treatments-xml/data/BD/09/14/BD091476FF8F805EFF7CFE2BDEB4F86C.xml

<document id="ED118D48D7669FD012F1023FDB34ABC8" ID-DOI="10.11646/zootaxa.3619.4.3" ID-GBIF-Dataset="d90b27f2-9c8a-411b-95bf-062e49931acb" ID-ISSN="1175-5326" ID-Zenodo-Dep="216757" ID-ZooBank="995ACA53-EB4C-4FE2-8A0E-8876FD2AA5D0" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460578460905" checkinUser="plazi" docAuthor="Giłka, Wojciech &amp; Zakrzewska, Marta" docDate="2013" docId="BD091476FF8F805EFF7CFE2BDEB4F86C" docLanguage="en" docName="zt03619p459.pdf" docOrigin="Zootaxa 3619 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus cotopaxi Giłka &amp; Zakrzewska, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="459" masterDocId="41306C0EFF8C8058FFEBFF9CDC00FF8F" masterDocTitle="A contribution to the systematics of Neotropical Tanytarsus van der Wulp: first descriptions from Ecuador (Diptera: Chironomidae: Tanytarsini)" masterLastPageNumber="459" masterPageNumber="453" pageNumber="456" updateTime="1698323720027" updateUser="plazi">
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<mods:title id="99C578AF4254E73B9DDAD612118C7BCF">A contribution to the systematics of Neotropical Tanytarsus van der Wulp: first descriptions from Ecuador (Diptera: Chironomidae: Tanytarsini)</mods:title>
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<mods:namePart id="3437E727D55F2D2F71071266C52C2AEA">Giłka, Wojciech</mods:namePart>
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<mods:namePart id="C10C7A8629356C05C091D3AB6366B03A">Zakrzewska, Marta</mods:namePart>
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<mods:date id="5366729FA60E23BE6791AA31E31A7FCA">2013</mods:date>
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<subSubSection id="7DBAF6EBFF8F805BFF7CFE2BDDEEFE5E" box="[151,494,439,465]" pageId="3" pageNumber="456" type="nomenclature">
<paragraph id="351FA560FF8F805BFF7CFE2BDDEEFE5E" blockId="3.[151,494,439,465]" box="[151,494,439,465]" pageId="3" pageNumber="456">
<heading id="6E57120CFF8F805BFF7CFE2BDDEEFE5E" bold="true" box="[151,494,439,465]" fontSize="11" level="1" pageId="3" pageNumber="456" reason="1">
<emphasis id="07D47972FF8F805BFF7CFE2BDDEEFE5E" bold="true" box="[151,494,439,465]" pageId="3" pageNumber="456">
<taxonomicName id="F2A0DEE3FF8F805BFF7CFE2BDD89FE5E" box="[151,393,439,465]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="cotopaxi" status="sp. nov.">
<emphasis id="07D47972FF8F805BFF7CFE2BDD89FE5E" bold="true" box="[151,393,439,465]" italics="true" pageId="3" pageNumber="456">Tanytarsus cotopaxi</emphasis>
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<taxonomicNameLabel id="1CE7C409FF8F805BFE7BFE2BDDEEFE5E" box="[400,494,439,465]" pageId="3" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="7DBAF6EBFF8F805BFF7CFE63D8D1FD0D" pageId="3" pageNumber="456" type="materials_examined">
<paragraph id="351FA560FF8F805BFF7CFE63DFD4FDB5" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF7CFE63DD4AFD98" bold="true" box="[151,330,510,535]" pageId="3" pageNumber="456">
<typeStatus id="EA1B1BC2FF8F805BFF7CFE63DCD2FD98" box="[151,210,511,535]" pageId="3" pageNumber="456">Type</typeStatus>
material.
</emphasis>
<typeStatus id="EA1B1BC2FF8F805BFEBDFE62DDC1FD99" box="[342,449,510,534]" pageId="3" pageNumber="456" type="holotype">Holotype</typeStatus>
, adult male: 
<collectingCountry id="4DB7E5F0FF8F805BFDB4FE62DEEAFD99" box="[607,746,510,534]" name="Ecuador" pageId="3" pageNumber="456">ECUADOR</collectingCountry>
, Pichincha province, San Miguel de Los Bancos (canton), Mindo, c. 
<quantity id="F2580885FF8F805BFEE1FDBEDD5FFDB5" box="[266,351,546,570]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="456" unit="m" value="1400.0">1400 m</quantity>
a.s.l., 
<date id="411E83A0FF8F805BFE43FDBEDE83FDB5" box="[424,643,546,570]" pageId="3" pageNumber="456" value="1999-09-16">16 September 1999</date>
, netting, leg. Dawid Graczyk.
</paragraph>
<paragraph id="351FA560FF8F805BFF2CFDDAD8D1FD0D" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF2CFDDADDE5FDD0" bold="true" box="[199,485,582,607]" pageId="3" pageNumber="456">Derivation of the name.</emphasis>
The specific name is referred to the 
<taxonomicName id="F2A0DEE3FF8F805BFC61FDDADFF3FDD1" box="[906,1011,582,606]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="458" phylum="Arthropoda" rank="species" species="cotopaxi">Cotopaxi</taxonomicName>
, the biggest volcano in the Pichincha province where the specimen was collected. The name should be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="7DBAF6EBFF8F805BFF2CFD12DD69FCB9" pageId="3" pageNumber="456" type="diagnosis">
<paragraph id="351FA560FF8F805BFF2CFD12DD69FCB9" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF2CFD12DD41FD28" bold="true" box="[199,321,654,679]" pageId="3" pageNumber="456">Diagnosis.</emphasis>
AR low, 0.48. Hypopygium with bristle-shaped median setae; lateral teeth absent; anal point expanded in proximal part, with spinulae arranged in regular row; microtrichia between crests absent; superior volsella ellipse-shaped, deeply excavated at apex, with small field of microtrichia at base; digitus long, with swollen roundish apex; inferior volsella parallel-sided, tapering to round apex; median volsella with setiform and pectinate lamellae.
</paragraph>
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<subSubSection id="7DBAF6EBFF8F805BFF2CFCDEDF2BF85D" pageId="3" pageNumber="456" type="description">
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<emphasis id="07D47972FF8F805BFF2CFCDEDDE5FCD4" bold="true" box="[199,485,834,859]" pageId="3" pageNumber="456">Description. Adult male</emphasis>
(n = 1)
</paragraph>
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Wing length 
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.
</paragraph>
<paragraph id="351FA560FF8F805BFF22FC17DDB6FC65" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF22FC17DD52FC2C" box="[201,338,907,931]" italics="true" pageId="3" pageNumber="456">Colouration</emphasis>
. Eyes black; antennal pedicel, head capsule, ground colour of thorax, scutellum, haltere, proximal parts of legs and abdomen greenish yellow; wing membrane with pale greenish undertone; remaining body parts slightly darker, brownish.
</paragraph>
<paragraph id="351FA560FF8F805BFF22FC6ADE2FFBBD" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF22FC6ADD06FB81" box="[201,262,1014,1038]" italics="true" pageId="3" pageNumber="456">Head</emphasis>
. Antenna with 13 flagellomeres; AR 0.48. Frontal tubercles absent. Length of palpomeres II–V (μm): 38, 94, 101, 179. Clypeus with 15 setae.
</paragraph>
<paragraph id="351FA560FF8F805BFF22FBA2D8F0FBD9" blockId="3.[151,1437,510,1297]" box="[201,1264,1086,1111]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF22FBA2DD98FBD9" box="[201,408,1086,1110]" italics="true" pageId="3" pageNumber="456">Thorax chaetotaxy</emphasis>
. Ac 15; Dc 8 on each side; Pa 1 on each side; Scts 4, outermost pair minute.
</paragraph>
<paragraph id="351FA560FF8F805BFF22FBF8DE91FB2E" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF22FBF8DD02FBF4" box="[201,258,1124,1147]" italics="true" pageId="3" pageNumber="456">Wing</emphasis>
. 
<figureCitation id="AD9BB9E5FF8F805BFEFBFBFEDD52FBF5" box="[272,338,1122,1146]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[254,1333,193,1912]" captionTargetId="figure@4.[254,1333,193,1947]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus cotopaxi sp. nov., adult male. A—wing, B—hypopygium, C—median volsella (magnified x 2 relative to hypopygium)." httpUri="https://zenodo.org/record/216759/files/figure.png" pageId="3" pageNumber="456">Fig. 2</figureCitation>
A. Membrane below R and An with sparse macrotrichia only in distal part, cells r4+5, m1+2 and distal half of m3+4 covered with dense macrotrichia.
</paragraph>
<paragraph id="351FA560FF8F805BFF22FB2DD977FA9F" blockId="3.[151,1437,510,1297]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF22FB2DDCFEFB47" box="[201,254,1201,1224]" italics="true" pageId="3" pageNumber="456">Legs</emphasis>
. Fore tibia with straight slender spur. Tibial combs of mid and hind leg separated; spurs of mid leg unequal: one long and straight and the second short, tooth-like; spurs of hind leg equal, long and slightly bent. Basitarsus of mid leg bearing 4 hook-shaped sensilla chaetica. Length of leg segments and leg ratios in 
<tableCitation id="782290DBFF8F805BFAF6FB64D973FA9F" box="[1309,1395,1272,1296]" captionStart="TABLE 2" captionStartId="3.[151,235,1344,1365]" captionTargetBox="[159,1305,1393,1551]" captionTargetPageId="3" captionText="TABLE 2. Lengths (μm) of leg segments and leg ratios of male Tanytarsus cotopaxi sp. nov." httpUri="http://table.plazi.org/id/61DFF5E8FF8F805BFF7CFADCD844FAD9" pageId="3" pageNumber="456" tableUuid="61DFF5E8FF8F805BFF7CFADCD844FAD9">Table 2</tableCitation>
.
</paragraph>
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<paragraph id="351FA560FF8F805BFF7CFADCD844FAD9" blockId="3.[151,1092,1343,1366]" box="[151,1092,1343,1366]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF7CFADCDD04FADA" bold="true" box="[151,260,1344,1365]" pageId="3" pageNumber="456">TABLE 2.</emphasis>
Lengths (μm) of leg segments and leg ratios of male 
<taxonomicName id="F2A0DEE3FF8F805BFCC9FADDDFEDFAD9" box="[802,1005,1345,1366]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="cotopaxi" status="sp. nov.">
<emphasis id="07D47972FF8F805BFCC9FADDDFEDFAD9" box="[802,1005,1345,1366]" italics="true" pageId="3" pageNumber="456">Tanytarsus cotopaxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1CE7C409FF8F805BFC1FFADDD844FAD9" box="[1012,1092,1345,1366]" pageId="3" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="351FA560FF8F805BFF32FAEDD8FAF984" pageId="3" pageNumber="456">
<table id="47A057C0FF8F7FA7FF74FAEDD919F980" box="[159,1305,1393,1551]" gridcols="8" gridrows="4" pageId="3" pageNumber="456">
<tr id="8B90A722FF8F7FA7FF74FAEDD919FA04" box="[159,1305,1393,1419]" gridrow="0" pageId="3" pageNumber="456">
<th id="C841CE5EFF8F7FA7FF74FAEDDCFFFA04" box="[159,255,1393,1419]" gridcol="0" gridrow="0" pageId="3" pageNumber="456">fe</th>
<th id="C841CE5EFF8F7FA7FE85FAEDDD94FA04" box="[366,404,1393,1419]" gridcol="1" gridrow="0" pageId="3" pageNumber="456">ti</th>
<th id="C841CE5EFF8F7FA7FDE8FAEDDE35FA04" box="[515,565,1393,1419]" gridcol="2" gridrow="0" pageId="3" pageNumber="456">ta1</th>
<th id="C841CE5EFF8F7FA7FD73FAEDDEBEFA04" box="[664,702,1393,1419]" gridcol="3" gridrow="0" pageId="3" pageNumber="456">ta2</th>
<th id="C841CE5EFF8F7FA7FCC6FAEDDF53FA04" box="[813,851,1393,1419]" gridcol="4" gridrow="0" pageId="3" pageNumber="456">ta3</th>
<th id="C841CE5EFF8F7FA7FC28FAEDDFE9FA04" box="[963,1001,1393,1419]" gridcol="5" gridrow="0" pageId="3" pageNumber="456">ta4</th>
<th id="C841CE5EFF8F7FA7FBB3FAEDD87EFA04" box="[1112,1150,1393,1419]" gridcol="6" gridrow="0" pageId="3" pageNumber="456">ta5</th>
<th id="C841CE5EFF8F7FA7FB06FAEDD919FA04" box="[1261,1305,1393,1419]" gridcol="7" gridrow="0" pageId="3" pageNumber="456">LR</th>
</tr>
<tr id="8B90A722FF8F7FA7FF74FA01D919FA38" box="[159,1305,1437,1463]" gridrow="1" pageId="3" pageNumber="456">
<th id="C841CE5EFF8F7FA7FF74FA01DCFFFA38" box="[159,255,1437,1463]" gridcol="0" gridrow="1" pageId="3" pageNumber="456">P1 840</th>
<td id="C841CE5EFF8F7FA7FE85FA01DD94FA38" box="[366,404,1437,1463]" gridcol="1" gridrow="1" pageId="3" pageNumber="456">380</td>
<td id="C841CE5EFF8F7FA7FDE8FA01DE35FA38" box="[515,565,1437,1463]" gridcol="2" gridrow="1" pageId="3" pageNumber="456">1270</td>
<td id="C841CE5EFF8F7FA7FD73FA01DEBEFA38" box="[664,702,1437,1463]" gridcol="3" gridrow="1" pageId="3" pageNumber="456">565</td>
<td id="C841CE5EFF8F7FA7FCC6FA01DF53FA38" box="[813,851,1437,1463]" gridcol="4" gridrow="1" pageId="3" pageNumber="456">445</td>
<td id="C841CE5EFF8F7FA7FC28FA01DFE9FA38" box="[963,1001,1437,1463]" gridcol="5" gridrow="1" pageId="3" pageNumber="456">355</td>
<td id="C841CE5EFF8F7FA7FBB3FA01D87EFA38" box="[1112,1150,1437,1463]" gridcol="6" gridrow="1" pageId="3" pageNumber="456">150</td>
<td id="C841CE5EFF8F7FA7FB06FA01D919FA38" box="[1261,1305,1437,1463]" gridcol="7" gridrow="1" pageId="3" pageNumber="456">3.34</td>
</tr>
<tr id="8B90A722FF8F7FA7FF74FA55D919FA6C" box="[159,1305,1481,1507]" gridrow="2" pageId="3" pageNumber="456">
<th id="C841CE5EFF8F7FA7FF74FA55DCFFFA6C" box="[159,255,1481,1507]" gridcol="0" gridrow="2" pageId="3" pageNumber="456">P2 795</th>
<td id="C841CE5EFF8F7FA7FE85FA55DD94FA6C" box="[366,404,1481,1507]" gridcol="1" gridrow="2" pageId="3" pageNumber="456">605</td>
<td id="C841CE5EFF8F7FA7FDE8FA55DE35FA6C" box="[515,565,1481,1507]" gridcol="2" gridrow="2" pageId="3" pageNumber="456">435</td>
<td id="C841CE5EFF8F7FA7FD73FA55DEBEFA6C" box="[664,702,1481,1507]" gridcol="3" gridrow="2" pageId="3" pageNumber="456">195</td>
<td id="C841CE5EFF8F7FA7FCC6FA55DF53FA6C" box="[813,851,1481,1507]" gridcol="4" gridrow="2" pageId="3" pageNumber="456">140</td>
<td id="C841CE5EFF8F7FA7FC28FA55DFE9FA6C" box="[963,1001,1481,1507]" gridcol="5" gridrow="2" pageId="3" pageNumber="456">80</td>
<td id="C841CE5EFF8F7FA7FBB3FA55D87EFA6C" box="[1112,1150,1481,1507]" gridcol="6" gridrow="2" pageId="3" pageNumber="456">55</td>
<td id="C841CE5EFF8F7FA7FB06FA55D919FA6C" box="[1261,1305,1481,1507]" gridcol="7" gridrow="2" pageId="3" pageNumber="456">0.72</td>
</tr>
<tr id="8B90A722FF8F7FA7FF74FA69D919F980" box="[159,1305,1525,1551]" gridrow="3" pageId="3" pageNumber="456">
<th id="C841CE5EFF8F7FA7FF74FA69DCFFF980" box="[159,255,1525,1551]" gridcol="0" gridrow="3" pageId="3" pageNumber="456">P3 820</th>
<td id="C841CE5EFF8F7FA7FE85FA69DD94F980" box="[366,404,1525,1551]" gridcol="1" gridrow="3" pageId="3" pageNumber="456">745</td>
<td id="C841CE5EFF8F7FA7FDE8FA69DE35F980" box="[515,565,1525,1551]" gridcol="2" gridrow="3" pageId="3" pageNumber="456">–</td>
<td id="C841CE5EFF8F7FA7FD73FA69DEBEF980" box="[664,702,1525,1551]" gridcol="3" gridrow="3" pageId="3" pageNumber="456">–</td>
<td id="C841CE5EFF8F7FA7FCC6FA69DF53F980" box="[813,851,1525,1551]" gridcol="4" gridrow="3" pageId="3" pageNumber="456">–</td>
<td id="C841CE5EFF8F7FA7FC28FA69DFE9F980" box="[963,1001,1525,1551]" gridcol="5" gridrow="3" pageId="3" pageNumber="456">–</td>
<td id="C841CE5EFF8F7FA7FBB3FA69D87EF980" box="[1112,1150,1525,1551]" gridcol="6" gridrow="3" pageId="3" pageNumber="456">–</td>
<td id="C841CE5EFF8F7FA7FB06FA69D919F980" box="[1261,1305,1525,1551]" gridcol="7" gridrow="3" pageId="3" pageNumber="456">–</td>
</tr>
</table>
</paragraph>
<paragraph id="351FA560FF8F805BFF2CF9CFDF2BF85D" blockId="3.[151,1436,1618,2002]" pageId="3" pageNumber="456">
<emphasis id="07D47972FF8F805BFF2CF9CFDD56F9E5" box="[199,342,1619,1642]" italics="true" pageId="3" pageNumber="456">Hypopygium</emphasis>
(
<figureCitation id="AD9BB9E5FF8F805BFE8DF9CEDDABF9E5" box="[358,427,1618,1642]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[254,1333,193,1912]" captionTargetId="figure@4.[254,1333,193,1947]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus cotopaxi sp. nov., adult male. A—wing, B—hypopygium, C—median volsella (magnified x 2 relative to hypopygium)." httpUri="https://zenodo.org/record/216759/files/figure.png" pageId="3" pageNumber="456">Fig. 2</figureCitation>
B, C). Gonostylus 100 μm long, slightly swollen in proximal part, tapering to slender tip. Anal tergite with 4 bristle-shaped median setae arranged in transversal row. Anal tergite bands of V-type, separated, with distal sections bent and parallelly tapering to the centre of anal tergite; lateral teeth absent. Anal point stout and long, slightly expanded in proximal part, roundish, with abruptly narrowed tip, apically blunt with 3–4 lateral setae on each side and 4 spinulae arranged in regular row and placed in pit framed by semicircular anal crests; entire area surrounding base of anal point covered with microtrichia, microtrichia between crests absent. Superior volsella ellipse-shaped, deeply excavated at apex, with small field of microtrichia at base, bearing 2 dorsal setae, 2 anteromedian setae and small tubercle (seta apparently broken) at base of digitus. Digitus straight and long, extending far beyond superior volsella, with swollen roundish apex. Inferior volsella parallel-sided, tapering to round apex, with slight dorsomedian ridge (
<figureCitation id="AD9BB9E5FF8F805BFD6AF80ADEC5F821" box="[641,709,1942,1966]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[254,1333,193,1912]" captionTargetId="figure@4.[254,1333,193,1947]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus cotopaxi sp. nov., adult male. A—wing, B—hypopygium, C—median volsella (magnified x 2 relative to hypopygium)." httpUri="https://zenodo.org/record/216759/files/figure.png" pageId="3" pageNumber="456">Fig. 2</figureCitation>
B). Stem of median volsella 12 μm long, slightly curved, bearing 1 short setiform and 3 slender pectinate lamellae (
<figureCitation id="AD9BB9E5FF8F805BFD2EF826DF09F85D" box="[709,777,1978,2002]" captionStart="FIGURE 2" captionStartId="4.[151,250,1968,1991]" captionTargetBox="[254,1333,193,1912]" captionTargetId="figure@4.[254,1333,193,1947]" captionTargetPageId="4" captionText="FIGURE 2. Tanytarsus cotopaxi sp. nov., adult male. A—wing, B—hypopygium, C—median volsella (magnified x 2 relative to hypopygium)." httpUri="https://zenodo.org/record/216759/files/figure.png" pageId="3" pageNumber="456">Fig. 2</figureCitation>
C).
</paragraph>
</subSubSection>
<caption id="61DFF5E8FF88805CFF7CF82CDD41F86A" httpUri="https://zenodo.org/record/216759/files/figure.png" pageId="4" pageNumber="457" targetBox="[254,1333,193,1912]" targetPageId="4">
<paragraph id="351FA560FF88805CFF7CF82CDD41F86A" blockId="4.[151,1436,1968,2021]" pageId="4" pageNumber="457">
<emphasis id="07D47972FF88805CFF7CF82CDD13F84A" bold="true" box="[151,275,1968,1991]" pageId="4" pageNumber="457">FIGURE 2.</emphasis>
<taxonomicName id="F2A0DEE3FF88805CFEF2F82DDDE5F849" box="[281,485,1969,1990]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="cotopaxi" status="sp. nov.">
<emphasis id="07D47972FF88805CFEF2F82DDDE5F849" box="[281,485,1969,1990]" italics="true" pageId="4" pageNumber="457">Tanytarsus cotopaxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1CE7C409FF88805CFE07F82DDE3CF849" box="[492,572,1969,1990]" pageId="4" pageNumber="457" rank="species">
<emphasis id="07D47972FF88805CFE07F82DDE3CF849" bold="true" box="[492,572,1969,1990]" pageId="4" pageNumber="457">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult male. A—wing, B—hypopygium, C—median volsella (magnified x 2 relative to hypopygium).
</paragraph>
</caption>
<subSubSection id="7DBAF6EBFF89805DFF2CFF0BDF1AFCD0" pageId="5" pageNumber="458" type="discussion">
<paragraph id="351FA560FF89805DFF2CFF0BD8C0FD94" blockId="5.[151,1437,151,755]" pageId="5" pageNumber="458">
<emphasis id="07D47972FF89805DFF2CFF0BDD4CFF3F" bold="true" box="[199,332,151,176]" pageId="5" pageNumber="458">Discussion.</emphasis>
We recognized 
<taxonomicName id="F2A0DEE3FF89805DFDE1FF05DEEEFF3F" box="[522,750,153,176]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="cotopaxi">
<emphasis id="07D47972FF89805DFDE1FF05DEEEFF3F" box="[522,750,153,176]" italics="true" pageId="5" pageNumber="458">Tanytarsus cotopaxi</emphasis>
</taxonomicName>
as a member of the 
<emphasis id="07D47972FF89805DFC0FFF05D83EFF3F" box="[996,1086,153,176]" italics="true" pageId="5" pageNumber="458">riopreto</emphasis>
species group that was so far known from four species (Fittkau &amp; Reiss 1973, Sanseverino 2006). Following the inclusion of 
<taxonomicName id="F2A0DEE3FF89805DFB2EFF21D941FF5B" box="[1221,1345,189,212]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="cotopaxi">
<emphasis id="07D47972FF89805DFB2EFF21D941FF5B" box="[1221,1345,189,212]" italics="true" pageId="5" pageNumber="458">T. cotopaxi</emphasis>
</taxonomicName>
into the group, we propose to delimit the recently emended group diagnosis (Sanseverino 2006) to several principal characters as follows: LR1 relatively high (c. 2.9–3.9); anal tergite bands parallelly tapering to the centre of anal tergite, ending close to anal point base; median setae strong or bristle-shaped; anal point elongate, with crests and spinulae well developed; superior volsella heart-shaped, ellipse-shaped or somewhat T-like, distinctly concave or excavated; two setae on median margin of superior volsella and single seta-bearing tubercle at base of digitus; digitus extending beyond superior volsella; median volsella with setiform and pectinate lamellae; stem of median volsella and digitus more or less of the same length. The 
<emphasis id="07D47972FF89805DFCA2FE20DFA4FE5C" box="[841,932,444,467]" italics="true" pageId="5" pageNumber="458">riopreto</emphasis>
group appears to be closely related to the Holarctic 
<taxonomicName id="F2A0DEE3FF89805DFEEDFE43DD7AFE78" box="[262,378,479,503]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="chinyensis">
<emphasis id="07D47972FF89805DFEEDFE43DD7AFE78" box="[262,378,479,503]" italics="true" pageId="5" pageNumber="458">chinyensis</emphasis>
</taxonomicName>
group, as suggested by Giłka and Paasivirta (2009). The median volsella / digitus proportions and the size of median setae are the best diagnostic characters for separation of the two groups.
</paragraph>
<paragraph id="351FA560FF89805DFF2CFDBBDE39FD7D" blockId="5.[151,1437,151,755]" pageId="5" pageNumber="458">
The adult male of 
<taxonomicName id="F2A0DEE3FF89805DFE77FDB4DE7FFDB0" box="[412,639,552,575]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="cotopaxi">
<emphasis id="07D47972FF89805DFE77FDB4DE7FFDB0" box="[412,639,552,575]" italics="true" pageId="5" pageNumber="458">Tanytarsus cotopaxi</emphasis>
</taxonomicName>
is easily determinable by the low AR (0.48 vs. 0.6–1.0 known in the 
<emphasis id="07D47972FF89805DFF7CFDD0DCF2FDEC" box="[151,242,588,611]" italics="true" pageId="5" pageNumber="458">riopreto</emphasis>
group so far), the bristle-shaped median setae arranged in transversal row, the anal point distinctly expanded in proximal part, bearing large spinulae placed in row, the superior volsella ellipse-shaped, deeply excavated at apex, the digitus long, with swollen apex, and the median volsella with single setiform and three pectinate lamellae. The field of microtrichia on the superior volsella found in 
<taxonomicName id="F2A0DEE3FF89805DFBFCFD24D896FD40" box="[1047,1174,696,719]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="cotopaxi">
<emphasis id="07D47972FF89805DFBFCFD24D896FD40" box="[1047,1174,696,719]" italics="true" pageId="5" pageNumber="458">T. cotopaxi</emphasis>
</taxonomicName>
is known in the group only from 
<taxonomicName id="F2A0DEE3FF89805DFEE6FD40DE39FD7D" authority="Fittkau et Reiss." authorityName="Fittkau et Reiss." box="[269,569,730,755]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="cururui">
<emphasis id="07D47972FF89805DFEE6FD40DD7CFD7C" box="[269,380,732,755]" italics="true" pageId="5" pageNumber="458">T. cururui</emphasis>
Fittkau 
<emphasis id="07D47972FF89805DFE31FD40DDEEFD7C" box="[474,494,732,755]" italics="true" pageId="5" pageNumber="458">et</emphasis>
Reiss.
</taxonomicName>
</paragraph>
<paragraph id="351FA560FF89805DFF7CFCDBDF1AFCD0" blockId="5.[151,794,838,863]" box="[151,794,838,863]" pageId="5" pageNumber="458">
<taxonomicName id="F2A0DEE3FF89805DFF7CFCDBDF1AFCD0" authority="Contreras-Lichtenberg, 1988" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[151,794,838,863]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="curvicristatus">
<emphasis id="07D47972FF89805DFF7CFCDBDF1AFCD0" bold="true" box="[151,794,838,863]" pageId="5" pageNumber="458">
<emphasis id="07D47972FF89805DFF7CFCDBDDB9FCD0" bold="true" box="[151,441,838,863]" italics="true" pageId="5" pageNumber="458">Tanytarsus curvicristatus</emphasis>
Contreras-Lichtenberg, 1988
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7DBAF6EBFF89805DFF7CFC0CDF4CFC4C" pageId="5" pageNumber="458" type="description">
<paragraph id="351FA560FF89805DFF7CFC0CDF4CFC4C" blockId="5.[151,1436,910,963]" pageId="5" pageNumber="458">
<taxonomicName id="F2A0DEE3FF89805DFF7CFC0CDEFDFC2B" authority="Contreras-Lichtenberg, 1988: 101" authorityName="Contreras-Lichtenberg" authorityPageNumber="101" authorityYear="1988" box="[151,765,910,933]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="curvicristatus">
<emphasis id="07D47972FF89805DFF7CFC0CDD98FC2A" box="[151,408,912,933]" italics="true" pageId="5" pageNumber="458">Tanytarsus curvicristatus</emphasis>
Contreras-Lichtenberg, 1988: 101
</taxonomicName>
(adult male; figs 1, 2; 
<collectingCountry id="4DB7E5F0FF89805DFC1BFC12D859FC2B" box="[1008,1113,910,932]" name="Colombia" pageId="5" pageNumber="458">Colombia</collectingCountry>
); Sanseverino 2006: 198 (adult male, fig. 62, 
<collectingCountry id="4DB7E5F0FF89805DFEB8FC32DD86FC4C" box="[339,390,942,963]" name="Peru" pageId="5" pageNumber="458">Peru</collectingCountry>
); Sanseverino &amp; Trivinho-Strixino 2010: 74.
</paragraph>
</subSubSection>
<caption id="61DFF5E8FF89805DFF7CF838DFDEF856" httpUri="https://zenodo.org/record/216760/files/figure.png" pageId="5" pageNumber="458" targetBox="[151,1435,999,1934]" targetPageId="5">
<paragraph id="351FA560FF89805DFF7CF838DFDEF856" blockId="5.[151,1436,1956,2009]" pageId="5" pageNumber="458">
<emphasis id="07D47972FF89805DFF7CF838DD1BF835" bold="true" box="[151,283,1956,1979]" pageId="5" pageNumber="458">FIGURE 3.</emphasis>
<taxonomicName id="F2A0DEE3FF89805DFEC1F83ADF70F835" authority="Contreras-Lichtenberg, 1988" authorityName="Contreras-Lichtenberg" authorityYear="1988" box="[298,880,1956,1979]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="curvicristatus">
<emphasis id="07D47972FF89805DFEC1F83ADE33F834" box="[298,563,1958,1979]" italics="true" pageId="5" pageNumber="458">Tanytarsus curvicristatus</emphasis>
Contreras-Lichtenberg, 1988
</taxonomicName>
, adult male. A—wing, B—hypopygial anal point, C—superior volsellae, D—median volsellae. AC—anal crests, Ba—anal point bar.
</paragraph>
</caption>
<subSubSection id="7DBAF6EBFF8A805EFF7CFF0BDF61FF5D" pageId="6" pageNumber="459" type="materials_examined">
<paragraph id="351FA560FF8A805EFF7CFF0BDF61FF5D" blockId="6.[151,1437,151,736]" pageId="6" pageNumber="459">
<emphasis id="07D47972FF8A805EFF7CFF0BDD84FF3F" bold="true" box="[151,388,151,176]" pageId="6" pageNumber="459">Material examined.</emphasis>
<collectingCountry id="4DB7E5F0FF8A805EFE65FF0BDE19FF20" box="[398,537,151,175]" name="Ecuador" pageId="6" pageNumber="459">ECUADOR</collectingCountry>
, Pichincha province, San Miguel de Los Bancos (canton), Mindo, c. 
<quantity id="F2580885FF8A805EFAAFFF0BD99CFF20" box="[1348,1436,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="459" unit="m" value="1400.0">1400 m</quantity>
a.s.l., 
<date id="411E83A0FF8A805EFF32FF26DDB9FF5D" box="[217,441,186,211]" pageId="6" pageNumber="459" value="1999-08-25" valueMax="1999-08-31" valueMin="1999-08-25">25–31 August 1999</date>
, netting, 
<specimenCount id="23A66EE9FF8A805EFDF4FF26DE6DFF5D" box="[543,621,186,210]" pageId="6" pageNumber="459" type="male">1 male</specimenCount>
, leg. Dawid Graczyk.
</paragraph>
</subSubSection>
<subSubSection id="7DBAF6EBFF8A805EFF2CFF41DF1CFC64" pageId="6" pageNumber="459" type="discussion">
<paragraph id="351FA560FF8A805EFF2CFF41DE04FD17" blockId="6.[151,1437,151,736]" pageId="6" pageNumber="459">
<emphasis id="07D47972FF8A805EFF2CFF41DD4CFF79" bold="true" box="[199,332,221,246]" pageId="6" pageNumber="459">Discussion.</emphasis>
Most of diagnostic characters found in the examined specimen of 
<taxonomicName id="F2A0DEE3FF8A805EFBB8FF42D970FF7A" box="[1107,1392,222,245]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="459" phylum="Arthropoda" rank="species" species="curvicristatus">
<emphasis id="07D47972FF8A805EFBB8FF42D970FF7A" box="[1107,1392,222,245]" italics="true" pageId="6" pageNumber="459">Tanytarsus curvicristatus</emphasis>
</taxonomicName>
are consistent with those originally described and recently complemented (Contreras-Lichtenberg 1988, Sanseverino 2006). The adult male of the species can be identified by the following feature combination: wing covered with macrotrichia only in distal part: sparse macrotrichia in 2/3 distal area of r4+5, m1+2 and along wing margin in cubital and anal cells (
<figureCitation id="AD9BB9E5FF8A805EFEA9FEF6DD85FE0D" box="[322,389,362,386]" captionStart="FIGURE 3" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[151,1435,999,1934]" captionTargetId="figure@5.[151,1436,991,1935]" captionTargetPageId="5" captionText="FIGURE 3. Tanytarsus curvicristatus Contreras-Lichtenberg, 1988, adult male. A—wing, B—hypopygial anal point, C—superior volsellae, D—median volsellae. AC—anal crests, Ba—anal point bar." httpUri="https://zenodo.org/record/216760/files/figure.png" pageId="6" pageNumber="459">Fig. 3</figureCitation>
A); hypopygial anal point with pair of short and broad oval crests above round pit, and with two bars (
<figureCitation id="AD9BB9E5FF8A805EFEFAFE11DD59FE2A" box="[273,345,397,421]" captionStart="FIGURE 3" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[151,1435,999,1934]" captionTargetId="figure@5.[151,1436,991,1935]" captionTargetPageId="5" captionText="FIGURE 3. Tanytarsus curvicristatus Contreras-Lichtenberg, 1988, adult male. A—wing, B—hypopygial anal point, C—superior volsellae, D—median volsellae. AC—anal crests, Ba—anal point bar." httpUri="https://zenodo.org/record/216760/files/figure.png" pageId="6" pageNumber="459">Fig. 3</figureCitation>
B); superior volsella round at base, with protruding posteromedian corner (
<figureCitation id="AD9BB9E5FF8A805EFB33FE11D920FE2A" box="[1240,1312,397,421]" captionStart="FIGURE 3" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[151,1435,999,1934]" captionTargetId="figure@5.[151,1436,991,1935]" captionTargetPageId="5" captionText="FIGURE 3. Tanytarsus curvicristatus Contreras-Lichtenberg, 1988, adult male. A—wing, B—hypopygial anal point, C—superior volsellae, D—median volsellae. AC—anal crests, Ba—anal point bar." httpUri="https://zenodo.org/record/216760/files/figure.png" pageId="6" pageNumber="459">Fig. 3</figureCitation>
C); digitus vestigial, pointed; median volsella short, with setiform, slender spindle-shaped and several ramose lamellae (
<figureCitation id="AD9BB9E5FF8A805EFA84FE2CDCA5FE65" captionStart="FIGURE 3" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[151,1435,999,1934]" captionTargetId="figure@5.[151,1436,991,1935]" captionTargetPageId="5" captionText="FIGURE 3. Tanytarsus curvicristatus Contreras-Lichtenberg, 1988, adult male. A—wing, B—hypopygial anal point, C—superior volsellae, D—median volsellae. AC—anal crests, Ba—anal point bar." httpUri="https://zenodo.org/record/216760/files/figure.png" pageId="6" pageNumber="459">Fig. 3</figureCitation>
D). Other metric and meristic diagnostic characters presently complemented: frontal tubercles absent, AR 0.56, 12 clypeals, 7–8 dorsocentrals, 4–5 scutellars and 1 prealar seta; tibial spur of fore leg 24 μm long, combs of mid and hind legs 12–20 μm long, spurs 24 and 40 μm long (mid leg), 32 and 44 μm long (hind leg); LR2 0.61, LR3 0.64; gonostylus c. 95 μm long, stem of median volsella 8 μm long. In proportion to the smaller size, the individual examined shows a distinctly shorter wing (
<quantity id="F2580885FF8A805EFD92FDC1DEDEFDFA" box="[633,734,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="459" unit="mm" value="1.25">1.25 mm</quantity>
vs. 
<quantity id="F2580885FF8A805EFCE7FDC1DFB4FDFA" box="[780,948,605,629]" metricMagnitude="-3" metricUnit="m" metricValue="1.815" metricValueMax="2.3" metricValueMin="1.33" pageId="6" pageNumber="459" unit="mm" value="1.815" valueMax="2.3" valueMin="1.33">1.33–2.30 mm</quantity>
), antennal and palp segments and a slightly higher LR1 (2.5 vs. 2.4) (
<emphasis id="07D47972FF8A805EFE44FD1CDDCBFD17" box="[431,459,640,664]" italics="true" pageId="6" pageNumber="459">cf.</emphasis>
l.c.).
</paragraph>
<paragraph id="351FA560FF8A805EFF2CFD39DFB5FD6F" blockId="6.[151,1437,151,736]" pageId="6" pageNumber="459">
The systematic position of several 
<taxonomicName id="F2A0DEE3FF8A805EFDA6FD3BDEC7FD31" box="[589,711,679,702]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="07D47972FF8A805EFDA6FD3BDEC7FD31" box="[589,711,679,702]" italics="true" pageId="6" pageNumber="459">Tanytarsus</emphasis>
</taxonomicName>
species having the characteristic structure of the anal point (
<figureCitation id="AD9BB9E5FF8A805EFA85FD39DCA4FD6F" captionStart="FIGURE 3" captionStartId="5.[151,250,1956,1979]" captionTargetBox="[151,1435,999,1934]" captionTargetId="figure@5.[151,1436,991,1935]" captionTargetPageId="5" captionText="FIGURE 3. Tanytarsus curvicristatus Contreras-Lichtenberg, 1988, adult male. A—wing, B—hypopygial anal point, C—superior volsellae, D—median volsellae. AC—anal crests, Ba—anal point bar." httpUri="https://zenodo.org/record/216760/files/figure.png" pageId="6" pageNumber="459">Fig. 3</figureCitation>
B) is discussed in detail by Sanseverino and Trivinho-Strixino (2010).
</paragraph>
<paragraph id="351FA560FF8A805EFF7CFCB9DD7FFCB1" blockId="6.[151,383,805,830]" box="[151,383,805,830]" pageId="6" pageNumber="459">
<emphasis id="07D47972FF8A805EFF7CFCB9DD7FFCB1" bold="true" box="[151,383,805,830]" pageId="6" pageNumber="459">Acknowledgements</emphasis>
</paragraph>
<paragraph id="351FA560FF8A805EFF7CFCF4DF1CFC64" blockId="6.[151,1436,871,1003]" pageId="6" pageNumber="459">
We thank the legator, Mr Dawid Graczyk of Gdynia, 
<collectingCountry id="4DB7E5F0FF8A805EFD00FCFBDF39FCF0" box="[747,825,871,895]" name="Poland" pageId="6" pageNumber="459">Poland</collectingCountry>
, Dr. Angela M. Sanseverino (Universidade Federal do Rio de Janeiro, 
<collectingCountry id="4DB7E5F0FF8A805EFEA3FC17DD90FC2C" box="[328,400,907,931]" name="Brazil" pageId="6" pageNumber="459">Brazil</collectingCountry>
), for supplying information on Neotropical 
<taxonomicName id="F2A0DEE3FF8A805EFC95FC10DFF8FC2C" box="[894,1016,908,931]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="07D47972FF8A805EFC95FC10DFF8FC2C" box="[894,1016,908,931]" italics="true" pageId="6" pageNumber="459">Tanytarsus</emphasis>
</taxonomicName>
. Thanks are directed to Dr. Diego A. Añón Suárez (Centro Regional Universitario Bariloche, 
<collectingCountry id="4DB7E5F0FF8A805EFCFDFC33DF88FC48" box="[790,904,943,967]" name="Argentina" pageId="6" pageNumber="459">Argentina</collectingCountry>
) and Dr. Mariano Donato (Museo de La Plata, 
<collectingCountry id="4DB7E5F0FF8A805EFF7CFC4FDD09FC64" box="[151,265,979,1003]" name="Argentina" pageId="6" pageNumber="459">Argentina</collectingCountry>
) who kindly agreed to evaluate the manuscript.
</paragraph>
</subSubSection>
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<collectingCountry id="4DB7E5F0FF8A805EFB2EFBEED937FB07" box="[1221,1335,1138,1160]" name="Colombia" pageId="6" pageNumber="459">Kolumbien</collectingCountry>
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, 
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, 
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<bibRef id="EC003196FF8A805EFF7CF8EDDE58F829" pageId="6" pageNumber="459" type="journal article">
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&amp; 
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(
<year id="4108E433FF8A805EFDC2F8EEDE5EF807" box="[553,606,1906,1928]" pageId="6" pageNumber="459">2004</year>
) 
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Two new species of 
<emphasis id="07D47972FF8A805EFCB3F8EFDFC4F807" box="[856,964,1907,1928]" italics="true" pageId="6" pageNumber="459">Tanytarsus</emphasis>
from southeast of 
<collectingCountry id="4DB7E5F0FF8A805EFB77F8EDD8DAF808" box="[1180,1242,1905,1927]" name="Brazil" pageId="6" pageNumber="459">Brazil</collectingCountry>
(Insecta, Diptera, Chironomidae)
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, 
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</paragraph>
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<bibRef id="EC003196FF8A805EFF7CF833DEB4F86C" pageId="6" pageNumber="459" type="journal article">
<author id="ED1D4D85FF8A805EFF7CF833DD16F84A" box="[151,278,1967,1989]" fns="I" npo="LF" pageId="6" pageNumber="459">Wirth, W.W.</author>
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(
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) 
<title id="47A44E56FF8A805EFE15F833D90DF84A" box="[510,1293,1967,1989]" pageId="6" pageNumber="459">
A method for mounting small insects on microscope slides in 
<collectingCountry id="4DB7E5F0FF8A805EFB9BF833D8BCF84A" box="[1136,1212,1967,1989]" name="Canada" pageId="6" pageNumber="459">Canada</collectingCountry>
balsam
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. 
<journalOrPublisher id="EF61AFACFF8A805EFAFDF833DE22F86C" pageId="6" pageNumber="459">
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Annals of the Entomological Society of 
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</paragraph>
</subSubSection>
</treatment>
</document>