85" pageNumber="86" type="description">
<paragraph blockId="85.[498,1542,1158,1902]" lastBlockId="86.[498,1542,234,1934]" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">
<emphasis bold="true" box="[530,1018,1676,1702]" pageId="85" pageNumber="86">
Worker measurements &amp; indices (
<emphasis bold="true" box="[941,957,1676,1702]" italics="true" pageId="85" pageNumber="86">n</emphasis>
= 3):
</emphasis>
SL =
<geoCoordinate box="[1084,1146,1676,1703]" degrees="0.533" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.533">0.533</geoCoordinate>
<geoCoordinate box="[1146,1223,1676,1702]" degrees="0.608" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.608">0.608</geoCoordinate>
(0.567); FRS =
<geoCoordinate box="[1402,1464,1676,1702]" degrees="0.201" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.201">0.201</geoCoordinate>
<geoCoordinate box="[1464,1541,1676,1702]" degrees="0.232" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.232">0.232</geoCoordinate>
(0.217); CW =
<geoCoordinate box="[678,740,1716,1742]" degrees="0.689" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.689">0.689</geoCoordinate>
<geoCoordinate box="[740,817,1716,1742]" degrees="0.770" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.77">0.770</geoCoordinate>
(0.725); CWb =
<geoCoordinate box="[1020,1082,1716,1742]" degrees="0.636" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.636">0.636</geoCoordinate>
<geoCoordinate box="[1082,1159,1716,1743]" degrees="0.725" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.725">0.725</geoCoordinate>
(0.675); PoOC =
<geoCoordinate box="[1371,1433,1716,1742]" degrees="0.287" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.287">0.287</geoCoordinate>
<geoCoordinate box="[1434,1511,1716,1742]" degrees="0.324" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.324">0.324</geoCoordinate>
(0.302); CL =
<geoCoordinate box="[665,727,1756,1782]" degrees="0.721" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.721">0.721</geoCoordinate>
<geoCoordinate box="[727,804,1756,1782]" degrees="0.816" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.816">0.816</geoCoordinate>
(0.764); EL =
<geoCoordinate box="[977,1039,1756,1783]" degrees="0.165" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.165">0.165</geoCoordinate>
<geoCoordinate box="[1039,1116,1756,1783]" degrees="0.195" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.195">0.195</geoCoordinate>
(0.179); EW =
<geoCoordinate box="[1302,1364,1756,1782]" degrees="0.122" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.122">0.122</geoCoordinate>
<geoCoordinate box="[1364,1441,1756,1782]" degrees="0.124" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.124">0.124</geoCoordinate>
(0.123); MD =
<geoCoordinate box="[580,642,1796,1822]" degrees="0.180" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.18">0.180</geoCoordinate>
<geoCoordinate box="[642,719,1796,1822]" degrees="0.210" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.21">0.210</geoCoordinate>
(0.198); WL =
<geoCoordinate box="[904,966,1796,1822]" degrees="0.867" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.867">0.867</geoCoordinate>
<geoCoordinate box="[966,1043,1796,1822]" degrees="0.964" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.964">0.964</geoCoordinate>
(0.899); SPST =
<geoCoordinate box="[1245,1307,1796,1822]" degrees="0.188" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.188">0.188</geoCoordinate>
<geoCoordinate box="[1308,1385,1796,1822]" degrees="0.233" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.233">0.233</geoCoordinate>
(0.211); MPST =
<geoCoordinate box="[607,669,1836,1862]" degrees="0.272" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.272">0.272</geoCoordinate>
<geoCoordinate box="[669,746,1835,1862]" degrees="0.338" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.338">0.338</geoCoordinate>
(0.298); PEL =
<geoCoordinate box="[935,997,1835,1862]" degrees="0.325" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.325">0.325</geoCoordinate>
<geoCoordinate box="[997,1074,1835,1862]" degrees="0.337" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.337">0.337</geoCoordinate>
(0.331); NOL =
<geoCoordinate box="[1274,1336,1836,1862]" degrees="0.183" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.183">0.183</geoCoordinate>
<geoCoordinate box="[1336,1413,1835,1862]" degrees="0.204" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.204">0.204</geoCoordinate>
(0.195); NOH =
<geoCoordinate box="[598,660,1876,1902]" degrees="0.137" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.137">0.137</geoCoordinate>
<geoCoordinate box="[660,737,1875,1902]" degrees="0.139" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.139">0.139</geoCoordinate>
(0.138); PEH =
<geoCoordinate box="[933,995,1876,1902]" degrees="0.238" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.238">0.238</geoCoordinate>
<geoCoordinate box="[995,1072,1875,1901]" degrees="0.263" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.263">0.263</geoCoordinate>
(0.25); PPL =
<geoCoordinate box="[1248,1310,1876,1902]" degrees="0.200" direction="north" orientation="latitude" pageId="85" pageNumber="86" precision="55" value="0.2">0.200</geoCoordinate>
<geoCoordinate box="[1310,1387,1875,1902]" degrees="0.231" direction="west" orientation="longitude" pageId="85" pageNumber="86" precision="55" value="-0.231">0.231</geoCoordinate>
(0.213); PW =
<geoCoordinate box="[577,639,234,260]" degrees="0.402" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.402">0.402</geoCoordinate>
<geoCoordinate box="[640,716,234,260]" degrees="0.464" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.464">0.464</geoCoordinate>
(0.431); SBPA =
<geoCoordinate box="[925,987,234,261]" degrees="0.157" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.157">0.157</geoCoordinate>
<geoCoordinate box="[987,1064,234,260]" degrees="0.180" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.18">0.180</geoCoordinate>
(0.166); SPTI =
<geoCoordinate box="[1263,1325,234,260]" degrees="0.208" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.208">0.208</geoCoordinate>
<geoCoordinate box="[1325,1402,234,260]" degrees="0.210" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.21">0.210</geoCoordinate>
(0.209); PEW =
<geoCoordinate box="[594,656,274,300]" degrees="0.148" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.148">0.148</geoCoordinate>
<geoCoordinate box="[656,733,274,300]" degrees="0.170" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.17">0.170</geoCoordinate>
(0.158); PNW =
<geoCoordinate box="[940,1002,274,301]" degrees="0.155" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.155">0.155</geoCoordinate>
<geoCoordinate box="[1002,1079,274,300]" degrees="0.163" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.163">0.163</geoCoordinate>
(0.158); PPW =
<geoCoordinate box="[1281,1343,274,301]" degrees="0.258" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.258">0.258</geoCoordinate>
<geoCoordinate box="[1344,1421,274,300]" degrees="0.271" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.271">0.271</geoCoordinate>
(0.263); HFL =
<geoCoordinate box="[586,648,314,341]" degrees="0.564" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.564">0.564</geoCoordinate>
<geoCoordinate box="[648,725,314,341]" degrees="0.652" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.652">0.652</geoCoordinate>
(0.599); HFWmax =
<geoCoordinate box="[981,1043,314,340]" degrees="0.144" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.144">0.144</geoCoordinate>
<geoCoordinate box="[1044,1121,314,340]" degrees="0.169" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.169">0.169</geoCoordinate>
(0.158); HFWmin =
<geoCoordinate box="[1374,1436,314,341]" degrees="0.050" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.05">0.050</geoCoordinate>
<geoCoordinate box="[1437,1514,314,340]" degrees="0.062" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.062">0.062</geoCoordinate>
(0.055); CS =
<geoCoordinate box="[659,721,354,380]" degrees="0.997" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.997">0.997</geoCoordinate>
<geoCoordinate box="[722,799,354,380]" degrees="1.133" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-1.133">1.133</geoCoordinate>
(1.057); ES =
<geoCoordinate box="[965,1027,354,380]" degrees="0.227" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="0.227">0.227</geoCoordinate>
<geoCoordinate box="[1027,1104,354,381]" degrees="0.257" direction="west" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="-0.257">0.257</geoCoordinate>
(0.240); SI = 84; OI = 23; CI = 8889 (88); WLI = 131136 (133); SBI = 2425 (25); PSI = 2227 (24); PWI = 159175 (166); PLI = 146163 (156); NI = 134149 (141); PNWI = 96105 (100); NLI = 5661 (59); FI = 258319 (288).
</paragraph>
<paragraph blockId="86.[498,1542,234,1934]" pageId="86" pageNumber="87">
<emphasis bold="true" box="[530,775,513,539]" pageId="86" pageNumber="87">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 8889). Mandibles densely, finely striate but shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin emarginated medially. Antennal scapes short: when fully retracted, failing to reach the posterior margin of the head capsule by about the maximum width of the scape (SI 84). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment slightly longer than the preceding two in combination. Compound eyes moderately protruding past the lateral margins of the head capsule. Frontal carinae short, extending past the antennal toruli by about one and a half times the maximum width of the antennal scape. Lateral margin of head weakly convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin flat but rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="86.[498,1542,234,1934]" lastBlockId="87.[498,1542,230,1934]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
In profile view, compound eyes ovular and moderately large (OI 23), with 11 ommatidia in longest row. Pronotal declivity distinct: dorsal margin of anterior face of pronotum marked by a weak carina; neck and anterior face of pronotum forming a ~120
<emphasis box="[1423,1433,1106,1124]" italics="true" pageId="86" pageNumber="87">
<superScript attach="left" box="[1423,1433,1106,1124]" fontSize="8" pageId="86" pageNumber="87">°</superScript>
</emphasis>
angle. Mesosoma very weakly sinuate: dorsum of promesonotum forming an even convexity between the pronotal declivity and the metanotum, metanotum and propodeum slightly depressed below the level of the promesonotum; metanotum slightly convex in some specimens; propodeal dorsum flat. Promesonotal suture extending from the posterior margin of the procoxal insertion to the mesothoracic spiracle, which is moderately well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae, continuing dorsally as a weak impression between the posterior margin of the mesonotum and the anterior margin of the metanotum. Propodeal spiracle weakly developed, directed posterolaterally, and separated from the propodeal declivity by about five spiracle diameters. Propodeal spines moderately long, about as long as the propodeal declivity (PSI 2227), tapering evenly from the base, directed dorsally, and acute. Propodeal declivity with a fine carina joining the base of the propodeal spines and the propodeal lobes; weakly concave, forming a ~120
<emphasis box="[1384,1394,1624,1642]" italics="true" pageId="86" pageNumber="87">
<superScript attach="left" box="[1384,1394,1624,1642]" fontSize="8" pageId="86" pageNumber="87">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole short (PLI 146163), with weakly developed tubercles anterodorsally. Subpetiolar process in the form of a moderately large, blunt, triangular tooth; ventral margin of petiole weakly bulging posterior to it. Petiolar peduncle short: comprising about a quarter of the total petiolar length. Petiolar node robust, erect, and rounded: transition between peduncle and node marked by a rounded angle of ~120
<emphasis box="[659,669,230,248]" italics="true" pageId="87" pageNumber="88">
<superScript attach="none" box="[659,669,230,248]" fontSize="8" pageId="87" pageNumber="88">°</superScript>
</emphasis>
, resulting in a weakly concave anterior node face; anterior face rounding evenly into the convex dorsal face; dorsal face rounding evenly into the posterior face, which forms a ~120
<emphasis box="[727,737,309,327]" italics="true" pageId="87" pageNumber="88">
<superScript attach="left" box="[727,737,309,327]" fontSize="8" pageId="87" pageNumber="88">°</superScript>
</emphasis>
angle with the caudal cylinder. Postpetiole evenly rounded anteriorly, strongly bulging anterodorsally, flattened posterodorsally; concave ventrally, with an acute, anteriorly directed subpostpetiolar process.
</paragraph>
<paragraph blockId="87.[498,1542,230,1934]" pageId="87" pageNumber="88">
In dorsal view, humeri moderately well developed and distinct: evenly rounded and wider than the rest of the mesosoma; mesothoracic spiracles very weakly protruding past the lateral margins of the mesosoma, visible as slight angles where the pronotum meets the mesonotum. Promesonotal groove represented by a weak sulcus. Metanotum delineated anteriorly and posteriorly by very faint impressions. Propodeal spines narrowly approximated basally and diverging apically, their apices separated from each other by about their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles weakly protruding past the lateral margins, but not noticeably constricted anterior to them. Petiolar node evenly ovular; node slightly wider than the peduncle, and evenly grading into the caudal cylinder, which the same width as the node. Postpetiole narrow (PWI 159175) and subquadrate. Anterior margin of the postpetiole weakly convex and meets the lateral margins at a rounded ~90
<emphasis box="[1151,1161,867,885]" italics="true" pageId="87" pageNumber="88">
<superScript attach="left" box="[1151,1161,867,885]" fontSize="8" pageId="87" pageNumber="88">°</superScript>
</emphasis>
angle; lateral margins parallel to each other; posterior margin flat. Metafemur moderately to strongly incrassate (FI 258319).
</paragraph>
<paragraph blockId="87.[498,1542,230,1934]" pageId="87" pageNumber="88">Sculpture: median clypeal carina present, extending posteriorly nearly to the frontal triangle, and flanked on either side by two equally strong carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture areolate. Antennal scapes shining through weak areolate ground sculpture. Cephalic dorsum areolate, with the areolae arranged into longitudinal rows by fine costulae. Lateral surfaces of head areolate, with sculpture similar to the dorsum of the head between the compound eye and the posterior margin of the head; sculpture between the compound eye and the mandibular insertion areolate-rugose. Ventral surface of head smooth and shining, with weak costulae. Pronotal neck areolate. Anterior face of the pronotum areolate. Lateral surface of the mesosoma areolate, with weak costulae overlying the ground sculpture on all surfaces except between the propodeal spiracle and the base of the propodeal spines. Dorsal surface of mesosoma areolate, with overlying rugose sculpture on the pronotum. Femora shining, with traces of weak areolate sculpture. Petiole and postpetiole predominantly areolate, but the anterior face of the postpetiole smooth and shining. First gastral tergite and sternite smooth and shining, with weak spectral iridescence.</paragraph>
<paragraph blockId="87.[498,1542,230,1934]" pageId="87" pageNumber="88">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about the width of the compound eye. The head bears ~28, mesosoma ~16, petiole 6, postpetiole ~14, and first gastral tergite ~44 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the densely sculptured integument.</paragraph>
<paragraph blockId="87.[498,1542,230,1934]" pageId="87" pageNumber="88">Color: predominantly dark brown, with antennae, apices of mandibles, and legs testaceous. Sting testaceous yellow.</paragraph>
<paragraph blockId="87.[498,1542,230,1934]" box="[530,734,1908,1934]" pageId="87" pageNumber="88">
<emphasis bold="true" box="[530,601,1908,1934]" pageId="87" pageNumber="88">Gyne:</emphasis>
Unknown.
</paragraph>
<paragraph blockId="88.[498,1542,234,619]" box="[530,730,234,260]" pageId="88" pageNumber="89">
<emphasis bold="true" box="[530,597,234,260]" pageId="88" pageNumber="89">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="88" pageNumber="89" type="etymology">
<paragraph blockId="88.[498,1542,234,619]" pageId="88" pageNumber="89">
<emphasis bold="true" box="[530,668,274,300]" pageId="88" pageNumber="89">Etymology:</emphasis>
Morphological, from the Latin acutus (= sharp) + spinosus (= thorny), a reference to the sharp, dorsally directed propodeal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="88" pageNumber="89" type="discussion">
<paragraph blockId="88.[498,1542,234,619]" pageId="88" pageNumber="89">
<emphasis bold="true" box="[530,670,353,379]" pageId="88" pageNumber="89">Comments:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[677,993,354,380]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis box="[677,993,354,380]" italics="true" pageId="88" pageNumber="89">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1000,1089,354,380]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
is known from a few collections in mid elevation mesic forest in
<collectingRegion box="[793,948,394,420]" country="Mexico" name="Chiapas" pageId="88" pageNumber="89">Chiapas state</collectingRegion>
in
<collectingCountry box="[990,1076,394,420]" name="Mexico" pageId="88" pageNumber="89">Mexico</collectingCountry>
and
<collectingRegion box="[1137,1275,394,420]" country="Honduras" name="Comayagua" pageId="88" pageNumber="89">Comayagua</collectingRegion>
in
<collectingCountry box="[1316,1436,394,420]" name="Honduras" pageId="88" pageNumber="89">Honduras</collectingCountry>
. This species is apparently a canopy dweller, and probably nests arboreally: the
<typeStatus box="[1369,1429,434,460]" pageId="88" pageNumber="89">types</typeStatus>
were collected from under epiphytes on a treefall or via canopy baiting.
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
<emphasis italics="true" pageId="88" pageNumber="89">Temnothorax acutispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[660,752,514,540]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
is closely related to the
<taxonomicName baseAuthorityName="Mackay" baseAuthorityYear="2000" box="[1039,1126,514,539]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis box="[1039,1126,514,539]" italics="true" pageId="88" pageNumber="89">rugosus</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1187,1283,514,539]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="annexus">
<emphasis box="[1187,1283,514,539]" italics="true" pageId="88" pageNumber="89">annexus</emphasis>
</taxonomicName>
species groups from
<collectingCountry box="[498,584,553,579]" name="Mexico" pageId="88" pageNumber="89">Mexico</collectingCountry>
north of the Isthmus of Tehuantepec and the Southwestern
<collectingCountry box="[1302,1461,553,579]" name="United States of America" pageId="88" pageNumber="89">United States</collectingCountry>
, all of which are apparently arboreal.
</paragraph>
</subSubSection>
</treatment>
</document>