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<document id="2D8DC1D598F34C7E58FB262959A48DDB" ID-CLB-Dataset="118647" ID-DOI="10.1111/zoj.12040" ID-GBIF-Dataset="dd75c7a5-e473-4119-8db8-ca35a11ca128" ID-ISSN="0024-4082" ID-Zenodo-Dep="5282985" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630045613184" checkinUser="marcus" docAuthor="Bolzern, Angelo, Burckhardt, Daniel &amp; Hänggi, Ambros" docDate="2013" docId="BD701413E264B61854D5FB15C0E91072" docLanguage="en" docName="zoj12040.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 168 (4)" docSource="http://dx.doi.org/10.1111/zoj.12040" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Tegenaria podoprygorai Revision" docType="treatment" docVersion="10" lastPageNumber="813" masterDocId="41496C6BE23DB6425411FF94C115171C" masterDocTitle="Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria-Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data" masterLastPageNumber="848" masterPageNumber="723" pageNumber="812" updateTime="1699146108430" updateUser="ExternalLinkService">
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<mods:title id="0D3E6AA9254A1449CEF1900A72E19B2B">Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria-Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data</mods:title>
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<mods:namePart id="DC9DDAE2CF98B4881EEF03773705FA71">Bolzern, Angelo</mods:namePart>
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<mods:namePart id="5F7E35ACA78BA3432FB2DC2B85606621">Burckhardt, Daniel</mods:namePart>
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<mods:namePart id="3437099834C3E3FBB5592731AB78915C">Hänggi, Ambros</mods:namePart>
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<mods:title id="877EA94DA3C322DB63EF296D9E563C5F">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="AE686DACF182CB29417A369A03EF957E">2013</mods:date>
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<mods:title id="A771C2C3790A52C46914E906853B61B5">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="0A77DDFF45A0B42E0649A46AE06D8EA5">2013-07-26</mods:number>
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<mods:number id="E3A5BF8CD930BC6602A80FC0295EE469">168</mods:number>
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<treatment id="BD701413E264B61854D5FB15C0E91072" ID-DOI="http://doi.org/10.5281/zenodo.6983379" ID-GBIF-Taxon="198975580" ID-Zenodo-Dep="6983379" LSID="urn:lsid:plazi:treatment:BD701413E264B61854D5FB15C0E91072" httpUri="http://treatment.plazi.org/id/BD701413E264B61854D5FB15C0E91072" lastPageId="90" lastPageNumber="813" pageId="89" pageNumber="812">
<subSubSection id="7DC3F68EE264B61B54D5FB15C30913A9" pageId="89" pageNumber="812" type="nomenclature">
<paragraph id="3566A505E264B61B54D5FB15C30913A9" blockId="89.[196,747,1152,1205]" pageId="89" pageNumber="812">
<heading id="6E2E1269E264B61B54D5FB15C30913A9" centered="true" fontSize="9" level="2" pageId="89" pageNumber="812" reason="4">
<taxonomicName id="F2D9DE86E264B61B54D5FB15C3FE138A" ID-CoL="7BMS8" authority="(KOVBLYUK, 2006)" authorityName="Revision" baseAuthorityName="Kovblyuk" baseAuthorityYear="2006" box="[196,747,1152,1174]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="844" phylum="Arthropoda" rank="species" species="podoprygorai">
<emphasis id="07AD7917E264B61B54D5FB15C31E1389" bold="true" box="[196,523,1153,1174]" italics="true" pageId="89" pageNumber="812">TEGENARIA PODOPRYGORAI</emphasis>
(
<bibRefCitation id="5148D8F4E264B61B560CFB14C3F6138A" author="Kovblyuk MM" box="[541,739,1152,1174]" pageId="89" pageNumber="812" pagination="23 - 37" refId="ref85145" refString="Kovblyuk MM. 2006. Malthonica podoprygorai sp. n. from the Crimea (Aranei: Agelenidae). Arthropoda Selecta 15: 23 - 37." type="journal article" year="2006">KOVBLYUK, 2006</bibRefCitation>
)
</taxonomicName>
<emphasis id="07AD7917E264B61B5582FB30C30913A9" bold="true" box="[403,540,1184,1205]" pageId="89" pageNumber="812">COMB. NOV.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7DC3F68EE264B61B54B5FB53C00513E7" pageId="89" pageNumber="812" type="reference_group">
