184 lines
22 KiB
XML
184 lines
22 KiB
XML
<document id="E8BF8B15780D900B411E9E3928EE0E77" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="FDBB31990A31CB6C53431B0CAAA30B2B" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus comes Guignot 1955" docType="treatment" docVersion="5" lastPageNumber="33" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="31" updateTime="1701314941532" updateUser="plazi">
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<mods:title id="14B7B34C69E8D097BB77336286C6A1DC">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="323EEE5D0312EF9DEB9323F43F79FF25">Bistroem, Olof</mods:namePart>
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<mods:namePart id="B09F18E8D6EDCDA998D10FBC8F8193EE">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="B038044DBE8B57AE5D036AE03DB4BE6C">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="B5D2F5B07011A62253FB4C8BA318D0B8">text</mods:typeOfResource>
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<mods:title id="00BF412F6673EB5E88FF0854EE242E33">ZooKeys</mods:title>
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<mods:date id="B2550EA713B8161A742D757B4D54C3C3">2015</mods:date>
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<mods:detail id="024F9D8EEB4BF46AB7B7D6CE750105F8" type="volume">
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<mods:number id="9235A27815E07A4C72B7837EDB84DEF4">542</mods:number>
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<mods:end id="D06E6E657517C63CBBD7D14B49742A22">379</mods:end>
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<mods:url id="3A56CA2DCAF7E38866D45162D9DF9B86">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:identifier id="D2AD21D1D4676B00A94276DA5A0DCC0F" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<mods:identifier id="A603169B584699165C0F2FDE903E1202" type="Pensoft-Pub">1313-2970-542-1</mods:identifier>
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<treatment id="FDBB31990A31CB6C53431B0CAAA30B2B" ID-GBIF-Taxon="127862698" LSID="urn:lsid:plazi:treatment:FDBB31990A31CB6C53431B0CAAA30B2B" httpUri="http://treatment.plazi.org/id/FDBB31990A31CB6C53431B0CAAA30B2B" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
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<subSubSection id="F31BB7B06DBB1E444E8130BB8382BAA0" pageId="30" pageNumber="31" type="multiple">
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<paragraph id="AB30364F9ED8A3DE0A742BDCB42C7CE9" pageId="30" pageNumber="31">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="118B8499E7673535198041FFB8F81607" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="59F93529E2FDD77457C3FB83A102AFF6" pageId="30" pageNumber="31">
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<taxonomicName id="520D5144F7A9E2B30173F1B682A5AFE5" ID-CoL="3RM67" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes Guignot, 1955</taxonomicName>
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Figs 33-34, 230-232, 391, 530
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</paragraph>
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</subSubSection>
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<subSubSection id="C4477F1BE0326BD4DA02BC066D776966" pageId="30" pageNumber="31" type="reference_group">
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<paragraph id="AA7452CB3E5901F6C21A0A8527EC63B9" pageId="30" pageNumber="31">
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<taxonomicName id="7D5552A1DC1AB5FA3AFC0F848FB2FB71" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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<bibRefCitation id="3A3F244523FEA9022F0F13D744AE1E10" author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="257" pageNumber="258" pagination="139 - 144" title="Nouveaux dytiscides de Madagascar (Coleopteres)." volume="7" year="1955 f">Guignot 1955f</bibRefCitation>
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: 141 (original description, faunistics);
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<bibRefCitation id="14C24E1E578D73EDCD10EF101ACC53B1" author="Rocchi, S" journalOrPublisher="Frustula Entomologica (n. s.)" pageId="262" pageNumber="263" pagination="71 - 89" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera)." volume="14" year="1991">Rocchi 1991</bibRefCitation>
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: 86 (faunistics, list);
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<bibRefCitation id="9FAC82C49992FA183D7AD91773E1E259" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 242 (catalogue, faunistics);
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<bibRefCitation id="E3DE200F8A04F4AAC96471C84E6E27F1" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 210 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="2A3D11EDC21B493FE0394E14C8CF8707" pageId="30" pageNumber="31" type="type locality">
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<paragraph id="6F56537113D5712CD4FD53C75B095A6B" pageId="30" pageNumber="31">Type locality.</paragraph>
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<paragraph id="35870287FD556AABB8A7CA7A6AF53D37" pageId="30" pageNumber="31">Madagascar, Tampolo.</paragraph>
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</subSubSection>
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<subSubSection id="82C81B6DED179901BE8FD6C177BCAC51" pageId="30" pageNumber="31" type="type material">
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<paragraph id="16FF30D033ED389AD7F2548759A04658" pageId="30" pageNumber="31">Type material, studied</paragraph>
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<paragraph id="27EDB5AAC1E04C16738245A253242F16" pageId="30" pageNumber="31">(2 exs.). Holotype: male: "Madagascar Tampolo VIII. 1949 / male symbol / Type" (MNHN). - Allotype (= Paratype), female: "Madagascar Tampolo VIII. 1949 / female symbol / Allotype" (1 ex. MNHN; habitus in Fig. 391).</paragraph>
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</subSubSection>
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<subSubSection id="2FB0ADCA0AD827448E164997F0825B50" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="materials_examined">
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<paragraph id="9AD4DA17B01DFD228167D397F273CA1C" pageId="30" pageNumber="31">Additional material studied</paragraph>
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<paragraph id="62FAB582CBAD15BD3DDC13ACC152F0A5" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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(116 exs.): Madagascar: "Andasibe 11.12. 2004 Lat -18.943 Lon 48.4063, Balke & Monaghan / DNA Voucher BMNH <670655> MSL008:B01 /
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<taxonomicName id="7248925C8205DBE4B5CC6BFA4261D7AB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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Bergsten det." (1 ex. NHRS); same data but
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<normalizedToken id="D0E3E5C64940E92E66BC56B63BF10D66" originalValue="“BMNH<670654>”">"BMNH<670654>"</normalizedToken>
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(1 ex. NHRS); "Toam, Ambatondrazaka, Zahamane, Zahamena NP river, P60BI04: N: E: m, 29.12. 2006 leg. Isambert et al / DNA Voucher BMNH <830741> MSL399:F2 /
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<taxonomicName id="2660F587492DD7DCD6168B6B4797E25B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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Bergsten det." (1 ex. NHRS); same data but "BMNH <830743> MSL399:F4" (1 ex. NHRS); "Toam, Ambatondrazaka, Zahamane, Zahamena NP stream, P60BI15:
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<pageBreakToken id="6F7D2D81783A20AE7832DDC9F4D9F1E2" pageId="31" pageNumber="32" start="start">N</pageBreakToken>
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: -
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<geoCoordinate id="14BBCC4B6FB5F5AD562B79BAA85CEDCA" direction="east" orientation="longitude" precision="555" value="17.52">17.52 E</geoCoordinate>
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: 48.721: 1075m / m, 29.12. 2006 leg. Isambert et al / DNA Voucher BMNH <830737> MSL399:E10 /
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<taxonomicName id="F45E9AF32F90517B6038794BBE84745F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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Bergsten det." (1 ex. NHRS); same data but "Analamaintsoa 3rd stream before Camp 1, stream of pools almost dry P60BI08, 30.12. 2006
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<geoCoordinate id="5C666A48A3BC6C68333C8CBF7E538290" direction="north" orientation="latitude" precision="1" value="17.505">N-17,50500</geoCoordinate>
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,
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<geoCoordinate id="CAEEB923458B9AB5603F7A92963AC188" direction="east" orientation="longitude" precision="1" value="48.7245">E48,72450</geoCoordinate>
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, 1054 m Isambert et al (1 ex. NHRS); same data but "BMNH <830748> MSL399:F)/P60BI12: N: -
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<geoCoordinate id="44212B65562BB5F99C3980573B959FE2" direction="east" orientation="longitude" precision="55" value="17.517">17.517 E</geoCoordinate>
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: 48.72: 1075 m, 31.12. 2006" (1 ex. NHRS); same data but "Zahamena NP 1st stream btw. Camp 1 and 2, stream of pools, 31.12. 2006,
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<geoCoordinate id="E2B2AEBB96A01000811D11137A3CD722" direction="north" orientation="latitude" precision="1" value="17.51733">N-17,51733</geoCoordinate>
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, 48,72067, 1075 m" (85 exx. NHRS, 2 exs. MZH); same data but "Analamaintsoa Forest 4th stream btw Camp 1 and 2, P6OBI15, 31.12. 2006,
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<geoCoordinate id="DF558C83F2F855BAA49A00F773B1203A" direction="north" orientation="latitude" precision="1" value="17.5205">N-17,52050</geoCoordinate>
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,
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<geoCoordinate id="B671BD05012237EAB5EC45CDEEC8A317" direction="east" orientation="longitude" precision="1" value="48.721336">E48,721337</geoCoordinate>
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, 1075 m" (14 exx. NHRS); "IF Anadiana: Sahamalaotra Ranomafana NP: small stream P27MD31, N -
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<geoCoordinate id="5963C26630A0E2D005D201A75EA19F9A" direction="east" orientation="longitude" precision="5" value="21.2359">21.2359 E</geoCoordinate>
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: 47.3963, alt. 1123 m, 6.12.2004 leg. Balke et al / BMNH(E) <794196> DNA Voucher" (1 ex. NHRS); same but" BMNH(E) <794198>" (1 ex. NHRS); same but" BMNH(E) <794197>" (1 ex. NHRS); Mahajanga melaky, Tsingy de Bemaraha NP,
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<geoCoordinate id="7856604B880DE9BB2589CCC0DDF49513" direction="south" orientation="latitude" precision="1" value="-19.03419">S19.03419</geoCoordinate>
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,
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<geoCoordinate id="31ACC5310D788B9F89E67388DBBB6D24" direction="east" orientation="longitude" precision="1" value="44.77499">E044.77499</geoCoordinate>
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, 41 m.a.o., 15.12. 2009, Water net, field, Bergsten et al. / 000000464 NHRS-JLKB" (1 ex. NHRS); same but "
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<geoCoordinate id="B403FA7A368A75905F7E9DED761E6F1C" direction="south" orientation="latitude" precision="1" value="-19.03572">S19.03572</geoCoordinate>
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,
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<geoCoordinate id="9FC3B02EFD8E6839FCE614D9C0CA00CA" direction="east" orientation="longitude" precision="1" value="44.77507">E044.77507</geoCoordinate>
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, 66 m.a.o., 15.12. 2009 / 000000467 NHRS/JLKB" (1 ex. NHRS); same but "
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<geoCoordinate id="354CB96D78426546A0009ADA738679BA" direction="south" orientation="latitude" precision="1" value="-18.75643">S18.75643</geoCoordinate>
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,
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<geoCoordinate id="9CACCE980D8149C163688D09BE2A8DF4" direction="east" orientation="longitude" precision="1" value="44.71398">E044.71398</geoCoordinate>
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, 119 m.a.o., 17.12. 2009 / 000000463 NHRS/JLKB" (1 ex. NHRS); "Ampasimpotsy Moramanga Antsabe 11.12. 2004
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<geoCoordinate id="D3F26A31A6BEFA6498E7C11D3EE61520" direction="north" orientation="latitude" precision="1" value="18.943">N-18,94300</geoCoordinate>
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,
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<geoCoordinate id="1112D5677A5886DBF60355BA57DC332B" direction="east" orientation="longitude" precision="1" value="48.4063">E48,40630</geoCoordinate>
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, 979 m, Balke et al" (2 exx. NHRS).
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</paragraph>
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</subSubSection>
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<subSubSection id="2A16F886DFA637B0F14AC7331A3610F9" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="49740DE8B3F9A481820A328C0961B704" pageId="31" pageNumber="32">Diagnosis.</paragraph>
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<paragraph id="8E340B9C548ED21BB699A07464A33A9F" pageId="31" pageNumber="32">
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<taxonomicName id="A6416980BA150E7570A83A170FD04621" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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forms together with
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<taxonomicName id="1B5EE80B9830C62675C8B3FE4E4DFC3C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
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,
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<taxonomicName id="6A0C04C547FB752E408C43895F8EFE57" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus tigrinus" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tigrinus">Laccophilus tigrinus</taxonomicName>
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,
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<taxonomicName id="C987981E59D87AE1EECC31480BBE8542" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pseustes" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pseustes">Laccophilus pseustes</taxonomicName>
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and
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<taxonomicName id="E57AE2B97A84787B3F006B2B08115E79" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
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a distinct group characterized by similar colour pattern of body and male genitalia.
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<taxonomicName id="9736F8702BD229E268E07913A6BB995A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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,
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<taxonomicName id="496C9CDDDB0FB54DFAF1016A671131B3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
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and
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<taxonomicName id="554DF883942348AFEC0B24D684EB6EA4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluadi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluadi">Laccophilus alluadi</taxonomicName>
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are smaller than the other species in the group. The three species are separated by small differences in shape of penis; see diagnosis of
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<taxonomicName id="E826CD81D0621C074FCF3801E632ECD0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
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on p. 41 and
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<taxonomicName id="407312E3200E2F725D18A273059E8AED" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
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on p. 42.
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</paragraph>
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</subSubSection>
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<subSubSection id="515B1D7FAD975C21B557E97CB9B4AF99" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="description">
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<paragraph id="7064F0839BEA6983EC6B30617FA1444A" pageId="31" pageNumber="32">Description.</paragraph>
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<paragraph id="92E3965E5A6407FAA9C54E116511CC08" pageId="31" pageNumber="32">Body length 3.3-3.9 mm, width 1.8-2.2 mm. Dorsal, colour pattern distinct and uniform (Fig. 391).</paragraph>
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<paragraph id="1BAABD69D9828F171BE92E3CF6F19F90" pageId="31" pageNumber="32">Head: Pale ferrugineous. At eyes with fine, irregularly distributed punctures. Rather shiny, although microsculptured. Reticulation double; coarse meshes only slightly stronger developed than fine meshes. Coarse meshes, when discernible, contain 2-3 fine meshes. In part, mesh categories cannot be distinguished.</paragraph>
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<paragraph id="349724E080C9034E51B20A9F04ACE7BC" pageId="31" pageNumber="32">Pronotum: Pale ferrugineous. Impunctate, except anteriorly, with scattered, fine punctures. Rather shiny, although finely microsculptured. Reticulation double. Coarse meshes only slightly more strongly developed than fine meshes; contain when discernible 3-4 fine meshes.</paragraph>
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<paragraph id="C9ED220635CE270C3AD1B26495021A7B" pageId="31" pageNumber="32">
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Elytra: Pale ferrugineous, with distinct, blackish ferrugineous to dark ferrugineous, longitudinal markings (Fig. 391). In a few specimens dark, longitudinal markings close to humeral region strongly reduced as in
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<taxonomicName id="4B3BF6FB6E0D44C30C61A5B2EC23DB30" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
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. Posteriorly with rather fine, scattered punctation. Submat, distinctly and densely microsculptured. In part, reticulation double. Two kinds of reticulation clearly visible at scutellar region; posteriorly and laterally meshes of reticulation approximately one kind.
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</paragraph>
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<paragraph id="E7D53E7429869C6ADEC2D1ED1804BA52" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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Ventral aspect: Metathorax and -coxal plates dark ferrugineous to ferrugineous; otherwise pale ferrugineous. Very fine, sparse punctures discernible on metacoxal plates and abdomen. Slightly mat due to very fine and hardly discernible microsculpture.
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<pageBreakToken id="93832C41969D643C3A9E5D21A664B5CC" pageId="32" pageNumber="33" start="start">Prosternal</pageBreakToken>
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process rather slender, apex pointed but not strongly extended backwards. Abdomen with fine, sparse, curved striae. Apical ventrite not distinctly modified (Fig. 33).
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</paragraph>
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<paragraph id="D75B2A546D86FB35DDD174BF0402F026" pageId="32" pageNumber="33">Legs: Pro- and mesotarsus slender, somewhat extended; provided with suckers.</paragraph>
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<paragraph id="28815D239E529A6668B1834436FEBD74" pageId="32" pageNumber="33">Male genitalia: Penis in lateral aspect medially, straight for a long distance, apex slightly bent; in dorsal aspect, penis from middle, strongly bent right (Figs 230-232).</paragraph>
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<paragraph id="260E899D591767F6B7467ED9170FC6C5" pageId="32" pageNumber="33">Female: Pro- and mesotarsus slender. Apex of apical ventrite more angular than in male (Fig. 34).</paragraph>
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</subSubSection>
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<subSubSection id="5A96EDA18A6B3CCEBA851A8B7246BA2E" pageId="32" pageNumber="33" type="distribution">
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<paragraph id="B84C672DA7AB70364A98E8AB28264B3F" pageId="32" pageNumber="33">Distribution.</paragraph>
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<paragraph id="9CB4FF3B68240E725F3B236F780A88FA" pageId="32" pageNumber="33">Madagascar (Fig. 530).</paragraph>
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</subSubSection>
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<subSubSection id="68EB2D5748001F435B3010B7FFEACE2F" pageId="32" pageNumber="33" type="collecting circumstances">
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<paragraph id="FB8468E7F213E16C57C1361B286CD159" pageId="32" pageNumber="33">Collecting circumstances.</paragraph>
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<paragraph id="98F6DD2AED78201E9D60D216FB4698E5" pageId="32" pageNumber="33">
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Label data indicate that
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<taxonomicName id="0FA154E22C00075EB64D4F79A10158F4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
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has been sampled in various sized, running waters as streams and rivers.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |