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<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FF16FF02B49C1C3E7C582048" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax nigricans" docType="treatment" docVersion="4" lastPageNumber="216" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="210" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prebus, Matthew M.</mods:namePart>
<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States &amp; Department of Entomology &amp; Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title>PeerJ</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="series">
<mods:title>e 11514</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2021-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>9</mods:number>
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<mods:start>1</mods:start>
<mods:end>462</mods:end>
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<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
<mods:identifier type="ISSN">2167-8359</mods:identifier>
<mods:identifier type="PMC">PMC8254503</mods:identifier>
<mods:identifier type="PubMed">34249486</mods:identifier>
<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102545" ID-GBIF-Taxon="183542667" ID-Zenodo-Dep="5102545" LSID="urn:lsid:plazi:treatment:054FDB70FF16FF02B49C1C3E7C582048" httpUri="http://treatment.plazi.org/id/054FDB70FF16FF02B49C1C3E7C582048" lastPageId="216" lastPageNumber="216" pageId="209" pageNumber="210">
<subSubSection box="[498,1058,1571,1598]" pageId="209" pageNumber="210" type="nomenclature">
<paragraph blockId="209.[498,1072,1571,1638]" box="[498,1058,1571,1598]" pageId="209" pageNumber="210">
<heading box="[498,1058,1571,1598]" level="2" pageId="209" pageNumber="210">
<taxonomicName authority="(Baroni Urbani, 1978)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,1058,1571,1598]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis bold="true" box="[498,1058,1571,1598]" pageId="209" pageNumber="210">
<emphasis bold="true" box="[498,778,1571,1598]" italics="true" pageId="209" pageNumber="210">Temnothorax nigricans</emphasis>
(
<bibRefCitation author="Baroni Urbani C." box="[798,1048,1571,1598]" journalOrPublisher="Entomologica Basiliensia" pageId="209" pageNumber="210" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis bold="true" box="[798,1048,1571,1598]" italics="true" pageId="209" pageNumber="210">Baroni Urbani, 1978</emphasis>
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[498,1072,1612,1638]" pageId="209" pageNumber="210" type="distribution">
<paragraph blockId="209.[498,1072,1571,1638]" box="[498,1072,1612,1638]" pageId="209" pageNumber="210">
Distribution:
<figureCitation box="[657,767,1612,1638]" captionStart="Figure 121" captionStartId="205.[116,181,1124,1146]" captionTargetBox="[90,1541,234,1111]" captionTargetId="figure-151@205.[90,1524,234,1096]" captionTargetPageId="205" captionText="Figure 121 Geographical and elevational distribution of the pastinifer group. (A) Temnothorax androsanus (B) T. nigricans (C) T. pastinifer (D) T. rutabulafer sp. nov. (E) T. schwarzi. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-121" figureDoi="http://doi.org/10.5281/zenodo.5102351" httpUri="https://zenodo.org/record/5102351/files/figure.png" pageId="209" pageNumber="210">Fig. 121B</figureCitation>
; worker &amp; gyne:
<figureCitation box="[972,1066,1612,1638]" captionStart="Figure 123" captionStartId="210.[524,589,1343,1365]" captionTargetBox="[498,1541,234,1330]" captionTargetId="figure-109@210.[498,1534,234,1315]" captionTargetPageId="210" captionText="Figure 123 Temnothorax nigricans. (AC) Lectotype worker (MCZENT00577111). (A) Full-face. (B) Profile view. (C) Dorsal view. (DF) Paralectotype gyne (MCZENT00577113). (D) Full-face. (E) Profile view. (F) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-123" figureDoi="http://doi.org/10.5281/zenodo.5102357" httpUri="https://zenodo.org/record/5102357/files/figure.png" pageId="209" pageNumber="210">Fig. 123</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="210" lastPageNumber="211" pageId="209" pageNumber="210" type="reference_group">
<paragraph blockId="209.[498,1542,1674,1860]" pageId="209" pageNumber="210">
<treatmentCitationGroup pageId="209" pageNumber="210">
<treatmentCitation author="Wheeler WM" box="[498,1050,1674,1701]" page="15" pageId="209" pageNumber="210" year="1931">
<taxonomicName authority="Wheeler, 1931: 15" authorityName="Wheeler" authorityPageNumber="15" authorityYear="1931" box="[498,1006,1674,1700]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="melanocephala">
<emphasis box="[498,1006,1674,1700]" italics="true" pageId="209" pageNumber="210">
Macromischa melanocephala
<bibRefCitation author="Wheeler WM" box="[842,1050,1674,1701]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="209" pageNumber="210" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">Wheeler, 1931: 15</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[1062,1154,1674,1700]" pageId="209" pageNumber="210">Syntype</typeStatus>
workers and gynes.
<location LSID="urn:lsid:plazi:treatment:054FDB70FF16FF02B49C1C3E7C582048:88393CBDFF16FF0BB01D1C977B50219B" country="Cuba" name="Ensenada de Cochinos" pageId="209" pageNumber="210">
Ensenada
<collectorName pageId="209" pageNumber="210">de Cochinos</collectorName>
</location>
,
<collectingCountry box="[624,689,1714,1740]" name="Cuba" pageId="209" pageNumber="210">Cuba</collectingCountry>
. One
<typeStatus box="[759,848,1715,1741]" pageId="209" pageNumber="210">syntype</typeStatus>
worker here designated
<emphasis bold="true" box="[1139,1248,1714,1740]" pageId="209" pageNumber="210">
<typeStatus box="[1139,1248,1714,1740]" pageId="209" pageNumber="210">lectotype</typeStatus>
</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="209.[498,1542,1674,1860]" pageId="209" pageNumber="210">
<treatmentCitationGroup pageId="209" pageNumber="210">
<treatmentCitation author="Baroni Urbani C." box="[498,1061,1754,1781]" page="467" pageId="209" pageNumber="210" year="1978">
<taxonomicName authority=": Baroni Urbani, 1978: 467" authorityName="Baroni Urbani" authorityPageNumber="467" authorityYear="1978" box="[498,1002,1754,1781]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[498,751,1754,1780]" italics="true" pageId="209" pageNumber="210">Leptothorax nigricans</emphasis>
:
<bibRefCitation author="Baroni Urbani C." box="[764,1061,1754,1781]" journalOrPublisher="Entomologica Basiliensia" pageId="209" pageNumber="210" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
<emphasis box="[764,1002,1754,1780]" italics="true" pageId="209" pageNumber="210">Baroni Urbani, 1978</emphasis>
: 467
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Transfer to
<taxonomicName authority="Mayr" box="[1210,1350,1754,1780]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis box="[1210,1350,1754,1780]" italics="true" pageId="209" pageNumber="210">Leptothorax</emphasis>
</taxonomicName>
. Nomen novum for
<taxonomicName authority="(Wheeler)" baseAuthorityName="Wheeler" box="[541,1012,1794,1821]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="melanocephalum">
<emphasis box="[541,885,1794,1820]" italics="true" pageId="209" pageNumber="210">Leptothorax melanocephalum</emphasis>
(Wheeler)
</taxonomicName>
(nec
<treatmentCitation author="Emery C." page="197" pageId="209" pageNumber="210" year="1870">
<taxonomicName authority="Emery, 1870: 197" authorityName="Emery" authorityPageNumber="197" authorityYear="1870" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="melanocephalum">
<emphasis italics="true" pageId="209" pageNumber="210">
Leptothorax melanocephalum
<bibRefCitation author="Emery C." journalOrPublisher="Bullettino della Societa Entomologica Italiana" pageId="209" pageNumber="210" pagination="193 - 201" part="2" refId="ref270637" refString="Emery C. 1870. Studi mirmecologici. Bullettino della Societa Entomologica Italiana 2: 193 - 201." title="Studi mirmecologici" type="journal article" year="1870">Emery, 1870: 197</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5102357" ID-Zenodo-Dep="5102357" httpUri="https://zenodo.org/record/5102357/files/figure.png" pageId="210" pageNumber="211" startId="210.[524,589,1343,1365]" targetBox="[498,1541,234,1330]" targetPageId="210">
<paragraph blockId="210.[524,1516,1343,1424]" pageId="210" pageNumber="211">
<emphasis bold="true" box="[524,890,1343,1365]" pageId="210" pageNumber="211">
Figure 123
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[649,884,1343,1365]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="210" pageNumber="211" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis bold="true" box="[649,884,1343,1365]" italics="true" pageId="210" pageNumber="211">Temnothorax nigricans</emphasis>
</taxonomicName>
.
</emphasis>
(AC) Lectotype worker (MCZENT00577111). (A) Full-face. (B) Profile view. (C) Dorsal view. (DF) Paralectotype gyne (MCZENT00577113). (D) Full-face. (E) Profile view. (F) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-123
</paragraph>
</caption>
<paragraph blockId="210.[498,1542,1479,1904]" pageId="210" pageNumber="211">
<treatmentCitationGroup pageId="210" pageNumber="211">
<taxonomicName authority="(Baroni Urbani)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,970,1479,1506]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="210" pageNumber="211" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[498,769,1479,1505]" italics="true" pageId="210" pageNumber="211">Temnothorax nigricans</emphasis>
(Baroni Urbani)
</taxonomicName>
:
<treatmentCitation author="Bolton B." box="[983,1188,1479,1505]" page="271" pageId="210" pageNumber="211" year="2003">
<bibRefCitation author="Bolton B." box="[983,1188,1479,1505]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="210" pageNumber="211" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
<emphasis box="[983,1128,1479,1505]" italics="true" pageId="210" pageNumber="211">
<collectingRegion box="[983,1061,1479,1505]" country="United Kingdom" name="Bolton" pageId="210" pageNumber="211">
<author box="[983,1061,1479,1505]" fns="*" inm="true" npo="*" pageId="210" pageNumber="211">Bolton</author>
</collectingRegion>
,
<year box="[1073,1128,1479,1505]" pageId="210" pageNumber="211">2003</year>
</emphasis>
: 271
</bibRefCitation>
</treatmentCitation>
. First combination in
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1519,1545]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="210" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis box="[498,655,1519,1545]" italics="true" pageId="210" pageNumber="211">Temnothorax</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="210" pageNumber="211" type="materials_examined">
<paragraph blockId="210.[498,1542,1479,1904]" pageId="210" pageNumber="211">
<emphasis bold="true" box="[530,834,1558,1585]" pageId="210" pageNumber="211">Type material examined:</emphasis>
<materialsCitation ID-GBIF-Occurrence="3321861810" collectingDate="1929-10-05" collectionCode="MCZC" collectorName="J. G. Myers &amp; M. C. Z. Cotype" country="Cuba" location="CUBA" pageId="210" pageNumber="211" specimenCount="1" typeStatus="lectotype">
<emphasis box="[843,1045,1558,1584]" italics="true" pageId="210" pageNumber="211">
<typeStatus box="[843,951,1559,1584]" pageId="210" pageNumber="211">Lectotype</typeStatus>
worker:
</emphasis>
<collectingCountry box="[1054,1135,1559,1585]" name="Cuba" pageId="210" pageNumber="211">CUBA</collectingCountry>
: Artemisia: Hacienda Jiqui, Ensenada de Cochinos,
<date box="[780,964,1598,1625]" pageId="210" pageNumber="211" value="1929-10-05">
<collectingDate box="[780,964,1598,1625]" pageId="210" pageNumber="211" value="1929-10-05">5 October 1929</collectingDate>
</date>
,
<collectorName box="[977,1103,1598,1625]" pageId="210" pageNumber="211">J.G. Myers</collectorName>
#C250, nest in fallen mahogany fruit,
<collectorName box="[498,672,1638,1664]" pageId="210" pageNumber="211">M.C.Z. Cotype</collectorName>
16365 (MCZENT00577110, middle specimen on pin) [
<collectionCode box="[1326,1417,1638,1665]" pageId="210" pageNumber="211">MCZC</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph blockId="210.[498,1542,1479,1904]" pageId="210" pageNumber="211">
<materialsCitation ID-GBIF-Occurrence="3321861709" collectingDate="1929-10-05" collectionCode="MCZC, USNM, MNHN" collectorName="J. G. Myers &amp; M. C. Z. Cotype" country="Cuba" location="CUBA" pageId="210" pageNumber="211" specimenCount="20" specimenCount-worker="20" typeStatus="paralectotype">
<emphasis box="[530,913,1678,1704]" italics="true" pageId="210" pageNumber="211">
<typeStatus box="[530,684,1678,1704]" pageId="210" pageNumber="211">Paralectotype</typeStatus>
workers and gynes:
</emphasis>
same pin as
<typeStatus box="[1069,1176,1678,1704]" pageId="210" pageNumber="211">lectotype</typeStatus>
,
<specimenCount box="[1189,1305,1678,1704]" pageId="210" pageNumber="211" type="worker">2 workers</specimenCount>
(top and bottom specimens on pin) [
<collectionCode box="[731,821,1717,1744]" pageId="210" pageNumber="211">MCZC</collectionCode>
]; same data as lectotype,
<specimenCount box="[1119,1235,1718,1744]" pageId="210" pageNumber="211" type="worker">3 workers</specimenCount>
(MCZENT00577111) [
<collectionCode box="[505,593,1757,1784]" pageId="210" pageNumber="211">MCZC</collectionCode>
]
<specimenCount box="[609,724,1758,1784]" pageId="210" pageNumber="211" type="worker">2 workers</specimenCount>
(MCZENT00577112) [
<collectionCode box="[999,1087,1757,1784]" pageId="210" pageNumber="211">MCZC</collectionCode>
] 2 dealate gynes (MCZENT00577113) [
<collectionCode box="[505,593,1797,1824]" pageId="210" pageNumber="211">MCZC</collectionCode>
]
<specimenCount box="[609,724,1797,1824]" pageId="210" pageNumber="211" type="worker">3 workers</specimenCount>
(MCZENT00577114) [
<collectionCode box="[999,1087,1797,1824]" pageId="210" pageNumber="211">MCZC</collectionCode>
]
<specimenCount box="[1102,1218,1797,1824]" pageId="210" pageNumber="211" type="worker">3 workers</specimenCount>
(MCZENT00016365) [
<collectionCode box="[505,593,1837,1864]" pageId="210" pageNumber="211">MCZC</collectionCode>
]
<specimenCount box="[606,719,1837,1864]" pageId="210" pageNumber="211" type="worker">3 workers</specimenCount>
(CASENT0758264) [
<collectionCode box="[965,1054,1837,1864]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="210" pageNumber="211" type="Museum">USNM</collectionCode>
]
<specimenCount box="[1067,1180,1837,1864]" pageId="210" pageNumber="211" type="worker">3 workers</specimenCount>
(CASENT0758265) [
<collectionCode box="[1426,1515,1837,1864]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="210" pageNumber="211" type="Museum">USNM</collectionCode>
]
<specimenCount pageId="210" pageNumber="211" type="worker">1 worker</specimenCount>
(images of CASENT0915381 examined on antweb.org) [
<collectionCode box="[1251,1351,1877,1904]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="210" pageNumber="211">MNHN</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[530,1328,234,261]" pageId="211" pageNumber="212" type="distribution">
<paragraph blockId="211.[498,1542,234,1935]" box="[530,1328,234,261]" pageId="211" pageNumber="212">
<emphasis bold="true" box="[530,755,234,260]" pageId="211" pageNumber="212">Geographic range:</emphasis>
Low elevations of
<collectingCountry box="[978,1040,234,260]" name="Cuba" pageId="211" pageNumber="212">Cuba</collectingCountry>
(Artemisia) (
<figureCitation box="[1201,1311,234,260]" captionStart="Figure 121" captionStartId="205.[116,181,1124,1146]" captionTargetBox="[90,1541,234,1111]" captionTargetId="figure-151@205.[90,1524,234,1096]" captionTargetPageId="205" captionText="Figure 121 Geographical and elevational distribution of the pastinifer group. (A) Temnothorax androsanus (B) T. nigricans (C) T. pastinifer (D) T. rutabulafer sp. nov. (E) T. schwarzi. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles; if present, types of synonyms are named; non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-121" figureDoi="http://doi.org/10.5281/zenodo.5102351" httpUri="https://zenodo.org/record/5102351/files/figure.png" pageId="211" pageNumber="212">Fig. 121B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="211" pageNumber="212" type="diagnosis">
<paragraph blockId="211.[498,1542,234,1935]" pageId="211" pageNumber="212">
<emphasis bold="true" box="[530,754,274,300]" pageId="211" pageNumber="212">Worker diagnosis:</emphasis>
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[763,1034,274,300]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[763,1034,274,300]" italics="true" pageId="211" pageNumber="212">Temnothorax nigricans</emphasis>
</taxonomicName>
can be separated from all other species in the
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[544,619,314,340]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[544,619,314,340]" italics="true" pageId="211" pageNumber="212">salvini</emphasis>
</taxonomicName>
clade by the following character combination: antennal scapes very long, surpassing the posterior margin of head by about one and a half times the maximum width of the antennal scape (SI 92100); anterior margin of clypeus entire; transition between anterior and dorsal faces of pronotum continuous, not marked by a transition or a weak transverse carina; in profile view, mesosoma strongly convex dorsally and compact (WLI 122132); in dorsal view, mesosoma posterior to the pronotum swollen, only slightly narrower than the pronotum; dorsum of propodeum with setae; in dorsal view, propodeal spines broadly approximated basally, negative space between them “U” shaped; propodeal spines about as long as the propodeal declivity, directed posterodorsally, and weakly downcurved; hind femora weakly to moderately incrassate (FI 224291); petiolar node weakly squamiform, dorsally rounded; dorsum of petiole with four erect setae; in dorsal view, petiolar node slightly broader than the caudal cylinder (PNWI &lt;160); postpetiole moderately to very broad (PWI 212239); dorsum of head uniformly areolate; dorsum of postpetiole with&gt; 10 erect setae; integument bicolored: antennal funiculus, clypeus, and distal third of the femora testaceous yellow; head capsule dark brown; mandibles, mesosoma, legs, waist segments, and gaster light yellow.
</paragraph>
<paragraph blockId="211.[498,1542,234,1935]" pageId="211" pageNumber="212">
<emphasis bold="true" box="[530,717,951,977]" pageId="211" pageNumber="212">Similar species:</emphasis>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[725,1040,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="misomoschus" status="sp. nov.">
<emphasis box="[725,1040,951,977]" italics="true" pageId="211" pageNumber="212">Temnothorax misomoschus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1048,1140,952,978]" pageId="211" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1152,1299,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1152,1299,951,977]" italics="true" pageId="211" pageNumber="212">T. subditivus</emphasis>
</taxonomicName>
, and other members of the
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[573,681,992,1018]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[573,681,992,1018]" italics="true" pageId="211" pageNumber="212">pastinifer</emphasis>
</taxonomicName>
group.
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[771,1041,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[771,1041,991,1017]" italics="true" pageId="211" pageNumber="212">Temnothorax nigricans</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1358,1506,991,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1358,1506,991,1017]" italics="true" pageId="211" pageNumber="212">T. subditivus</emphasis>
</taxonomicName>
by the narrower petiolar node in dorsal view (PNWI &lt;160 vs.&gt;
<quantity box="[1231,1305,1031,1057]" metricMagnitude="0" metricUnit="m" metricValue="4.064" pageId="211" pageNumber="212" unit="in" value="160.0">160 in</quantity>
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1314,1463,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1314,1463,1031,1057]" italics="true" pageId="211" pageNumber="212">T. subditivus</emphasis>
</taxonomicName>
), the somewhat shorter petiolar peduncle (~2/3 of the total petiole length in
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1340,1478,1072,1097]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[1340,1478,1072,1097]" italics="true" pageId="211" pageNumber="212">T. nigricans</emphasis>
</taxonomicName>
vs. ~ 3/
<quantity box="[546,593,1111,1137]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="211" pageNumber="212" unit="in" value="4.0">4 in</quantity>
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[602,751,1111,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[602,751,1111,1137]" italics="true" pageId="211" pageNumber="212">T. subditivus</emphasis>
</taxonomicName>
) and the sculpture of the head dorsum (uniformly areolate in
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,636,1152,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[498,636,1152,1177]" italics="true" pageId="211" pageNumber="212">T. nigricans</emphasis>
</taxonomicName>
vs. variable in
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[816,965,1151,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[816,965,1151,1177]" italics="true" pageId="211" pageNumber="212">T. subditivus</emphasis>
</taxonomicName>
, but never uniformly areolate). In contrast to
<emphasis box="[498,830,1191,1217]" italics="true" pageId="211" pageNumber="212">
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,636,1191,1216]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">T. nigricans</taxonomicName>
,
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[648,830,1191,1217]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="misomoschus" status="sp. nov.">T. misomoschus</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[838,927,1191,1217]" pageId="211" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
is devoid of erect setae on most surfaces of the body, has a depressed propodeum, and a posteriorly leaning, subquadrate petiolar node.
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,769,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[498,769,1270,1296]" italics="true" pageId="211" pageNumber="212">Temnothorax nigricans</emphasis>
</taxonomicName>
can be separated from other members of the
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1312,1420,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[1312,1420,1270,1296]" italics="true" pageId="211" pageNumber="212">pastinifer</emphasis>
</taxonomicName>
group by the areolate head (smooth and shining in
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[994,1127,1310,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis box="[994,1127,1310,1336]" italics="true" pageId="211" pageNumber="212">T. schwarzi</emphasis>
</taxonomicName>
), weakly convex anterior clypeus margin (strongly emarginate in
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[875,1016,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[875,1016,1350,1376]" italics="true" pageId="211" pageNumber="212">T. pastinifer</emphasis>
</taxonomicName>
), moderately long antennal scape, which surpasses the posterior margin of the head by about one and a half times the maximum width of the antennal scape (barely surpassing the posterior margin in
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1905" box="[1317,1481,1430,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="androsanus">
<emphasis box="[1317,1481,1430,1456]" italics="true" pageId="211" pageNumber="212">T. androsanus</emphasis>
</taxonomicName>
), the continuous transition between the anterior and dorsal faces of the pronotum in profile view (transition distinct in
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1905" box="[820,985,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="androsanus">
<emphasis box="[820,985,1509,1535]" italics="true" pageId="211" pageNumber="212">T. androsanus</emphasis>
</taxonomicName>
).
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1011,1282,1509,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[1011,1282,1509,1535]" italics="true" pageId="211" pageNumber="212">Temnothorax nigricans</emphasis>
</taxonomicName>
can also be separated from
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[564,726,1549,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="rutabulafer" status="sp. nov.">
<emphasis box="[564,726,1549,1575]" italics="true" pageId="211" pageNumber="212">T. rutabulafer</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[735,826,1550,1576]" pageId="211" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
by the relatively narrow petiolar node in dorsal view (PNWI 130149 vs.&gt;
<quantity box="[670,743,1589,1615]" metricMagnitude="0" metricUnit="m" metricValue="4.572" pageId="211" pageNumber="212" unit="in" value="180.0">180 in</quantity>
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[753,915,1589,1615]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="rutabulafer" status="sp. nov.">
<emphasis box="[753,915,1589,1615]" italics="true" pageId="211" pageNumber="212">T. rutabulafer</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[924,1019,1590,1616]" pageId="211" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="215" lastPageNumber="216" pageId="211" pageNumber="212" type="description">
<paragraph blockId="211.[498,1542,234,1935]" lastBlockId="212.[498,1542,234,1895]" lastPageId="212" lastPageNumber="213" pageId="211" pageNumber="212">
<emphasis bold="true" box="[530,1018,1629,1655]" pageId="211" pageNumber="212">
Worker measurements &amp; indices (
<emphasis bold="true" box="[941,957,1629,1655]" italics="true" pageId="211" pageNumber="212">n</emphasis>
= 7):
</emphasis>
SL =
<geoCoordinate box="[1084,1146,1629,1656]" degrees="0.454" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.454">0.454</geoCoordinate>
<geoCoordinate box="[1146,1223,1629,1656]" degrees="0.512" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.512">0.512</geoCoordinate>
(0.477); FRS =
<geoCoordinate box="[1402,1464,1629,1655]" degrees="0.169" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.169">0.169</geoCoordinate>
<geoCoordinate box="[1464,1541,1629,1655]" degrees="0.189" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.189">0.189</geoCoordinate>
(0.179); CW =
<geoCoordinate box="[678,740,1669,1696]" degrees="0.502" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.502">0.502</geoCoordinate>
<geoCoordinate box="[740,817,1669,1696]" degrees="0.573" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.573">0.573</geoCoordinate>
(0.537); CWb =
<geoCoordinate box="[1020,1082,1669,1696]" degrees="0.452" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.452">0.452</geoCoordinate>
<geoCoordinate box="[1082,1159,1669,1696]" degrees="0.524" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.524">0.524</geoCoordinate>
(0.489); PoOC =
<geoCoordinate box="[1372,1434,1669,1695]" degrees="0.211" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.211">0.211</geoCoordinate>
<geoCoordinate box="[1434,1511,1669,1695]" degrees="0.237" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.237">0.237</geoCoordinate>
(0.226); CL =
<geoCoordinate box="[665,727,1708,1735]" degrees="0.532" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.532">0.532</geoCoordinate>
<geoCoordinate box="[727,804,1709,1735]" degrees="0.602" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.602">0.602</geoCoordinate>
(0.570); EL =
<geoCoordinate box="[977,1039,1708,1735]" degrees="0.115" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.115">0.115</geoCoordinate>
<geoCoordinate box="[1039,1116,1709,1735]" degrees="0.131" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.131">0.131</geoCoordinate>
(0.123); EW =
<geoCoordinate box="[1302,1364,1709,1735]" degrees="0.090" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.09">0.090</geoCoordinate>
<geoCoordinate box="[1364,1441,1709,1735]" degrees="0.104" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.104">0.104</geoCoordinate>
(0.097); MD =
<geoCoordinate box="[580,642,1748,1775]" degrees="0.135" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.135">0.135</geoCoordinate>
<geoCoordinate box="[642,719,1748,1775]" degrees="0.154" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.154">0.154</geoCoordinate>
(0.142); WL =
<geoCoordinate box="[904,966,1748,1775]" degrees="0.574" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.574">0.574</geoCoordinate>
<geoCoordinate box="[966,1043,1748,1775]" degrees="0.667" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.667">0.667</geoCoordinate>
(0.619); SPST =
<geoCoordinate box="[1245,1307,1749,1775]" degrees="0.244" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.244">0.244</geoCoordinate>
<geoCoordinate box="[1308,1385,1749,1775]" degrees="0.289" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.289">0.289</geoCoordinate>
(0.263); MPST =
<geoCoordinate box="[607,669,1788,1814]" degrees="0.192" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.192">0.192</geoCoordinate>
<geoCoordinate box="[669,746,1788,1814]" degrees="0.233" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.233">0.233</geoCoordinate>
(0.213); PEL =
<geoCoordinate box="[935,997,1788,1814]" degrees="0.212" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.212">0.212</geoCoordinate>
<geoCoordinate box="[997,1074,1788,1814]" degrees="0.278" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.278">0.278</geoCoordinate>
(0.248); NOL =
<geoCoordinate box="[1274,1336,1788,1814]" degrees="0.131" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.131">0.131</geoCoordinate>
<geoCoordinate box="[1336,1413,1788,1814]" degrees="0.177" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.177">0.177</geoCoordinate>
(0.149); NOH =
<geoCoordinate box="[598,660,1828,1855]" degrees="0.105" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.105">0.105</geoCoordinate>
<geoCoordinate box="[660,737,1828,1854]" degrees="0.141" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.141">0.141</geoCoordinate>
(0.120); PEH =
<geoCoordinate box="[933,995,1828,1854]" degrees="0.182" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.182">0.182</geoCoordinate>
<geoCoordinate box="[995,1072,1828,1854]" degrees="0.234" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.234">0.234</geoCoordinate>
(0.210); PPL =
<geoCoordinate box="[1262,1324,1828,1854]" degrees="0.131" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.131">0.131</geoCoordinate>
<geoCoordinate box="[1324,1401,1828,1855]" degrees="0.158" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.158">0.158</geoCoordinate>
(0.147); PPH =
<geoCoordinate box="[588,650,1868,1894]" degrees="0.163" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.163">0.163</geoCoordinate>
<geoCoordinate box="[650,727,1868,1894]" degrees="0.198" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.198">0.198</geoCoordinate>
(0.182); PW =
<geoCoordinate box="[912,974,1868,1894]" degrees="0.347" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.347">0.347</geoCoordinate>
<geoCoordinate box="[975,1052,1868,1894]" degrees="0.393" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.393">0.393</geoCoordinate>
(0.369); SBPA =
<geoCoordinate box="[1260,1322,1868,1894]" degrees="0.143" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.143">0.143</geoCoordinate>
<geoCoordinate box="[1322,1399,1868,1895]" degrees="0.175" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.175">0.175</geoCoordinate>
(0.156); SPTI =
<geoCoordinate box="[590,652,1908,1934]" degrees="0.226" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.226">0.226</geoCoordinate>
<geoCoordinate box="[653,730,1908,1934]" degrees="0.264" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.264">0.264</geoCoordinate>
(0.244); PEW =
<geoCoordinate box="[932,994,1908,1934]" degrees="0.110" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.11">0.110</geoCoordinate>
<geoCoordinate box="[994,1071,1908,1934]" degrees="0.142" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.142">0.142</geoCoordinate>
(0.124); PNW =
<geoCoordinate box="[1278,1340,1908,1935]" degrees="0.154" direction="north" orientation="latitude" pageId="211" pageNumber="212" precision="55" value="0.154">0.154</geoCoordinate>
<geoCoordinate box="[1340,1417,1908,1934]" degrees="0.189" direction="west" orientation="longitude" pageId="211" pageNumber="212" precision="55" value="-0.189">0.189</geoCoordinate>
(0.173); PPW =
<geoCoordinate box="[590,652,234,261]" degrees="0.259" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="55" value="0.259">0.259</geoCoordinate>
<geoCoordinate box="[653,730,234,260]" degrees="0.303" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="55" value="-0.303">0.303</geoCoordinate>
(0.283); HFL =
<geoCoordinate box="[917,979,234,260]" degrees="0.419" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="55" value="0.419">0.419</geoCoordinate>
<geoCoordinate box="[979,1056,234,260]" degrees="0.491" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="55" value="-0.491">0.491</geoCoordinate>
(0.465); HFWmax =
<geoCoordinate box="[1306,1368,234,260]" degrees="0.122" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="55" value="0.122">0.122</geoCoordinate>
<geoCoordinate box="[1368,1445,234,261]" degrees="0.153" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="55" value="-0.153">0.153</geoCoordinate>
(0.136); HFWmin =
<geoCoordinate box="[645,707,274,300]" degrees="0.047" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="55" value="0.047">0.047</geoCoordinate>
<geoCoordinate box="[708,785,274,301]" degrees="0.059" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="55" value="-0.059">0.059</geoCoordinate>
(0.054); CS =
<geoCoordinate box="[958,1020,274,300]" degrees="0.718" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="55" value="0.718">0.718</geoCoordinate>
<geoCoordinate box="[1021,1098,274,301]" degrees="0.825" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="55" value="-0.825">0.825</geoCoordinate>
(0.774); ES =
<geoCoordinate box="[1269,1331,274,300]" degrees="0.160" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="55" value="0.16">0.160</geoCoordinate>
<geoCoordinate box="[1331,1408,274,300]" degrees="0.183" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="55" value="-0.183">0.183</geoCoordinate>
(0.172); SI = 92100 (98); OI = 2223 (22); CI = 8489 (86); WLI = 122132 (127); SBI = 3035 (32); PSI = 4145 (42); PWI = 212239 (229); PLI = 159180 (168); NI = 115131 (124); PNWI = 130149 (140); NLI = 5571 (60); FI = 224291 (254).
</paragraph>
<paragraph blockId="212.[498,1542,234,1895]" pageId="212" pageNumber="213">
<emphasis bold="true" box="[530,775,433,460]" pageId="212" pageNumber="213">Worker description:</emphasis>
In full-face view, head subquadrate, longer than broad (CI 8489). Mandibles finely striate, shining, and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin flat to evenly convex medially. Antennal scapes very long: when fully retracted, surpassing the posterior margin of the head capsule by about one and a half times the maximum width of the antennal scape (SI 92100). Antennae 12-segmented; antennal club of three segments, with the apical-most segment longer than the preceding two in combination. Frontal carinae short, extending past the antennal toruli by about two times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head weakly convex, converging evenly to the mandibular insertions. Posterior head margin flat to very slightly concave, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="212.[498,1542,234,1895]" pageId="212" pageNumber="213">
In profile view, compound eyes ovular and moderately large (OI 2223), with 12 ommatidia in longest row. Pronotal declivity indistinct, but pronotal neck and anterior face of pronotum forming a rounded ~120
<emphasis box="[1000,1010,987,1005]" italics="true" pageId="212" pageNumber="213">
<superScript attach="left" box="[1000,1010,987,1005]" fontSize="8" pageId="212" pageNumber="213">°</superScript>
</emphasis>
angle. Mesosoma evenly, but weakly, convex dorsally from where it joins the pronotal neck to the propodeal spines. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which moderately well developed. Metanotal groove nearly entirely absent: only visible as a faint disruption in the surface sculpture between meso- and metacoxal insertions to the minute metathoracic spiracle, which is indistinguishable from the ground sculpture. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about two and a half spiracle diameters. Propodeal spines well developed and long (PSI 4145), longer than the propodeal declivity, tapering evenly from the base, downcurved for most of their length but turning up slightly at the tips, and acute. Propodeal declivity weakly concave, forming a rounded ~110
<emphasis box="[1482,1492,1385,1403]" italics="true" pageId="212" pageNumber="213">
<superScript attach="left" box="[1482,1492,1385,1403]" fontSize="8" pageId="212" pageNumber="213">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed, but with an angulate dorsal flange. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole moderately long (PLI 159180), without tubercles anterodorsally. Subpetiolar process in the form of a weakly developed, acute tooth, which grades evenly into the ventral margin of the petiole posteriorly. Ventral surface of petiole bulging medially. Petiolar peduncle long, comprising about two thirds of the total length of the petiole. Petiolar node squamiform: transition between peduncle and node marked by a rounded angle of ~110
<emphasis box="[659,669,1744,1762]" italics="true" pageId="212" pageNumber="213">
<superScript attach="none" box="[659,669,1744,1762]" fontSize="8" pageId="212" pageNumber="213">°</superScript>
</emphasis>
; anterior face of the node is straight and does not overhang the peduncle; anterior face rounds evenly into the strongly convex dorsal face, which rounds evenly into the posterior face. Postpetiole evenly rounded anterodorsally, before flattening posterodorsally; flat ventrally.
</paragraph>
<paragraph blockId="213.[498,1542,234,1935]" pageId="213" pageNumber="214">In dorsal view, humeri very weakly developed: rounded and only slightly wider than the rest of the mesosoma; mesothoracic spiracles not protruding past the lateral margins of the mesosoma. Metanotal groove absent: mesonotum and propodeum completely fused and converging evenly to the bases of the propodeal spines. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by slightly less than their length; negative space between them a “U”. Petiolar peduncle with spiracles not protruding past the lateral margins. Petiolar node squamiform: rounded anteriorly, with a flattened posterior face, much wider than the peduncle, and narrowing into the caudal cylinder, which is narrower than the node. Postpetiole moderately to very broad (PWI 212239) and campaniform, articulating with nearly the entire anterior margin of the gaster, but leaving angulate corners of the gaster exposed on each side. Anterior margin of the postpetiole broadly convex, meeting the lateral margins at rounded angles; lateral margins diverging evenly posteriorly; posterior margin flat. Metafemur weakly to moderately incrassate (FI 224291).</paragraph>
<paragraph blockId="213.[498,1542,234,1935]" pageId="213" pageNumber="214">Sculpture: median clypeal carina present, flanked by two slightly weaker carinae over areolate-costulate ground sculpture. Antennal scapes weakly areolate and dull. Cephalic dorsum and lateral surfaces of head with dense areolae arranged into longitudinal rows by costulae; ventral surface shining through weaker areolate sculpture. Pronotal neck areolate. Lateral face of pronotum with sculpture similar to the cephalic dorsum. Lateral face of propodeum, meso- and metapleurae uniformly areolate. Dorsally, mesosoma sculptured similarly to the cephalic dorsum. Areolate sculpture weaker on the propodeal declivity. Femora weakly shining through areolate sculpture. Petiolar peduncle, venter, anterior, and posterior faces of the petiolar node weakly shining through shallow areolate sculpture; dorsal face of the petiolar node more densely areolate. Dorsal surface of postpetiole dull with areolate sculpture and costulae. Gaster smooth and shining, with weak spectral iridescence on the first tergite. Surface of the first gastral sternite smooth and shining.</paragraph>
<paragraph blockId="213.[498,1542,234,1935]" pageId="213" pageNumber="214">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of head, mesosoma, waist segments and gaster with short, erect, blunt-tipped setae, the longest of which are roughly the width of the compound eye. The head bears ~32, mesosoma ~22, petiole 6, postpetiole ~10, and first gastral tergite ~36 setae. Sparse, adpressed pubescence present on the entire body, but may be difficult to discern from the dense ground sculpture and light integument on most of the body.</paragraph>
<paragraph blockId="213.[498,1542,234,1935]" pageId="213" pageNumber="214">Color: funiculus of the antenna, clypeus, and distal third of the femora testaceous yellow. Head capsule dark brown. Mandibles, mesosoma, legs, waist segments, and gaster light yellow.</paragraph>
<paragraph blockId="213.[498,1542,234,1935]" lastBlockId="214.[498,1542,234,1934]" lastPageId="214" lastPageNumber="215" pageId="213" pageNumber="214">
<emphasis bold="true" box="[530,1001,1668,1695]" pageId="213" pageNumber="214">
Gyne measurements &amp; indices (
<emphasis bold="true" box="[919,935,1669,1695]" italics="true" pageId="213" pageNumber="214">n</emphasis>
= 2):
</emphasis>
SL =
<geoCoordinate box="[1073,1135,1669,1695]" degrees="0.469" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.469">0.469</geoCoordinate>
<geoCoordinate box="[1136,1213,1669,1696]" degrees="0.520" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.52">0.520</geoCoordinate>
(0.495); FRS =
<geoCoordinate box="[1400,1462,1669,1695]" degrees="0.212" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.212">0.212</geoCoordinate>
<geoCoordinate box="[1463,1539,1669,1695]" degrees="0.230" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.23">0.230</geoCoordinate>
(0.221); CW =
<geoCoordinate box="[678,740,1708,1734]" degrees="0.637" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.637">0.637</geoCoordinate>
<geoCoordinate box="[740,817,1708,1735]" degrees="0.687" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.687">0.687</geoCoordinate>
(0.662); CWb =
<geoCoordinate box="[1020,1082,1708,1735]" degrees="0.590" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.59">0.590</geoCoordinate>
<geoCoordinate box="[1082,1159,1708,1735]" degrees="0.646" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.646">0.646</geoCoordinate>
(0.618); PoOC =
<geoCoordinate box="[1372,1434,1708,1735]" degrees="0.245" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.245">0.245</geoCoordinate>
<geoCoordinate box="[1434,1511,1708,1735]" degrees="0.251" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.251">0.251</geoCoordinate>
(0.248); CL =
<geoCoordinate box="[665,727,1748,1774]" degrees="0.627" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.627">0.627</geoCoordinate>
<geoCoordinate box="[727,804,1748,1775]" degrees="0.675" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.675">0.675</geoCoordinate>
(0.651); EL =
<geoCoordinate box="[977,1039,1749,1775]" degrees="0.173" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.173">0.173</geoCoordinate>
<geoCoordinate box="[1039,1116,1749,1775]" degrees="0.183" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.183">0.183</geoCoordinate>
(0.178); EW =
<geoCoordinate box="[1302,1364,1749,1775]" degrees="0.132" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.132">0.132</geoCoordinate>
<geoCoordinate box="[1364,1441,1748,1775]" degrees="0.158" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.158">0.158</geoCoordinate>
(0.145); MD =
<geoCoordinate box="[580,642,1788,1815]" degrees="0.155" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.155">0.155</geoCoordinate>
<geoCoordinate box="[642,719,1788,1814]" degrees="0.160" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.16">0.160</geoCoordinate>
(0.158); WL =
<geoCoordinate box="[904,966,1788,1814]" degrees="0.903" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.903">0.903</geoCoordinate>
<geoCoordinate box="[966,1043,1788,1814]" degrees="1.008" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-1.008">1.008</geoCoordinate>
(0.956); SPST =
<geoCoordinate box="[1245,1307,1788,1815]" degrees="0.259" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.259">0.259</geoCoordinate>
<geoCoordinate box="[1308,1385,1788,1814]" degrees="0.269" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.269">0.269</geoCoordinate>
(0.264); MPST =
<geoCoordinate box="[607,669,1828,1854]" degrees="0.262" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.262">0.262</geoCoordinate>
<geoCoordinate box="[669,746,1828,1855]" degrees="0.265" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.265">0.265</geoCoordinate>
(0.264); PEL =
<geoCoordinate box="[935,997,1828,1854]" degrees="0.287" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.287">0.287</geoCoordinate>
<geoCoordinate box="[997,1074,1828,1854]" degrees="0.317" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.317">0.317</geoCoordinate>
(0.302); NOL =
<geoCoordinate box="[1274,1336,1828,1854]" degrees="0.160" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.16">0.160</geoCoordinate>
<geoCoordinate box="[1336,1413,1828,1855]" degrees="0.175" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.175">0.175</geoCoordinate>
(0.168); NOH =
<geoCoordinate box="[598,660,1868,1894]" degrees="0.131" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.131">0.131</geoCoordinate>
<geoCoordinate box="[660,737,1868,1894]" degrees="0.164" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.164">0.164</geoCoordinate>
(0.148); PEH =
<geoCoordinate box="[933,995,1868,1895]" degrees="0.257" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.257">0.257</geoCoordinate>
<geoCoordinate box="[995,1072,1868,1894]" degrees="0.287" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.287">0.287</geoCoordinate>
(0.272); PPL =
<geoCoordinate box="[1262,1324,1868,1894]" degrees="0.182" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.182">0.182</geoCoordinate>
<geoCoordinate box="[1324,1401,1868,1894]" degrees="0.187" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.187">0.187</geoCoordinate>
(0.185); PPH =
<geoCoordinate box="[588,650,1908,1934]" degrees="0.226" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.226">0.226</geoCoordinate>
<geoCoordinate box="[650,727,1908,1935]" degrees="0.250" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.25">0.250</geoCoordinate>
(0.238); PW =
<geoCoordinate box="[912,974,1908,1935]" degrees="0.554" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.554">0.554</geoCoordinate>
<geoCoordinate box="[975,1052,1908,1934]" degrees="0.628" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.628">0.628</geoCoordinate>
(0.591); SBPA =
<geoCoordinate box="[1260,1322,1908,1934]" degrees="0.287" direction="north" orientation="latitude" pageId="213" pageNumber="214" precision="55" value="0.287">0.287</geoCoordinate>
<geoCoordinate box="[1322,1399,1908,1935]" degrees="0.345" direction="west" orientation="longitude" pageId="213" pageNumber="214" precision="55" value="-0.345">0.345</geoCoordinate>
(0.316); SPTI =
<geoCoordinate box="[590,652,234,260]" degrees="0.273" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.273">0.273</geoCoordinate>
<geoCoordinate box="[653,730,234,260]" degrees="0.312" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.312">0.312</geoCoordinate>
(0.293); PEW =
<geoCoordinate box="[932,994,234,260]" degrees="0.149" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.149">0.149</geoCoordinate>
<geoCoordinate box="[994,1071,234,260]" degrees="0.174" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.174">0.174</geoCoordinate>
(0.162); PNW =
<geoCoordinate box="[1278,1340,234,260]" degrees="0.194" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.194">0.194</geoCoordinate>
<geoCoordinate box="[1340,1417,234,260]" degrees="0.213" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.213">0.213</geoCoordinate>
(0.204); PPW =
<geoCoordinate box="[590,652,274,300]" degrees="0.371" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.371">0.371</geoCoordinate>
<geoCoordinate box="[653,730,274,300]" degrees="0.432" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.432">0.432</geoCoordinate>
(0.402); HFL =
<geoCoordinate box="[917,979,274,301]" degrees="0.535" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.535">0.535</geoCoordinate>
<geoCoordinate box="[979,1056,274,301]" degrees="0.589" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.589">0.589</geoCoordinate>
(0.562); HFWmax =
<geoCoordinate box="[1306,1368,274,300]" degrees="0.128" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.128">0.128</geoCoordinate>
<geoCoordinate box="[1368,1445,274,300]" degrees="0.144" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.144">0.144</geoCoordinate>
(0.136); HFWmin =
<geoCoordinate box="[645,707,314,341]" degrees="0.056" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.056">0.056</geoCoordinate>
<geoCoordinate box="[708,785,314,341]" degrees="0.059" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.059">0.059</geoCoordinate>
(0.058); CS =
<geoCoordinate box="[958,1020,314,340]" degrees="0.904" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.904">0.904</geoCoordinate>
<geoCoordinate box="[1021,1098,314,340]" degrees="0.984" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.984">0.984</geoCoordinate>
(0.944); ES =
<geoCoordinate box="[1269,1331,314,340]" degrees="0.239" direction="north" orientation="latitude" pageId="214" pageNumber="215" precision="55" value="0.239">0.239</geoCoordinate>
<geoCoordinate box="[1331,1408,314,340]" degrees="0.262" direction="west" orientation="longitude" pageId="214" pageNumber="215" precision="55" value="-0.262">0.262</geoCoordinate>
(0.251); SI = 7980 (80); OI = 2627 (27); CI = 9496 (95); WLI = 153156 (155); SBI = 4953 (51); PSI = 2729 (28); PWI = 248249 (249); PLI = 153174 (164); NI = 107122 (114); PNWI = 122130 (126); NLI = 5556 (55); FI = 229244 (236).
</paragraph>
<paragraph blockId="214.[498,1542,234,1934]" pageId="214" pageNumber="215">
<emphasis bold="true" box="[530,747,473,499]" pageId="214" pageNumber="215">Gyne description:</emphasis>
In full-face view, head subquadrate, about as long as broad (CI 9496). Mandibles densely, finely striate, weakly shining, and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin flat to weakly emarginated medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 7980). Antennae 12-segmented; antennal club composed of three segments, with the apical-most longer than the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head evenly convex, converging from below the compound eyes to the mandibular insertions. Posterior head margin flat, rounding evenly into the lateral margins.
</paragraph>
<paragraph blockId="214.[498,1542,234,1934]" pageId="214" pageNumber="215">
In profile view, compound eyes ovular and large (OI 2627), with 17 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, barely covering the dorsal surface of the pronotum, and weakly convex dorsally. Mesoscutellum slightly depressed below the level of the mesoscutum. Posterior margin of metanotum extending slightly past the posterior margin of the mesoscutum, the two sclerites form an even convexity. Propodeal spiracle well developed, directed posterolaterally, and separated from the propodeal declivity by about three and a half spiracle diameters. Propodeal spines stout, well developed, and moderately long (PSI 2729), about as long as the propodeal declivity, tapering evenly from the base, directed posteriorly, straight, and acute. Propodeal declivity weakly concave, forming a rounded ~90
<emphasis box="[972,982,1306,1324]" italics="true" pageId="214" pageNumber="215">
<superScript attach="left" box="[972,982,1306,1324]" fontSize="8" pageId="214" pageNumber="215">°</superScript>
</emphasis>
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed, but with a flange that is slightly angulate dorsally. Metapleural gland bulla moderately large, extending from the metacoxal insertion two thirds of the way to the propodeal spiracle. Petiole moderately long (PLI 153174), with peduncle evenly rounded anterodorsally. Subpetiolar process in the form of a weakly developed, acute tooth, which grades evenly into the ventral margin of the petiole posteriorly. Petiolar peduncle long, comprising about two thirds of the total length of the petiole. Petiolar node squamiform: transition between peduncle and node marked by a rounded angle of ~120
<emphasis box="[764,774,1624,1642]" italics="true" pageId="214" pageNumber="215">
<superScript attach="none" box="[764,774,1624,1642]" fontSize="8" pageId="214" pageNumber="215">°</superScript>
</emphasis>
; anterior face transitioning joining the short dorsal face at a ~90
<emphasis box="[1530,1540,1624,1642]" italics="true" pageId="214" pageNumber="215">
<superScript attach="none" box="[1530,1540,1624,1642]" fontSize="8" pageId="214" pageNumber="215">°</superScript>
</emphasis>
angle, which rounds evenly into the posterior face. Postpetiole evenly rounded anterodorsally, bulging slightly before it transitions into the flattened dorsal face; ventral surface weakly lobed.
</paragraph>
<paragraph blockId="214.[498,1542,234,1934]" lastBlockId="215.[498,1542,234,1934]" lastPageId="215" lastPageNumber="216" pageId="214" pageNumber="215">In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as rounded sclerites. Propodeal spines weakly diverging basally but parallel apically, their apices separated from each other by about one and a half times their length; negative space between them a broad “U”. Petiolar peduncle with spiracles weakly protruding past the lateral margins. Petiolar node squamiform: flat anteriorly, with lateral faces strongly convex, and with a flattened posterior face, much wider than the peduncle, and narrowing into the caudal cylinder, which is narrower than the node. Postpetiole very broad (PWI 248249) and campaniform, articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side exposed. Anterior margin of postpetiole weakly convex, with corners marked by rounded angles as it transitions to the lateral margins, which evenly diverge to the angulate posterior corners; posterior margin flat. Metafemur weakly incrassate (FI 229244).</paragraph>
<paragraph blockId="215.[498,1542,234,1934]" pageId="215" pageNumber="216">Sculpture: median clypeal lobe with about seven equally strong fine carinae, extending from the anterior margin nearly to frontal triangle. Lateral clypeal lobes with additional fine carinae; ground sculpture weakly areolate. Antennal scapes weakly shining through areolate sculpture. Cephalic dorsum and lateral surfaces of head with dense costulae over areolate sculpture; ventral surface shining through weaker areolate-costulate sculpture. Pronotal neck areolate. Pronotum, a posterior two thirds of the anepisternum and katepisternum, and lateral face of the propodeum with sculpture similar to that of the cephalic dorsum, but slightly weaker. Propodeal declivity weakly areolate-costulate. Mesoscutum and mesoscutellum with sculpture similar to that of the cephalic dorsum. Mesoscutellum smooth and shining medially, surrounded by costulae and areolae. Femora shining through weak areolate sculpture. Petiole with weak areolate sculpture on all surfaces, becoming stronger on the dorsoposterior surface of the node. Postpetiole densely areolate dorsally. Gaster predominantly smooth and shining, but with the anterior of the first gastral tergite finely areolate-costulate, and with faint spectral iridescence. Surface of the first gastral sternite smooth and shining.</paragraph>
<paragraph blockId="215.[498,1542,234,1934]" pageId="215" pageNumber="216">Setae: antennal scapes and funiculi with short, adpressed pilosity. Dorsum of head, mesosoma, waist segments and gaster with short, erect, blunt-tipped setae, the longest of which are roughly three quarters of the compound eye width. Sparse, adpressed pubescence present on the entire body, but may be difficult to discern from the dense ground sculpture and light integument on most of the body.</paragraph>
<paragraph blockId="215.[498,1542,234,1934]" pageId="215" pageNumber="216">Color: funiculus of the antenna, clypeus, and distal third of the femora testaceous yellow. Meso- and metapleurae, anteromedial and posterolateral portions of the mesoscutum, posterior edge of the mesoscutellum, metanotum, and posterior third of all gastral sclerites testaceous. Head capsule dark brown. Mandibles, mesosoma, legs, waist segments, and gaster light yellow.</paragraph>
<paragraph blockId="215.[498,1542,234,1934]" box="[530,730,1549,1575]" pageId="215" pageNumber="216">
<emphasis bold="true" box="[530,597,1549,1575]" pageId="215" pageNumber="216">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="215" pageNumber="216" type="etymology">
<paragraph blockId="215.[498,1542,234,1934]" pageId="215" pageNumber="216">
<emphasis bold="true" box="[530,668,1589,1615]" pageId="215" pageNumber="216">Etymology:</emphasis>
Morphological, from the Latin nigricans (= blackish), presumably in reference to the dark head.
</paragraph>
</subSubSection>
<subSubSection lastPageId="216" lastPageNumber="217" pageId="215" pageNumber="216" type="discussion">
<paragraph blockId="215.[498,1542,234,1934]" lastBlockId="216.[498,1542,1359,1823]" lastPageId="216" lastPageNumber="217" pageId="215" pageNumber="216">
<emphasis bold="true" box="[530,670,1669,1695]" pageId="215" pageNumber="216">Comments:</emphasis>
In the original description by
<bibRefCitation author="Wheeler WM" box="[1033,1210,1669,1695]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="215" pageNumber="216" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[1033,1210,1669,1695]" italics="true" pageId="215" pageNumber="216">Wheeler (1931)</emphasis>
</bibRefCitation>
, the
<typeStatus box="[1271,1321,1669,1695]" pageId="215" pageNumber="216">type</typeStatus>
series of
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[498,768,1709,1735]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis box="[498,768,1709,1735]" italics="true" pageId="215" pageNumber="216">Temnothorax nigricans</emphasis>
</taxonomicName>
was collected from the ground in a fruit of
<taxonomicName box="[1281,1514,1709,1735]" class="Magnoliopsida" family="Meliaceae" genus="Swietenia" kingdom="Plantae" order="Sapindales" pageId="215" pageNumber="216" phylum="Tracheophyta" rank="species" species="mahogani">
<emphasis box="[1281,1514,1709,1735]" italics="true" pageId="215" pageNumber="216">Swietenia mahogani</emphasis>
</taxonomicName>
, a hardwood species endemic to the Caribbean islands and southern Florida. The
<typeStatus box="[1435,1485,1749,1775]" pageId="215" pageNumber="216">type</typeStatus>
locality is in northern Artemisia province on the west side of
<collectingCountry box="[1227,1291,1788,1814]" name="Cuba" pageId="215" pageNumber="216">Cuba</collectingCountry>
, which experiences a dry season typical of many islands in the Caribbean but receives substantial precipitation year-round, leading to this area being classified as having a tropical rainforest climate. In addition to the
<typeStatus box="[716,766,1908,1934]" pageId="215" pageNumber="216">type</typeStatus>
locality, this species has been reported from numerous other localities across the island by
<bibRefCitation author="Alayo P." box="[848,998,1359,1385]" journalOrPublisher="Series Biologia" pageId="216" pageNumber="217" pagination="1 - 58" part="53" refId="ref269672" refString="Alayo P. 1974. Introduccion al estudio de los hymenopteros de Cuba. Superfamilia Formicoidea. Series Biologia 53: 1 - 58." title="Introduccion al estudio de los hymenopteros de Cuba. Superfamilia Formicoidea" type="journal article" year="1974">
<emphasis box="[848,998,1359,1385]" italics="true" pageId="216" pageNumber="217">Alayo (1974)</emphasis>
</bibRefCitation>
, including Ciénaga de Zapata,
<collectingRegion box="[1365,1496,1359,1385]" country="Cuba" name="Cienfuegos" pageId="216" pageNumber="217">Cienfuegos</collectingRegion>
, Caibarién, Buenos Aires, Escambray, and Seibabo. I have only been able to examine the original nest series, but the presence of multiple gynes suggests that this species may be polygynous. In his remarks on this species,
<bibRefCitation author="Wheeler WM" box="[1009,1185,1478,1505]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="216" pageNumber="217" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
<emphasis box="[1009,1185,1478,1505]" italics="true" pageId="216" pageNumber="217">Wheeler (1931)</emphasis>
</bibRefCitation>
mused about the resemblance of the structure of the mesosoma to
<taxonomicName authority="Mann" authorityName="Mann" authorityYear="1920" box="[929,1194,1518,1545]" class="Insecta" family="Formicidae" genus="Antillaemyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis box="[929,1114,1518,1544]" italics="true" pageId="216" pageNumber="217">Antillaemyrmex</emphasis>
Mann
</taxonomicName>
(a now defunct genus applied to what is presently referred to as the
<taxonomicName baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1043,1156,1558,1584]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis box="[1043,1156,1558,1584]" italics="true" pageId="216" pageNumber="217">pulchellus</emphasis>
</taxonomicName>
group +
<taxonomicName authority="(Mann))" authorityName="" baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1268,1521,1558,1584]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="allardycei">
<emphasis box="[1268,1412,1558,1584]" italics="true" pageId="216" pageNumber="217">T. allardycei</emphasis>
(Mann))
</taxonomicName>
and in petiolar shape to
<taxonomicName authorityName="Roger" authorityYear="1863" box="[788,944,1598,1624]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis box="[788,944,1598,1624]" italics="true" pageId="216" pageNumber="217">Macromischa</emphasis>
</taxonomicName>
(probably referring
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1185,1325,1598,1624]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[1185,1325,1598,1624]" italics="true" pageId="216" pageNumber="217">T. pastinifer</emphasis>
</taxonomicName>
). Our current understanding is that
<taxonomicName authorityName="Roger" authorityYear="1863" box="[750,906,1638,1664]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis box="[750,906,1638,1664]" italics="true" pageId="216" pageNumber="217">Macromischa</emphasis>
</taxonomicName>
, as defined by
<bibRefCitation author="Mann WM" box="[1074,1224,1638,1664]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="216" pageNumber="217" pagination="403 - 439" part="42" refId="ref271797" refString="Mann WM. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403 - 439." title="Additions to the ant fauna of the West Indies and Central America" type="journal article" year="1920">
<emphasis box="[1074,1224,1638,1664]" italics="true" pageId="216" pageNumber="217">Mann (1920)</emphasis>
</bibRefCitation>
with
<taxonomicName authority="(Roger)" baseAuthorityName="Roger" baseAuthorityYear="1863" box="[1288,1542,1638,1664]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="purpuratus">
<emphasis box="[1288,1447,1639,1664]" italics="true" pageId="216" pageNumber="217">T. purpuratus</emphasis>
(Roger)
</taxonomicName>
as the
<typeStatus box="[571,621,1678,1704]" pageId="216" pageNumber="217">type</typeStatus>
for the genus, is polyphyletic: namely
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1061,1207,1678,1704]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="subditivus">
<emphasis box="[1061,1207,1678,1704]" italics="true" pageId="216" pageNumber="217">T. subditivus</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1905" box="[1220,1384,1678,1704]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="androsanus">
<emphasis box="[1220,1384,1678,1704]" italics="true" pageId="216" pageNumber="217">T. androsanus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Prebus" authorityYear="2021" baseAuthorityName="Emery" baseAuthorityYear="1894" box="[1397,1535,1678,1704]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="pastinifer">
<emphasis box="[1397,1535,1678,1704]" italics="true" pageId="216" pageNumber="217">T. pastinifer</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Prebus" authorityYear="2017" box="[498,623,1717,1743]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis box="[498,623,1717,1743]" italics="true" pageId="216" pageNumber="217">T. fuscatus</emphasis>
</taxonomicName>
, and
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[691,799,1717,1743]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="salvini">
<emphasis box="[691,799,1717,1743]" italics="true" pageId="216" pageNumber="217">T. salvini</emphasis>
</taxonomicName>
are all more closely related to
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1165,1292,1717,1743]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="terricola">
<emphasis box="[1165,1292,1717,1743]" italics="true" pageId="216" pageNumber="217">T. terricola</emphasis>
</taxonomicName>
than they are to
<taxonomicName baseAuthorityName="Roger" baseAuthorityYear="1863" box="[498,660,1758,1783]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="purpuratus">
<emphasis box="[498,660,1758,1783]" italics="true" pageId="216" pageNumber="217">T. purpuratus</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Prebus M." box="[722,870,1757,1783]" journalOrPublisher="BMC Evolutionary Biology" pageId="216" pageNumber="217" pagination="250" part="17" refId="ref272195" refString="Prebus M. 2017. Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae). BMC Evolutionary Biology 17 (1): 250 DOI 10.1186 / s 12862 - 017 - 1095 - 8." title="Insights into the evolution, biogeography and natural history of the acorn ants, genus Temnothorax Mayr (Hymenoptera: Formicidae)" type="journal article" year="2017">
<emphasis box="[722,870,1757,1783]" italics="true" pageId="216" pageNumber="217">Prebus, 2017</emphasis>
</bibRefCitation>
), mostly concurring with
<bibRefCitation author="Fontenla Rizo JL" box="[1180,1424,1757,1783]" journalOrPublisher="Avicennia" pageId="216" pageNumber="217" pagination="35 - 44" part="13" refId="ref270970" refString="Fontenla Rizo JL. 2000. Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax). Avicennia 13: 35 - 44." title="Definition, phylogenetic relationships and morphological species groups of taxon &quot; Macromischa &quot; (Hymenoptera: Formicidae: Leptothorax)" type="journal article" year="2000">
<emphasis box="[1180,1424,1757,1783]" italics="true" pageId="216" pageNumber="217">Fontenla Rizo (2000)</emphasis>
</bibRefCitation>
, which inferred phylogenetic relationships within
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[999,1156,1797,1823]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis box="[999,1156,1797,1823]" italics="true" pageId="216" pageNumber="217">Temnothorax</emphasis>
</taxonomicName>
using morphology.
</paragraph>
</subSubSection>
</treatment>
</document>