<paragraph id="3566A505E264B61B54B5FB53C00513E7" blockId="89.[164,778,1223,1306]" pageId="89" pageNumber="812">
<treatmentCitationGroup id="15C9822BE264B61B54B5FB53C00513E7" pageId="89" pageNumber="812">
<treatmentCitation id="B4788314E264B61B54B5FB53C21313C1" author="Kovblyuk MM" box="[164,774,1223,1245]" page="26" pageId="89" pageNumber="812" year="2006">
<taxonomicName id="F2D9DE86E264B61B54B5FB53C21313C1" authority="Kovblyuk, 2006: 24 - 26" authorityName="Kovblyuk" authorityPageNumber="24 - 26" authorityYear="2006" baseAuthorityName="Kovblyuk" baseAuthorityYear="2006" box="[164,774,1223,1245]" class="Arachnida" family="Agelenidae" genus="Malthonica" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="812" phylum="Arthropoda" rank="species" species="podoprygorai">
<emphasis id="07AD7917E264B61B54B5FB53C0C513C0" box="[164,464,1223,1244]" italics="true" pageId="89" pageNumber="812">Malthonica podoprygorai</emphasis>
<bibRefCitation id="5148D8F4E264B61B55F4FB53C21313C1" author="Kovblyuk MM" box="[485,774,1223,1245]" pageId="89" pageNumber="812" pagination="23 - 37" refId="ref85145" refString="Kovblyuk MM. 2006. Malthonica podoprygorai sp. n. from the Crimea (Aranei: Agelenidae). Arthropoda Selecta 15: 23 - 37." type="journal article" year="2006">Kovblyuk, 2006: 24–26</bibRefCitation>
</taxonomicName>
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, figs 1–10.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7DC3F68EE264B61B54ADFA91C0BD1186" pageId="89" pageNumber="812" type="materials_examined">
<paragraph id="3566A505E264B61B54ADFA91C0DD1206" blockId="89.[164,778,1223,1306]" box="[188,456,1284,1306]" pageId="89" pageNumber="812">No material examined.</paragraph>
<paragraph id="3566A505E264B61B54B5FADCC0301241" blockId="89.[164,779,1352,1434]" box="[164,293,1352,1373]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B54B5FADCC0301241" box="[164,293,1352,1373]" italics="true" pageId="89" pageNumber="812">Description</emphasis>
</paragraph>
<paragraph id="3566A505E264B61B54B5FAF2C1E71286" blockId="89.[164,779,1352,1434]" pageId="89" pageNumber="812">
A detailed description was provided by
<bibRefCitation id="5148D8F4E264B61B568FFAF2C1FA1286" author="Kovblyuk MM" pageId="89" pageNumber="812" pagination="23 - 37" refId="ref85145" refString="Kovblyuk MM. 2006. Malthonica podoprygorai sp. n. from the Crimea (Aranei: Agelenidae). Arthropoda Selecta 15: 23 - 37." type="journal article" year="2006">Kovblyuk (2006)</bibRefCitation>
.
</paragraph>
<paragraph id="3566A505E264B61B54B5FA53C03A12C0" blockId="89.[164,442,1479,1532]" box="[164,303,1479,1500]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B54B5FA53C03A12C0" box="[164,303,1479,1500]" italics="true" pageId="89" pageNumber="812">Distribution</emphasis>
</paragraph>
<paragraph id="3566A505E264B61B54B5FA72C0AF12E0" blockId="89.[164,442,1479,1532]" box="[164,442,1510,1532]" pageId="89" pageNumber="812">
Reported from
<collectingCountry id="4DCEE595E264B61B5545FA72C0A312E0" box="[340,438,1510,1532]" name="Ukraine" pageId="89" pageNumber="812">Ukraine</collectingCountry>
.
</paragraph>
<paragraph id="3566A505E264B61B54B5F9BEC00B1123" blockId="89.[164,778,1578,1691]" box="[164,286,1578,1599]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B54B5F9BEC00B1123" box="[164,286,1578,1599]" italics="true" pageId="89" pageNumber="812">Discussion</emphasis>
</paragraph>
<paragraph id="3566A505E264B61B54B5F9DCC0BD1186" blockId="89.[164,778,1578,1691]" pageId="89" pageNumber="812">
The presence of a colulus, the number of cheliceral teeth, and the genital characters place this species clearly into
<taxonomicName id="F2D9DE86E264B61B5520F911C0B41186" authorityName="Latreille" authorityYear="1804" box="[305,417,1669,1690]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="812" phylum="Arthropoda" rank="genus">
<emphasis id="07AD7917E264B61B5520F911C0B41186" box="[305,417,1669,1690]" italics="true" pageId="89" pageNumber="812">Tegenaria</emphasis>
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.
</paragraph>
</subSubSection>
<subSubSection id="7DC3F68EE264B61B5513F942C26C1621" pageId="89" pageNumber="812" type="reference_group">
<paragraph id="3566A505E264B61B5513F942C3B811F6" blockId="89.[258,685,1749,1802]" box="[258,685,1749,1771]" pageId="89" pageNumber="812">
<taxonomicName id="F2D9DE86E264B61B5513F942C3B811F6" ID-CoL="7BN3W" authority="SIMON, 1907" authorityName="SIMON" authorityYear="1907" box="[258,685,1749,1771]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="846" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E264B61B5513F942C31A11F5" box="[258,527,1750,1771]" italics="true" pageId="89" pageNumber="812">TEGENARIA RACOVITZAI</emphasis>
<bibRefCitation id="5148D8F4E264B61B5608F941C3B811F6" author="Simon E" box="[537,685,1749,1771]" pageId="89" pageNumber="812" pagination="537 - 553" refId="ref88402" refString="Simon E. 1907. Araneae, Chernetes et Opiliones (Premiere serie). Archives de zoologie experimentale et generale 4: 537 - 553." type="journal article" year="1907">SIMON, 1907</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="3566A505E264B61B5542F961C3491016" blockId="89.[258,685,1749,1802]" box="[339,604,1781,1802]" pageId="89" pageNumber="812">
(
<figureCitation id="ADE2B980E264B61B554AF961C3461016" box="[347,595,1781,1802]" captionStart="Figure 17" captionStartId="61.[164,243,1707,1726]" captionTargetId="figure-1@62.[158,1408,197,1818]" captionTargetPageId="62" captionText="Figure 17. A, B, Tegenaria domestica; C, F, G, M, N, Tegenaria racovitzai; D, E, Q, R, Tegenaria hauseri; H-J, Tegenaria bozhkovi; K, L, O, P, Tegenaria campestris; S, T, Tegenaria pieperi; U, Tegenaria schmalfussi; V, Tegenaria faniapollinis; W, Tegenaria percuriosa (holotype of Tegenaria bithyniae); X-Z, Tegenaria croatica sp. nov. Left male palp in ventral (M, O, Q) and retrolateral views (N, P, R); epigyne in ventral (D, F, H, K, S, U–W, Y) and posterioventral views (T), vulva in dorsal (E, G, I, L, U, Z) and lateral views (J); face of female Teg. domestica (A) and Teg. racovitzai (C); cheliceral margins of female Teg. domestica in posterioventral view (B); habitus, dorsal view (X). Scale bars = 0.5 mm (no scale for Teg. faniapollinis, V, and Teg. schmalfussi, U)." figureDoi="http://doi.org/10.5281/zenodo.6983672" httpUri="https://zenodo.org/record/6983672/files/figure.png" pageId="89" pageNumber="812">FIG. 17C, F, G, M, N</figureCitation>
)
</paragraph>
<paragraph id="3566A505E264B61B54B5F888C1E21073" blockId="89.[164,779,1819,1903]" pageId="89" pageNumber="812">
<treatmentCitationGroup id="15C9822BE264B61B54B5F888C1E21073" pageId="89" pageNumber="812">
<treatmentCitation id="B4788314E264B61B54B5F888C364102D" author="Simon E" box="[164,625,1819,1842]" page="548" pageId="89" pageNumber="812" year="1907">
<taxonomicName id="F2D9DE86E264B61B54B5F888C364102D" ID-CoL="7BN3W" authority="Simon, 1907: 548" authorityName="SIMON" authorityPageNumber="548" authorityYear="1907" box="[164,625,1819,1842]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="812" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E264B61B54B5F888C084102D" box="[164,401,1820,1841]" italics="true" pageId="89" pageNumber="812">Tegenaria racovitzai</emphasis>
<bibRefCitation id="5148D8F4E264B61B558FF888C364102D" author="Simon E" box="[414,625,1819,1842]" pageId="89" pageNumber="812" pagination="537 - 553" refId="ref88402" refString="Simon E. 1907. Araneae, Chernetes et Opiliones (Premiere serie). Archives de zoologie experimentale et generale 4: 537 - 553." type="journal article" year="1907">Simon, 1907: 548</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 549, fig. 3b, male;
<treatmentCitation id="B4788314E264B61B54FCF8AEC38C1053" author="Ribera C &amp; Barrientos JA" box="[237,665,1850,1872]" page="191" pageId="89" pageNumber="812" year="1986">
<bibRefCitation id="5148D8F4E264B61B54FCF8AEC38C1053" author="Ribera C &amp; Barrientos JA" box="[237,665,1850,1872]" pageId="89" pageNumber="812" pagination="187 - 197" refId="ref87472" refString="Ribera C, Barrientos JA. 1986. Sobre algunas Tegenarias cavernicolas poco conocidas del Mediterraneo Espanol. Memoires de la Societe royale belge d'Entomologie 33: 187 - 197." type="journal article" year="1986">Ribera &amp; Barrientos, 1986: 191–193</bibRefCitation>
</treatmentCitation>
, figs 6, 7, female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3566A505E264B61B5742FF52C2B517E6" blockId="89.[826,1441,198,250]" pageId="89" pageNumber="812">
<treatmentCitationGroup id="15C9822BE264B61B5742FF52C2B517E6" pageId="89" pageNumber="812">
<treatmentCitation id="B4788314E264B61B5742FF52C42317C7" author="Simon E" box="[851,1334,198,220]" page="211" pageId="89" pageNumber="812" year="1916">
<taxonomicName id="F2D9DE86E264B61B5742FF52C42317C7" ID-CoL="5548C" authority="Simon, 1916: 211" authorityName="Simon" authorityPageNumber="211" authorityYear="1916" box="[851,1334,198,220]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="812" phylum="Arthropoda" rank="species" species="antrorum">
<emphasis id="07AD7917E264B61B5742FF52C55117C0" box="[851,1092,198,220]" italics="true" pageId="89" pageNumber="812">Tegenaria antrorum</emphasis>
<bibRefCitation id="5148D8F4E264B61B5046FF52C42317C7" author="Simon E" box="[1111,1334,198,220]" pageId="89" pageNumber="812" pagination="209 - 211" refId="ref88429" refString="Simon E. 1916. Descriptions de plusiers especes d'arachnides recemment decouvertes en France (Troisieme note). Bulletin de la Societe Entomologique de France 1916: 209 - 211." type="journal article" year="1916">Simon, 1916: 211</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, female, syn. nov.
</treatmentCitationGroup>
</paragraph>
<paragraph id="3566A505E264B61B572BFEBCC26C1621" blockId="89.[826,1441,296,532]" box="[826,889,296,317]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B572BFEBCC26C1621" box="[826,889,296,317]" italics="true" pageId="89" pageNumber="812">Types</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="7DC3F68EE264B618572BFED2C0E91072" lastPageId="90" lastPageNumber="813" pageId="89" pageNumber="812" type="materials_examined">
<paragraph id="3566A505E264B61B572BFED2C4691686" blockId="89.[826,1441,296,532]" pageId="89" pageNumber="812">
<materialsCitation id="85B1AF58E264B61B572BFED2C4691686" ID-GBIF-Occurrence="3886835602" collectingDate="1905-08-20" collectionCode="MNHN, 1965, 45, 23627" collectorName="Racovitza &amp; Janel." country="Spain" county="Huesca" location="Fanlo" municipality="Hoya de Huesca" pageId="89" pageNumber="812" specimenCount="2" specimenCount-male="1" stateProvince="Aragon" typeStatus="holotype">
<typeStatus id="EA621BA7E264B61B572BFED2C2B11640" box="[826,932,326,348]" pageId="89" pageNumber="812" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="4DCEE595E264B61B57A5FED2C2EC1640" box="[948,1017,326,348]" name="Spain" pageId="89" pageNumber="812">Spain</collectingCountry>
:
<collectingRegion id="F71D6BE7E264B61B5019FED3C54B1640" box="[1032,1118,327,348]" country="Spain" name="Aragon" pageId="89" pageNumber="812">Aragon</collectingRegion>
:
<collectingCounty id="DC07DD89E264B61B507EFED3C5D31640" box="[1135,1222,327,348]" pageId="89" pageNumber="812">Huesca</collectingCounty>
,
<collectingMunicipality id="D5023F7FE264B61B50C7FED3C4881640" box="[1238,1437,326,348]" pageId="89" pageNumber="812">Hoya de Huesca</collectingMunicipality>
,
<location id="3006F3DEE264B61B572BFEF1C26B1667" LSID="urn:lsid:plazi:treatment:BD701413E264B61854D5FB15C0E91072:3006F3DEE264B61B572BFEF1C26B1667" box="[826,894,357,379]" country="Spain" county="Huesca" municipality="Hoya de Huesca" name="Fanlo" pageId="89" pageNumber="812" stateProvince="Aragon">Fanlo</location>
, ‘
<location id="3006F3DEE264B61B579FFEF1C5D31667" LSID="urn:lsid:plazi:treatment:BD701413E264B61854D5FB15C0E91072:3006F3DEE264B61B579FFEF1C5D31667" box="[910,1222,357,379]" country="Spain" county="Huesca" municipality="Hoya de Huesca" name="Cueva abaho de los Gloces'" pageId="89" pageNumber="812" stateProvince="Aragon">Cueva abaho de los Gloces’</location>
,
<specimenCount id="23DF6E8CE264B61B50C3FEF0C5FE1666" box="[1234,1259,356,378]" pageId="89" pageNumber="812" type="male">♂</specimenCount>
,
<specimenCount id="23DF6E8CE264B61B50E8FEF1C4221667" box="[1273,1335,357,379]" pageId="89" pageNumber="812" type="generic">1 juv.</specimenCount>
(
<collectionCode id="53C83DC0E264B61B5154FEF1C2E31685" pageId="89" pageNumber="812">MNHN, 1965, 45, 23627</collectionCode>
),
<date id="416783C5E264B61B501AFE17C5871685" box="[1035,1170,387,409]" pageId="89" pageNumber="812" value="1905-08-20">
<collectingDate id="51237A2DE264B61B501AFE17C5871685" box="[1035,1170,387,409]" pageId="89" pageNumber="812" value="1905-08-20">20.viii.1905</collectingDate>
</date>
,
<collectorName id="982CC0D3E264B61B50B1FE10C41A1686" box="[1184,1295,388,410]" pageId="89" pageNumber="812">Racovitza</collectorName>
&amp;
<collectorName id="982CC0D3E264B61B5127FE10C4691686" box="[1334,1404,388,410]" pageId="89" pageNumber="812">Janel.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="3566A505E264B61B5742FE36C2DF1508" blockId="89.[826,1441,296,532]" pageId="89" pageNumber="812">
<materialsCitation id="85B1AF58E264B61B5742FE36C2D31508" ID-GBIF-Occurrence="3886835495" collectionCode="MNHN" collectorName="de Villefranche'" country="France" location="Grotte de Villefranche'" pageId="89" pageNumber="812" specimenCount="2" specimenCount-female="2" typeStatus="syntype">
Sub
<taxonomicName id="F2D9DE86E264B61B578FFE37C58E16A4" authorityName="Simon" authorityYear="1916" box="[926,1179,419,440]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="812" phylum="Arthropoda" rank="species" species="antrorum">
<emphasis id="07AD7917E264B61B578FFE37C58E16A4" box="[926,1179,419,440]" italics="true" pageId="89" pageNumber="812">Tegenaria antrorum</emphasis>
</taxonomicName>
:
<typeStatus id="EA621BA7E264B61B50D1FE37C43D16A4" box="[1216,1320,419,440]" pageId="89" pageNumber="812" type="syntype">syntypes</typeStatus>
.
<collectingCountry id="4DCEE595E264B61B515AFE37C48B16A4" box="[1355,1438,419,440]" name="France" pageId="89" pageNumber="812">France</collectingCountry>
: Pyrénées-Orientales: Prades, Villefranche-de- Conflent, ‘
<location id="3006F3DEE264B61B57BEFE4BC5A216E9" LSID="urn:lsid:plazi:treatment:BD701413E264B61854D5FB15C0E91072:3006F3DEE264B61B57BEFE4BC5A216E9" box="[943,1207,479,501]" country="France" name="Grotte de Villefranche'" pageId="89" pageNumber="812">
Grotte
<collectorName id="982CC0D3E264B61B5013FE4BC5A216E9" box="[1026,1207,479,501]" pageId="89" pageNumber="812">de Villefranche’</collectorName>
</location>
,
<specimenCount id="23DF6E8CE264B61B50D5FE4BC5E516E9" box="[1220,1264,479,501]" pageId="89" pageNumber="812" type="female">2 ♀</specimenCount>
(
<collectionCode id="53C83DC0E264B61B50EEFE4BC44316E8" box="[1279,1366,479,500]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="89" pageNumber="812">MNHN</collectionCode>
, 1965, 581), Simon
</materialsCitation>
.
</paragraph>
<paragraph id="3566A505E264B61B572BFDD5C549154A" blockId="89.[826,1116,577,630]" box="[826,1116,577,598]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B572BFDD5C549154A" box="[826,1116,577,598]" italics="true" pageId="89" pageNumber="812">Other material examined</emphasis>
</paragraph>
<paragraph id="3566A505E264B61B572BFDF4C2DB1569" blockId="89.[826,1116,577,630]" box="[826,974,607,630]" pageId="89" pageNumber="812">
<materialsCitation id="85B1AF58E264B61B572BFDF4C2DD1569" ID-GBIF-Occurrence="3886835508" box="[826,968,607,630]" country="Spain" location="Spain" pageId="89" pageNumber="812" specimenCount="3" specimenCount-male="3">
<collectingCountry id="4DCEE595E264B61B572BFDF4C269156A" box="[826,892,608,630]" name="Spain" pageId="89" pageNumber="812">Spain</collectingCountry>
(
<specimenCount id="23DF6E8CE264B61B579EFDF4C2AA1569" box="[911,959,607,629]" pageId="89" pageNumber="812" type="male">3 ♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3566A505E264B61B572BFD30C2BF15A5" blockId="89.[826,1441,676,819]" box="[826,938,676,697]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B572BFD30C2BF15A5" box="[826,938,676,697]" italics="true" pageId="89" pageNumber="812">Diagnosis</emphasis>
</paragraph>
<paragraph id="3566A505E264B61B572BFD56C5D4142F" blockId="89.[826,1441,676,819]" pageId="89" pageNumber="812">
<taxonomicName id="F2D9DE86E264B61B572BFD56C53715CB" authorityName="SIMON" authorityYear="1907" box="[826,1058,706,727]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="89" pageNumber="812" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E264B61B572BFD56C53715CB" box="[826,1058,706,727]" italics="true" pageId="89" pageNumber="812">Tegenaria racovitzai</emphasis>
</taxonomicName>
can be recognized by the distinct reduced conductor, the distinct epigyne with sharply pointed ‘pseudo teeth’, and the posterior sclerite having a convex anterior margin.
</paragraph>
<paragraph id="3566A505E264B61B572BFCF5C2AE146A" blockId="89.[826,1441,865,1437]" box="[826,955,865,886]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B572BFCF5C2AE146A" box="[826,955,865,886]" italics="true" pageId="89" pageNumber="812">Description</emphasis>
</paragraph>
<paragraph id="3566A505E264B61B572BFC14C2F714CF" blockId="89.[826,1441,865,1437]" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B572BFC14C2F11489" box="[826,996,896,917]" italics="true" pageId="89" pageNumber="812">Measurements:</emphasis>
A description of female, including some measurements, was provided by
<bibRefCitation id="5148D8F4E264B61B50EDFC0AC2CB14CF" author="Ribera C &amp; Barrientos JA" pageId="89" pageNumber="812" pagination="187 - 197" refId="ref87472" refString="Ribera C, Barrientos JA. 1986. Sobre algunas Tegenarias cavernicolas poco conocidas del Mediterraneo Espanol. Memoires de la Societe royale belge d'Entomologie 33: 187 - 197." type="journal article" year="1986">Ribera &amp; Barrientos (1986)</bibRefCitation>
.
</paragraph>
<paragraph id="3566A505E264B61B5742FC4FC4871281" blockId="89.[826,1441,865,1437]" pageId="89" pageNumber="812">
Male (
<emphasis id="07AD7917E264B61B57B0FC48C2A014ED" box="[929,949,988,1009]" italics="true" pageId="89" pageNumber="812">N</emphasis>
= 1): CL 4.22, CW 2.98, STL 2.16, STW 1.81, OL 4.70, OW 2.78. Leg I (5.24, 1.66, 5.26, 5.55, 3.01), II (5.05, 1.60, 4.82, 5.56, 2.78), III (4.79, 1.49, 4.40, 5.75, 2.60), IV (6.01, 1.84, 5.38, 7.00, 3.40). Pedipalp (1.84, 0.66, 0.92, 1.77), bulbL 1.01. Female (
<emphasis id="07AD7917E264B61B5753FBE1C2431396" box="[834,854,1141,1162]" italics="true" pageId="89" pageNumber="812">N</emphasis>
= 1): CL 3.66, CW 2.65, STL 1.92, STW 1.68. Leg I (4.41, 1.28, 3.90, 4.40, 2.36), II (4.25, 1.35, 3.79, 4.27, 2.24), III (3.86, 1.23, 3.36, 4.31, 1.85), IV (4.83, 1.26, 4.39, 5.76, 1.78). Pedipalp (1.51, 0.60, 0.98, 1.61). EPL 0.72, EPW 1.11, ATL 0.22, ATW 0.39. Eyes: PME 0.14–0.15, PLE 0.15–0.16, AME 0.10–0.11, ALE 0.16– 0.17. Eye distances: PME–PME 1–1.5 x PME, PME– AME 1 x PME, PME–PLE 1–1.5 x PME, PME–ALE 0.5–1 x PME, AME–AME 0.5–1 x AME, AME–ALE 0.5–1 x AME. CLY1 2.5 x AME, CLY2 1–1.5 x ALE.
</paragraph>
<paragraph id="3566A505E264B618572BFA5FC06517E6" blockId="89.[826,1441,1483,1903]" lastBlockId="90.[144,759,198,250]" lastPageId="90" lastPageNumber="813" pageId="89" pageNumber="812">
<emphasis id="07AD7917E264B61B572BFA5FC2A312FC" box="[826,950,1483,1504]" italics="true" pageId="89" pageNumber="812">Male palp:</emphasis>
RTA with three branches, ventral branch long, drawn-out bulge with distinct ventral ridge, lateral branch much smaller, protruding and distally pointed, dorsal branch strongly sclerotized, broadly protruding, moderately longer than wide, distally broadly truncated. Filiform embolus length about 1.5 x CB, originating at 8–9 o’clock position, distal tip at 2 o’clock position. Conductor reduced to transparent lamelliform appendage with lateral margin moderately folded. Terminal end simple and moderately pointed. Connection of conductor to tegulum membranous or moderately sclerotized. MA originating at 4–5 o’clock position, moderately protruding, distally with spoon-like sclerite. MA membranously connected to tegulum. Basal part of tegulum clearly visible, with continuous margin.
</paragraph>
<paragraph id="3566A505E267B6185481FEA1C0EC1586" blockId="90.[144,759,308,666]" pageId="90" pageNumber="813">
<emphasis id="07AD7917E267B6185481FEA1C0971655" box="[144,386,308,330]" italics="true" pageId="90" pageNumber="813">Epigyne and vulva:</emphasis>
Epigyne medially with pale, membranous area. Posterior sclerite expressed as sclerotized bulge with anterior margin convex. CO between the membranous median area and the posterior sclerite. Epigynal ‘pseudo teeth’ present and sharply pointed. Vulva consists of distinguishable CD, RC, and FD. CD very short leading into globular, smoothly sclerotized RC. RC almost touching each other. An additional small globular structure (second pair of RC) attached to the large RC, which are visible through the epigynal plate. FD only represented by small, leaf-shaped appendages.
</paragraph>
<paragraph id="3566A505E267B6185481FD40C03013C8" blockId="90.[144,759,724,1236]" pageId="90" pageNumber="813">
<emphasis id="07AD7917E267B6185481FD40C0F115F5" box="[144,484,724,746]" italics="true" pageId="90" pageNumber="813">Other important characters:</emphasis>
Cheliceral promargin with three, retromargin with four to five teeth. Colulus developed as trapezoidal plate with the distal margin medially notched. Same pattern of distal spigots on PMS (in females) as described for the
<typeStatus id="EA621BA7E267B61856D4FCDBC3E31478" box="[709,758,847,868]" pageId="90" pageNumber="813">type</typeStatus>
species. PLS with distal segment shorter than basal segment. Tarsal trichobothria on cymbium and palp tarsus absent. Tarsal trichobothria seven to nine. Leg spination: male palp (2–0–0–0 or 3–0–0, 2–0–0, 1–0–2–0), female palp (2–1–0–0 or 3–0–0–0, 2–0–0, 2–2–0–0), leg femora (2–2–2–0 or 2–3–2–0, 2–2–2–0, 2–2–2–0, 2–1–1–0 or 2–2–1–0), patellae (all 2–0–0), tibiae (1–2–0–2p+1 or 1–2–2–2p+1 or 1–2–0–3p, 1–2– 2–1+2p or 1–2–2–3p, 1–2–2–2+1p or 1–2–2–3p, 1–2– 2–2+1p or 1–2–2–3p), metatarsi (0–2–2–3p+1, 0–3–2– 3p+1, 2–3–3–3p+1, 2–3–3–3p+1), tarsi (I &amp; II 0, III &amp; IV 0–0–1–0).
</paragraph>
<paragraph id="3566A505E267B6185481FA99C059125E" blockId="90.[144,759,1293,1346]" pageId="90" pageNumber="813">
<emphasis id="07AD7917E267B6185481FA99C01B123E" box="[144,270,1293,1314]" italics="true" pageId="90" pageNumber="813">Coloration:</emphasis>
No coloration pattern visible on the specimens examined.
</paragraph>
<paragraph id="3566A505E267B6185481FAEFC00E128C" blockId="90.[144,758,1403,1486]" box="[144,283,1403,1424]" pageId="90" pageNumber="813">
<emphasis id="07AD7917E267B6185481FAEFC00E128C" box="[144,283,1403,1424]" italics="true" pageId="90" pageNumber="813">Distribution</emphasis>
</paragraph>
<paragraph id="3566A505E267B6185481FA0EC07412D2" blockId="90.[144,758,1403,1486]" pageId="90" pageNumber="813">
Reported from northern
<collectingCountry id="4DCEE595E267B61855B9FA0EC0FF12AC" box="[424,490,1434,1456]" name="Spain" pageId="90" pageNumber="813">Spain</collectingCountry>
and south-west
<collectingCountry id="4DCEE595E267B61856B6FA0EC3E312B3" box="[679,758,1434,1455]" name="France" pageId="90" pageNumber="813">France</collectingCountry>
(Pyrenees region).
</paragraph>
<paragraph id="3566A505E267B6185481F99DC01F1102" blockId="90.[144,759,1545,1902]" box="[144,266,1545,1566]" pageId="90" pageNumber="813">
<emphasis id="07AD7917E267B6185481F99DC01F1102" box="[144,266,1545,1566]" italics="true" pageId="90" pageNumber="813">Discussion</emphasis>
</paragraph>
<paragraph id="3566A505E267B6185481F9B3C0E91072" blockId="90.[144,759,1545,1902]" pageId="90" pageNumber="813">
<bibRefCitation id="5148D8F4E267B6185481F9B3C0331121" author="Simon E" box="[144,294,1575,1597]" pageId="90" pageNumber="813" pagination="537 - 553" refId="ref88402" refString="Simon E. 1907. Araneae, Chernetes et Opiliones (Premiere serie). Archives de zoologie experimentale et generale 4: 537 - 553." type="journal article" year="1907">Simon (1907)</bibRefCitation>
assigned
<taxonomicName id="F2D9DE86E267B6185587F9B3C32E1120" authorityName="SIMON" authorityYear="1907" box="[406,571,1575,1596]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E267B6185587F9B3C32E1120" box="[406,571,1575,1596]" italics="true" pageId="90" pageNumber="813">Teg. racovitzai</emphasis>
</taxonomicName>
to his
<taxonomicName id="F2D9DE86E267B6185697F9B3C0141146" authorityName=": Walckenaer" authorityYear="1805" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="domestica">
<emphasis id="07AD7917E267B6185697F9B3C0141146" italics="true" pageId="90" pageNumber="813">Tegenaria domestica</emphasis>
</taxonomicName>
-group. Later, he described
<taxonomicName id="F2D9DE86E267B6185644F9D2C0201166" authority="(Simon, 1916)" baseAuthorityName="Simon" baseAuthorityYear="1916" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="antrorum">
<emphasis id="07AD7917E267B6185644F9D2C3E31147" box="[597,758,1606,1627]" italics="true" pageId="90" pageNumber="813">Teg. antrorum</emphasis>
(
<bibRefCitation id="5148D8F4E267B6185486F9F0C0381166" author="Simon E" box="[151,301,1636,1658]" pageId="90" pageNumber="813" pagination="209 - 211" refId="ref88429" refString="Simon E. 1916. Descriptions de plusiers especes d'arachnides recemment decouvertes en France (Troisieme note). Bulletin de la Societe Entomologique de France 1916: 209 - 211." type="journal article" year="1916">Simon, 1916</bibRefCitation>
)
</taxonomicName>
, which he placed near
<taxonomicName id="F2D9DE86E267B6185643F9F0C3E21165" authorityName="SIMON" authorityYear="1907" box="[594,759,1636,1657]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E267B6185643F9F0C3E21165" box="[594,759,1636,1657]" italics="true" pageId="90" pageNumber="813">Teg. racovitzai</emphasis>
</taxonomicName>
within his
<taxonomicName id="F2D9DE86E267B6185533F917C3141184" authorityName="SIMON" authorityYear="1873" box="[290,513,1667,1688]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="armigera">
<emphasis id="07AD7917E267B6185533F917C3141184" box="[290,513,1667,1688]" italics="true" pageId="90" pageNumber="813">Tegenaria armigera</emphasis>
</taxonomicName>
-group.
<bibRefCitation id="5148D8F4E267B6185674F917C3E31185" author="Fage L" box="[613,758,1667,1689]" pageId="90" pageNumber="813" pagination="99 - 291" refId="ref83837" refString="Fage L. 1931. Araneae, 5 e serie, precedee d'un essai sur l'evolution souterraine et son determinisme. Archives de Zoologie Experimentale et Generale 71: 99 - 291." type="journal article" year="1931">Fage (1931)</bibRefCitation>
agreed with this placement, in contrast to
<bibRefCitation id="5148D8F4E267B618568BF935C1F711CA" author="Brignoli PM" pageId="90" pageNumber="813" pagination="69 - 71" refId="ref82457" refString="Brignoli PM. 1977 c. Sur quelqes Tegenaria cavernicoles d'Espagne (Araneae, Agelenidae) Comunicacions del 6 e. Simposium d'Espeleologia. Teressa: Bioespeleologia, 69 - 71." type="book chapter" year="1977">Brignoli (1977c)</bibRefCitation>
who mentioned that
<taxonomicName id="F2D9DE86E267B61855F4F954C39C11C9" authorityName="SIMON" authorityYear="1907" box="[485,649,1728,1749]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E267B61855F4F954C39C11C9" box="[485,649,1728,1749]" italics="true" pageId="90" pageNumber="813">Teg. racovitzai</emphasis>
</taxonomicName>
does not show close affinities to any of the
<taxonomicName id="F2D9DE86E267B6185621F94BC01F100E" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="variety" species="domestica" variety="atrica">
<emphasis id="07AD7917E267B6185621F94BC1AF100E" italics="true" pageId="90" pageNumber="813">Tegenaria domestica</emphasis>
,
<emphasis id="07AD7917E267B61854D6F969C01F100E" box="[199,266,1789,1810]" italics="true" pageId="90" pageNumber="813">atrica</emphasis>
</taxonomicName>
, or
<emphasis id="07AD7917E267B6185527F969C0EA100E" box="[310,511,1789,1810]" italics="true" pageId="90" pageNumber="813">
<taxonomicName id="F2D9DE86E267B6185527F969C08A100E" authorityName="SIMON" authorityYear="1873" box="[310,415,1789,1810]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="armigera">armigera</taxonomicName>
/
<taxonomicName id="F2D9DE86E267B61855BCF969C0EA100E" authorityName=": Lehtinen" authorityYear="1967" box="[429,511,1789,1810]" class="Arachnida" family="Agelenidae" genus="Pseudotegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="oribata">oribata</taxonomicName>
</emphasis>
-groups. In our analyses, the original hypothesis of Simon had most support because
<taxonomicName id="F2D9DE86E267B6185571F8AFC310104C" authorityName="SIMON" authorityYear="1907" box="[352,517,1851,1872]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="species" species="racovitzai">
<emphasis id="07AD7917E267B6185571F8AFC310104C" box="[352,517,1851,1872]" italics="true" pageId="90" pageNumber="813">Teg. racovitzai</emphasis>
</taxonomicName>
represents a basal branch of
<taxonomicName id="F2D9DE86E267B6185516F8CDC0621072" authorityName="Latreille" authorityYear="1804" box="[263,375,1881,1902]" class="Arachnida" family="Agelenidae" genus="Tegenaria" kingdom="Animalia" order="Araneae" pageId="90" pageNumber="813" phylum="Arthropoda" rank="genus">
<emphasis id="07AD7917E267B6185516F8CDC0621072" box="[263,375,1881,1902]" italics="true" pageId="90" pageNumber="813">Tegenaria</emphasis>
</taxonomicName>
(
<figureCitation id="ADE2B980E267B6185598F8CCC0C51072" box="[393,464,1880,1902]" captionStart="Figure 3" captionStartId="22.[144,223,198,217]" captionTargetId="graphics-2@21.[210,1354,194,1812]" captionTargetPageId="21" captionText="Figure 3. Most parsimonious strict consensus tree based on morphological data with equal character weights. Branch support is indicated by jack-knife values (1000 pseudoreplicates)/Bremer support at the nodes. Analysis with implied weighting (K = 10) resulted in additional supported clades (jack-knife support): Tegenaria mercanturensis + Tegenaria mirifica (69), Tegenaria montana + Tegenaria rilaensis (52), Tegenaria parvula + Tegenaria silvestris (55). Black dots indicate apomorpies, white dots indicate homoplasies. Numbers above the dots correspond to the character number, numbers below the dots to the character state." figureDoi="http://doi.org/10.5281/zenodo.6984004" httpUri="https://zenodo.org/record/6984004/files/figure.png" pageId="90" pageNumber="813">Figs 3</figureCitation>
,
<figureCitation id="ADE2B980E267B61855CEF8CDC0FB1072" box="[479,494,1881,1902]" captionStart="Figure 7" captionStartId="29.[164,243,1338,1357]" captionTargetBox="[164,1441,198,1284]" captionTargetId="graphics-186@29.[164,1441,198,1284]" captionTargetPageId="29" captionText="Figure 7. Combined DNA and morphological data (cytochrome c oxidase subunit 1, nicotinamide adenine dinucleotide dehydrogenase subunit 1, 28S, and morphological data) Bayesian tree. Posterior probabilities of clades are expressed in percentages and given above branches. Clade support (&gt; 50) from the resampling method (jack-knife, 1000 replications) based on parsimony analysis with implied weighting (K = 10) is given below the branches. Bremer support (&gt; 4) is given to the right of the corresponding node. Abbreviations: AT, Austria; CH, Switzerland; DE, Germany; ES, Spain; FR, France; GR, Greece; IT, Italy; PT, Portugal; SE, Sweden; US, United States." figureDoi="http://doi.org/10.5281/zenodo.5282989" httpUri="https://zenodo.org/record/5282989/files/figure.png" pageId="90" pageNumber="813">7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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