262 lines
19 KiB
XML
262 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.14.2520" ID-PMC="PMC3492924" ID-Pensoft-Pub="1314-2003-14-23" ID-Pensoft-UUID="FFD6B463B03FBB5EFFD4FFC3FFC1FFC4" ID-PubMed="23170071" ID-Zenodo-Dep="576131" ModsDocID="1314-2003-14-23" checkinTime="1451252000297" checkinUser="pensoft" docAuthor="Robinson, Harold & Funk, Vicki A." docDate="2012" docId="D9CF6FABB2FC5C5A93A6BCD9327C3559" docLanguage="en" docName="PhytoKeys 14: 23-41" docOrigin="PhytoKeys 14" docPubDate="2012-07-30" docSource="http://dx.doi.org/10.3897/phytokeys.14.2520" docTitle="Cuatrecasanthus flexipappus (Gleason) H. Rob., Revista Colomb. Ci. Exact. 17 (65): 210. 1989." docType="treatment" docVersion="3" id="FFD6B463B03FBB5EFFD4FFC3FFC1FFC4" lastPageNumber="28" masterDocId="FFD6B463B03FBB5EFFD4FFC3FFC1FFC4" masterDocTitle="Cuatrecasanthus (Vernonieae, Compositae): A revision of a north-central Andean genus" masterLastPageNumber="41" masterPageNumber="23" pageNumber="25" updateTime="1668139705674" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cuatrecasanthus (Vernonieae, Compositae): A revision of a north-central Andean genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Robinson, Harold</mods:namePart>
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<mods:affiliation>Department of Botany, MRC 166, National Museum of Natural History, P. O. Box 37012, Smithsonian Institution, Washington, DC. 20013 - 7012</mods:affiliation>
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<mods:nameIdentifier type="email">robinsoh@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Funk, Vicki A.</mods:namePart>
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<mods:affiliation>Department of Botany, MRC 166, National Museum of Natural History, P. O. Box 37012, Smithsonian Institution, Washington, DC. 20013 - 7012</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2012-07-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>14</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>41</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.14.2520</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.14.2520</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-14-23</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFD6B463B03FBB5EFFD4FFC3FFC1FFC4</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576131</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="182229322" LSID="urn:lsid:plazi:treatment:D9CF6FABB2FC5C5A93A6BCD9327C3559" httpUri="http://treatment.plazi.org/id/D9CF6FABB2FC5C5A93A6BCD9327C3559" lastPageId="5" lastPageNumber="28" pageId="2" pageNumber="25">
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<subSubSection pageId="2" pageNumber="25" type="nomenclature">
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<paragraph pageId="2" pageNumber="25">
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1.
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<taxonomicName LSID="D9CF6FAB-B2FC-5C5A-93A6-BCD9327C3559" authority="(Gleason) H. Rob., Revista Colomb. Ci. Exact. 17 (65): 210. 1989." authorityName="(Gleason) H. Rob., Revista Colomb. Ci. Exact. 17 (65): 210. 1989." class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Cuatrecasanthus flexipappus" order="Asterales" pageId="2" pageNumber="25" phylum="Tracheophyta" rank="species" species="flexipappus">Cuatrecasanthus flexipappus (Gleason) H. Rob., Revista Colomb. Ci. Exact. 17 (65): 210. 1989.</taxonomicName>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photographs of Cuatrecasanthus types: A Cuatrecasanthus flexipappus, holotype (NY) B Cuatrecasanthus giannasii, holotype (S)." httpUri="https://binary.pensoft.net/fig/10122" pageId="2" pageNumber="25">Figs 5A</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution map of Cuatrecasanthus species." httpUri="https://binary.pensoft.net/fig/10127" pageId="2" pageNumber="25">10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="25" type="type - based on">
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<paragraph pageId="2" pageNumber="25">
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Type: Based on
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<taxonomicName authorityName="Gleason" authorityYear="1925" class="Insecta" family="Philopteridae" genus="Vernonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Phthiraptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="flexipappa">
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<emphasis italics="true" pageId="2" pageNumber="25">Vernonia flexipappa</emphasis>
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</taxonomicName>
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Gleason
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</paragraph>
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<paragraph pageId="2" pageNumber="25">
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<taxonomicName authorityName="Gleason" authorityYear="1925" class="Insecta" family="Philopteridae" genus="Vernonia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Phthiraptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="flexipappa">
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<emphasis italics="true" pageId="2" pageNumber="25">Vernonia flexipappa</emphasis>
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</taxonomicName>
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Gleason, Bull. Torrey Bot. Club 52(5): 186.1925.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="25" type="type">
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<paragraph pageId="2" pageNumber="25">Type:</paragraph>
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<paragraph pageId="2" pageNumber="25">
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Ecuador. Loja: sin. loc.,
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<emphasis italics="true" pageId="2" pageNumber="25">
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E.
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<normalizedToken originalValue="André">Andre</normalizedToken>
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2250
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</emphasis>
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(holotype: NY, image US!; isotype: K).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25" type="description">
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<paragraph pageId="2" pageNumber="25">Description.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25">
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Shrubs or small trees, 1.0-3.0 m tall;
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<emphasis italics="true" pageId="2" pageNumber="25">stems</emphasis>
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densely pilose with dark brown trichomes, becoming glabrous with age.
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<emphasis italics="true" pageId="2" pageNumber="25">Leaves</emphasis>
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with petioles 0.5-1.2 cm long;
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<emphasis italics="true" pageId="2" pageNumber="25">blades</emphasis>
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narrowly to broadly elliptical, mostly 3-9 cm long, 1-3 cm wide, narrowly acuminate at base and apex, margin narrowly but strongly recurved, without evident teeth or with in-turned teeth, adaxial surface dark green, glabrous or with minute appressed pubescence, secondary and tertiary veins insculpate, abaxial surface pale greenish covered with mostly appressed, stiff, brownish trichomes (rarely straw colored) intermixed with less evident whitish prostrate myceliiform branching trichomes, midvein with dense antrorse pubescence mostly on sides;
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<emphasis italics="true" pageId="2" pageNumber="25">secondary veins</emphasis>
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ca. 5 pairs, spreading from midvein at ca. 45° angles, strongly curved.
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<emphasis italics="true" pageId="2" pageNumber="25">Inflorescence</emphasis>
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scarcely exceeding vegetative leaves;
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<emphasis italics="true" pageId="2" pageNumber="25">branches</emphasis>
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densely pilosulous or hirtellous.
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<emphasis italics="true" pageId="2" pageNumber="25">Heads</emphasis>
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sessile in clusters of 3-7, ca. 10-11 mm tall
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<normalizedToken originalValue="×">x</normalizedToken>
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2 mm wide;
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<emphasis italics="true" pageId="2" pageNumber="25">involucres</emphasis>
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cylindrical to fusiform; bracts mostly deciduous, ca. 15 in ca. 5 series, 1.0-5.5 mm long, ca. 1.2 mm wide, apices short-acute, ovate to narrowly elliptical, yellowish or with reddish median stripe, puberulous to nearly glabrous outside.
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<emphasis italics="true" pageId="2" pageNumber="25">Florets</emphasis>
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with corollas white to bluish white or lavender, ca. 5.5 mm long, with glandular dots on basal tube and tips of lobes, few small trichomes
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<pageBreakToken pageId="3" pageNumber="26" start="start">on</pageBreakToken>
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lobe tips, tubes ca. 2 mm long, lobes ca. 4 mm long, with some non-glandular trichomes;
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<emphasis italics="true" pageId="3" pageNumber="26">anther thecae</emphasis>
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ca. 2 mm long.
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<emphasis italics="true" pageId="3" pageNumber="26">Achenes</emphasis>
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2.0-2.5 mm long;
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<emphasis italics="true" pageId="3" pageNumber="26">pappus</emphasis>
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white, of ca. 50 bristles mostly ca. 6 mm long, not or scarcely broadened toward tips.
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<emphasis italics="true" pageId="3" pageNumber="26">Pollen</emphasis>
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grains 37-47
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<normalizedToken originalValue="µm">µm</normalizedToken>
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in diam.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/10122" pageId="3" pageNumber="26" start="Figure 5" startId="F5">
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<paragraph pageId="3" pageNumber="26">
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<emphasis bold="true" pageId="3" pageNumber="26">Figure 5.</emphasis>
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Photographs of
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<taxonomicName authorityName="H. Rob., Revista Acad. Colomb. Ci. Exact. 17 (65): 209" authorityYear="1989" class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="26" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="3" pageNumber="26">Cuatrecasanthus</emphasis>
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</taxonomicName>
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types:
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<emphasis bold="true" pageId="3" pageNumber="26">A</emphasis>
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="26" phylum="Tracheophyta" rank="species" species="flexipappus">
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<emphasis italics="true" pageId="3" pageNumber="26">Cuatrecasanthus flexipappus</emphasis>
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</taxonomicName>
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, holotype (NY)
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<emphasis bold="true" pageId="3" pageNumber="26">B</emphasis>
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<taxonomicName authorityName="H. Rob. & V. A. Funk" authorityYear="2012" baseAuthorityName="H. Rob. & V. A. Funk" baseAuthorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="3" pageNumber="26" phylum="Tracheophyta" rank="species" species="giannasii">
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<emphasis italics="true" pageId="3" pageNumber="26">Cuatrecasanthus giannasii</emphasis>
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</taxonomicName>
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, holotype (S).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="28" pageId="3" pageNumber="26" type="additional specimens examined">
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<paragraph pageId="3" pageNumber="26">Additional specimens examined.</paragraph>
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<paragraph lastPageId="5" lastPageNumber="28" pageId="3" pageNumber="26">
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Ecuador.
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<emphasis bold="true" pageId="3" pageNumber="26">Loja:</emphasis>
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along road between Loja and Zamora, ca. km 11 [
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<geoCoordinate degrees="03" direction="south" minutes="59" orientation="latitude" precision="15" seconds="0" value="-3.9833333">03°59'0"S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="08" orientation="longitude" precision="15" seconds="16" value="-79.13778">79°08'16"W</geoCoordinate>
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, estimated], 2600 m, 2 August 1978,
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<emphasis italics="true" pageId="3" pageNumber="26">Zarucchi & Andrade 2304</emphasis>
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(US); Carretera Loja-Zamora, km 13, 2500 m [
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<geoCoordinate degrees="03" direction="south" minutes="59" orientation="latitude" precision="15" seconds="00" value="-3.9833333">03°59'00"S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="07" orientation="longitude" precision="15" seconds="00" value="-79.11667">79°07'00"W</geoCoordinate>
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, estimated], 16 August 1983,
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<emphasis italics="true" pageId="3" pageNumber="26">Jaramillo & Winnerskojold 5812</emphasis>
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(AAU); Loja-Zamora road, ca. km 15,
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<geoCoordinate degrees="03" direction="south" minutes="58" orientation="latitude" precision="925" value="-3.9666667">03°58'S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="08" orientation="longitude" precision="925" value="-79.13333">79°08'W</geoCoordinate>
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, 2400-2700 m, 22-23 April 1984,
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<emphasis italics="true" pageId="3" pageNumber="26">Madsen 74081</emphasis>
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(AAU, QCA, US); In the
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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of "El Tiro," located at northern terminus of Podocarpus National Park, 500 m from the Loja-Zamora highway,
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<geoCoordinate degrees="03" direction="south" minutes="59" orientation="latitude" precision="925" value="-3.9833333">03°59'S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="08" orientation="longitude" precision="925" value="-79.13333">79°08'W</geoCoordinate>
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, 2940-2970 m, 14 April 1992,
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<emphasis italics="true" pageId="3" pageNumber="26">Keating 143</emphasis>
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(US); In the
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<pageBreakToken pageId="4" pageNumber="27" start="start">
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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</pageBreakToken>
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and shrub
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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above the Refugio de Cajanuma (Centro de
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<normalizedToken originalValue="Información">Informacion</normalizedToken>
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), Podocarpus National Park,
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<geoCoordinate degrees="04" direction="south" minutes="07" orientation="latitude" precision="15" seconds="00" value="-4.116667">04°07'00"S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="09" orientation="longitude" precision="15" seconds="30" value="-79.15833">79°09'30"W</geoCoordinate>
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, 2800 m, 31 July 1993,
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<emphasis italics="true" pageId="4" pageNumber="27">Keating 409</emphasis>
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(US).
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<emphasis bold="true" pageId="4" pageNumber="27">Zamora-Chinchipe:</emphasis>
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14.8 km from transit control out of Loja on ro
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<pageBreakToken pageId="5" pageNumber="28" start="start">ad</pageBreakToken>
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to Zamora [
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<geoCoordinate degrees="03" direction="south" minutes="59" orientation="latitude" precision="15" seconds="10" value="-3.9861112">03°59'10"S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="08" orientation="longitude" precision="15" seconds="02" value="-79.13389">79°08'02"W</geoCoordinate>
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, estimated], 2500 m, 8 July 1983,
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<emphasis italics="true" pageId="5" pageNumber="28">Keeley & Keeley 4104, 4105, 4106, 4107, 4108, 4109, 4110, 4111, 4115</emphasis>
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(K, US);
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<emphasis italics="true" pageId="5" pageNumber="28">Keeley & Keeley 4112, 4114</emphasis>
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(US);Zamora, carretera Loja-Zamora,
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<normalizedToken originalValue="Estación">Estacion</normalizedToken>
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<normalizedToken originalValue="Científica">Cientifica</normalizedToken>
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San Francisco, sendero hacia las antenas. Colecciones cerca del Francisco 4, en Transecto 2,
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<geoCoordinate degrees="03" direction="south" minutes="58" orientation="latitude" precision="925" value="-3.9666667">03°58'S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="04" orientation="longitude" precision="925" value="-79.066666">79°04'W</geoCoordinate>
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, 3000 m, 29 April 2000;
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<emphasis italics="true" pageId="5" pageNumber="28">Freire Fierro 3121</emphasis>
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(MO, US).
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</paragraph>
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<paragraph pageId="5" pageNumber="28">
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Peru.
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<emphasis bold="true" pageId="5" pageNumber="28">Cajamarca:</emphasis>
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Prov. Jaen; E slope of Paso de Huascarai, head of Quebrada Granadillas, 15 km SE of Huancabamba,
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<geoCoordinate degrees="05" direction="south" minutes="22" orientation="latitude" precision="925" value="-5.366667">05°22'S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="20" orientation="longitude" precision="925" value="-79.333336">79°20'W</geoCoordinate>
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, 3000 m, 10 June 1947,
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<emphasis italics="true" pageId="5" pageNumber="28">Fosberg 27852</emphasis>
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(US).
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<emphasis bold="true" pageId="5" pageNumber="28">Piura:</emphasis>
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Prov. Huancabamba; Los Llanos to Chorro Blanco (Sapalache - Chiguelas), 2650 m,
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<geoCoordinate degrees="5" direction="south" minutes="08.2" orientation="latitude" precision="92" value="-5.136667">5°08.2'S</geoCoordinate>
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,
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<geoCoordinate degrees="79" direction="west" minutes="24.6" orientation="longitude" precision="92" value="-79.41">79°24.6'W</geoCoordinate>
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, 19 Oct. 2001,
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<emphasis italics="true" pageId="5" pageNumber="28">
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<normalizedToken originalValue="Sagasteguí">Sagastegui</normalizedToken>
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, Dillon, Leiva, & Zapata 16781
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</emphasis>
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(F, HUT).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="28" type="habitat">
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<paragraph pageId="5" pageNumber="28">Habitat.</paragraph>
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<paragraph pageId="5" pageNumber="28">
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Roadside, burned over cloud forest on steep south-facing slope; shrub
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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at 2400-3000 m in elevation.
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</paragraph>
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<paragraph pageId="5" pageNumber="28">
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The species is the most commonly collected member of the genus but apparently is sympatric with both
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<taxonomicName authorityName="H. Rob. & V. A. Funk" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="28" phylum="Tracheophyta" rank="species" species="kingii">
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<emphasis italics="true" pageId="5" pageNumber="28">Cuatrecasanthus kingii</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="H. Rob. & V. A. Funk" authorityYear="2012" class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="28" phylum="Tracheophyta" rank="species" species="lanceolatus">
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<emphasis italics="true" pageId="5" pageNumber="28">Cuatrecasanthus lanceolatus</emphasis>
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</taxonomicName>
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in the area near the border between Loja and Zamora/Chinchipe. The species is very closely related to the northern Peruvian
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<taxonomicName authorityName="H. Rob" authorityYear="1989" baseAuthorityName="Hieron." class="Magnoliopsida" family="Asteraceae" genus="Cuatrecasanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="5" pageNumber="28" phylum="Tracheophyta" rank="species" species="jelskii">
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<emphasis italics="true" pageId="5" pageNumber="28">Cuatrecasanthus jelskii</emphasis>
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</taxonomicName>
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(Hieron.) H. Rob. The latter differs most obviously by the densely hirsute abaxial surface of the midvein of the leaves and erect rather than appressed trichomes of the abaxial blade surface. The adaxial leaf surface of the latter also has less strongly insculpate veins.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="28" type="preliminary conservation status">
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<paragraph pageId="5" pageNumber="28">Preliminary conservation status.</paragraph>
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<paragraph pageId="5" pageNumber="28">Data Deficient</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |