treatments-xml/data/AF/6A/87/AF6A87D543766151FF6ADFFA4B0C3BFC.xml
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<document id="C702A78223E80F834593FA9ABB132461" ID-CLB-Dataset="22814" ID-DOI="10.11646/zootaxa.4766.4.1" ID-GBIF-Dataset="1e2996e2-e31f-45e3-9149-60ebce934708" ID-ISSN="1175-5334" ID-Zenodo-Dep="3765652" ID-ZooBank="urn:lsid:zoobank.org:pub:2F8FEDC1-6891-46D1-B372-858CDCBE4051" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587754498722" checkinUser="carolina" docAuthor="Kompier, Tom, Dow, Rory A. &amp; Steinhoff, Philip O. M." docDate="2020" docId="AF6A87D543766151FF6ADFFA4B0C3BFC" docLanguage="en" docName="zootaxa.4766.4.1.pdf.imf" docOrigin="Zootaxa 4766 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia pulchella Kompier &amp; Dow &amp; Steinhoff 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="534" masterDocId="5353FFAD436C6170FFFDDB7048083A53" masterDocTitle="Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus" masterLastPageNumber="538" masterPageNumber="501" pageNumber="527" updateTime="1698835840532" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E44BFE805E8A3EAB98AD7FB511177D19">Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus</mods:title>
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<mods:namePart id="8DEC9D5973D7F3FD000BC914B32171FC">Kompier, Tom</mods:namePart>
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<mods:namePart id="69B44353DDBFCE403FB6B2231BF854AE">Dow, Rory A.</mods:namePart>
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<mods:namePart id="C8CEB173C8DF11F71252C80AD60D561F">Steinhoff, Philip O. M.</mods:namePart>
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<mods:date id="FD86D8E25042F81F6DF22B23AA0A0D8B">2020</mods:date>
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<treatment id="AF6A87D543766151FF6ADFFA4B0C3BFC" ID-DOI="http://doi.org/10.5281/zenodo.3803430" ID-GBIF-Taxon="163739035" ID-Zenodo-Dep="3803430" LSID="urn:lsid:plazi:treatment:AF6A87D543766151FF6ADFFA4B0C3BFC" httpUri="http://treatment.plazi.org/id/AF6A87D543766151FF6ADFFA4B0C3BFC" lastPageId="33" lastPageNumber="534" pageId="26" pageNumber="527">
<subSubSection id="6FD965484376616AFF6ADFFA4A0A3EF7" box="[151,514,1161,1188]" pageId="26" pageNumber="527" type="nomenclature">
<paragraph id="277C36C34376616AFF6ADFFA4A0A3EF7" blockId="26.[151,812,1161,1224]" box="[151,514,1161,1188]" pageId="26" pageNumber="527">
<heading id="7C3481AF4376616AFF6ADFFA4A0A3EF7" bold="true" box="[151,514,1161,1188]" fontSize="10" level="2" pageId="26" pageNumber="527" reason="2">
<emphasis id="15B7EAD14376616AFF6ADFFA4A0A3EF7" bold="true" box="[151,514,1161,1188]" pageId="26" pageNumber="527">
8.
<taxonomicName id="E0C34D404376616AFF48DFF949943EF7" ID-CoL="9X2BX" authority="Kompier &amp; Dow &amp; Steinhoff, 2020" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[181,412,1161,1188]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="pulchella" status="sp. nov.">
<emphasis id="15B7EAD14376616AFF48DFF949943EF7" bold="true" box="[181,412,1161,1188]" italics="true" pageId="26" pageNumber="527">Coeliccia pulchella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E8457AA4376616AFE59DFFA4A0A3EF7" box="[420,514,1162,1188]" pageId="26" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="6FD965484376616AFF6ADFDD4B243E9B" box="[151,812,1197,1224]" pageId="26" pageNumber="527" type="description">
<paragraph id="277C36C34376616AFF6ADFDD4B243E9B" blockId="26.[151,812,1161,1224]" box="[151,812,1197,1224]" pageId="26" pageNumber="527">
(
<figureCitation id="BFF82A464376616AFF62DFDD48E03E94" box="[159,232,1197,1223]" captionStart="FIGURE 6" captionStartId="8.[151,250,1923,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure@8.[151,1436,993,1898]" captionTargetPageId="8" captionText="FIGURE 6. Holotype of Coeliccia pulchella (a) thorax in lateral view; (b) thorax in dorsal view; (c) head in dorsolateral view; (d) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765664" httpUri="https://zenodo.org/record/3765664/files/figure.png" pageId="26" pageNumber="527">Figs 6</figureCitation>
,
<figureCitation id="BFF82A464376616AFF09DFDD49093E94" box="[244,257,1197,1223]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="26" pageNumber="527">7</figureCitation>
cd, 8gh, 11cd, 14e, 16e, 17e, 19e, 21b, 27df)
</paragraph>
</subSubSection>
<subSubSection id="6FD965484376616AFF6ADF864C7E3C58" pageId="26" pageNumber="527" type="materials_examined">
<paragraph id="277C36C34376616AFF6ADF864D4C3F5C" blockId="26.[151,1437,1269,2016]" box="[151,1348,1269,1296]" pageId="26" pageNumber="527">
<materialsCitation id="97AB3C9E4376616AFF6ADF864D4C3F5C" ID-GBIF-Occurrence="2608670279" box="[151,1348,1269,1296]" collectingDate="2016-07-23" collectorName="Huu Lien NR" elevation="250" latitude="21.662" location="Lang Son Prov." longLatPrecision="75" longitude="106.373" pageId="26" pageNumber="527" specimenCount="1" specimenCount-male="1" stateProvince="Lang Son Prov." typeStatus="holotype">
<emphasis id="15B7EAD14376616AFF6ADF86490C3F43" bold="true" box="[151,260,1270,1296]" pageId="26" pageNumber="527">
<typeStatus id="F87888614376616AFF6ADF86490C3F43" box="[151,260,1270,1296]" pageId="26" pageNumber="527" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="31C5FD4A4376616AFEF6DF8649283F43" box="[267,288,1270,1296]" pageId="26" pageNumber="527" type="male"></specimenCount>
<collectorName id="8A3653154376616AFEDADF8649CB3F43" box="[295,451,1269,1296]" pageId="26" pageNumber="527">Huu Lien NR</collectorName>
,
<collectingRegion id="E507F8214376616AFE30DF864A733F43" box="[461,635,1270,1296]" country="Vietnam" name="Lang Son" pageId="26" pageNumber="527">Lang Son Prov.</collectingRegion>
(appr.
<geoCoordinate id="42F750044376616AFD34DF854B263F5C" box="[713,814,1269,1295]" degrees="21.662" direction="north" orientation="latitude" pageId="26" pageNumber="527" precision="55" value="21.662">21.662N</geoCoordinate>
,
<geoCoordinate id="42F750044376616AFCC5DF854BA03F5C" box="[824,936,1269,1295]" degrees="106.373" direction="east" orientation="longitude" pageId="26" pageNumber="527" precision="55" value="106.373">106.373E</geoCoordinate>
; alt.
<elevation id="ACEED1F04376616AFC20DF854C463F43" box="[989,1102,1269,1296]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="527" unit="m" value="250.0">
<quantity id="E03B9B264376616AFC20DF854C2C3F43" box="[989,1060,1269,1296]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="527" unit="m" value="250.0">250 m</quantity>
asl
</elevation>
.)
<date id="537D10034376616AFB9FDF864CEC3F5C" box="[1122,1252,1269,1296]" pageId="26" pageNumber="527" value="2016-07-23">
<collectingDate id="4339E9EB4376616AFB9FDF864CEC3F5C" box="[1122,1252,1269,1296]" pageId="26" pageNumber="527" value="2016-07-23">23 vii 2016</collectingDate>
</date>
, TK leg.
</materialsCitation>
</paragraph>
<paragraph id="277C36C34376616AFF3ADE6A49B23F28" blockId="26.[151,1437,1269,2016]" pageId="26" pageNumber="527">
<emphasis id="15B7EAD14376616AFF3ADE6A49373F67" bold="true" box="[199,319,1306,1332]" pageId="26" pageNumber="527">
<typeStatus id="F87888614376616AFF3ADE6A49373F67" box="[199,319,1306,1332]" pageId="26" pageNumber="527" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="31C5FD4A4376616AFEB8DE6A49783F67" box="[325,368,1306,1332]" pageId="26" pageNumber="527" type="male">1 ♂</specimenCount>
,
<collectorName id="8A3653154376616AFE86DE6A4A1D3F67" box="[379,533,1305,1332]" pageId="26" pageNumber="527">Huu Lien NR</collectorName>
,
<collectingRegion id="E507F8214376616AFDE2DE6A4AC63F67" box="[543,718,1306,1332]" country="Vietnam" name="Lang Son" pageId="26" pageNumber="527">Lang Son Prov.</collectingRegion>
,
<date id="537D10034376616AFD2ADE6A4B403F67" box="[727,840,1305,1332]" pageId="26" pageNumber="527" value="2015-10-11">
<collectingDate id="4339E9EB4376616AFD2ADE6A4B403F67" box="[727,840,1305,1332]" pageId="26" pageNumber="527" value="2015-10-11">11 x 2015</collectingDate>
</date>
, TK leg.;
<specimenCount id="31C5FD4A4376616AFC49DE6A4BD73F67" box="[948,991,1306,1332]" pageId="26" pageNumber="527" type="male">1 ♂</specimenCount>
, same location and collector,
<date id="537D10034376616AFADBDE694D9F3F67" box="[1318,1431,1305,1332]" pageId="26" pageNumber="527" value="2015-10-25">
<collectingDate id="4339E9EB4376616AFADBDE694D9F3F67" box="[1318,1431,1305,1332]" pageId="26" pageNumber="527" value="2015-10-25">25 x 2015</collectingDate>
</date>
;
<specimenCount id="31C5FD4A4376616AFF6ADE4E48CA3F0B" box="[151,194,1342,1368]" pageId="26" pageNumber="527" type="female">1 ♀</specimenCount>
, same location, date and collector as
<typeStatus id="F87888614376616AFD90DE4D4AC73F04" box="[621,719,1341,1367]" pageId="26" pageNumber="527" type="holotype">holotype</typeStatus>
(used for description);
<specimenCount id="31C5FD4A4376616AFC27DE4E4C143F0B" box="[986,1052,1342,1368]" pageId="26" pageNumber="527" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="31C5FD4A4376616AFBD4DE4E4C5C3F0B" box="[1065,1108,1342,1368]" pageId="26" pageNumber="527" type="female">1 ♀</specimenCount>
,
<collectorName id="8A3653154376616AFB9EDE4E4D123F0B" box="[1123,1306,1341,1368]" pageId="26" pageNumber="527">Cuc Phuong NP</collectorName>
,
<collectingRegion id="E507F8214376616AFADBDE4D48DD3F2F" country="Vietnam" name="Ninh Binh" pageId="26" pageNumber="527">Ninh Binh Prov.</collectingRegion>
,
<date id="537D10034376616AFF22DE1249523F28" box="[223,346,1377,1404]" pageId="26" pageNumber="527" value="2016-09-18">
<collectingDate id="4339E9EB4376616AFF22DE1249523F28" box="[223,346,1377,1404]" pageId="26" pageNumber="527" value="2016-09-18">18 ix 2016</collectingDate>
</date>
, TK leg.
</paragraph>
<paragraph id="277C36C34376616AFF3ADEF54C7E3C58" blockId="26.[151,1437,1269,2016]" pageId="26" pageNumber="527">
<emphasis id="15B7EAD14376616AFF3ADEF54AEF3FF3" bold="true" box="[199,743,1413,1440]" pageId="26" pageNumber="527">Other material not examined by the authors.</emphasis>
<specimenCount id="31C5FD4A4376616AFD0DDEF64B3A3FF3" box="[752,818,1414,1440]" pageId="26" pageNumber="527" type="male">2 ♂♂</specimenCount>
, Huu Lien NR,
<collectingRegion id="E507F8214376616AFC16DEF64C973FF3" box="[1003,1183,1414,1440]" country="Vietnam" name="Lang Son" pageId="26" pageNumber="527">Lang Son Prov.</collectingRegion>
,
<date id="537D10034376616AFB51DEF64D2A3FF3" box="[1196,1314,1413,1440]" pageId="26" pageNumber="527" value="2015-05-21">21 v 2015</date>
, QTP leg.;
<specimenCount id="31C5FD4A4376616AFF6ADEDA48D13F97" box="[151,217,1450,1476]" pageId="26" pageNumber="527" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="31C5FD4A4376616AFF1BDEDA492E3F97" box="[230,294,1450,1476]" pageId="26" pageNumber="527" type="female">2 ♀♀</specimenCount>
, Cuc Phuong NP,
<collectingRegion id="E507F8214376616AFE0BDED94AB93F97" box="[502,689,1449,1476]" country="Vietnam" name="Ninh Binh" pageId="26" pageNumber="527">Ninh Binh Prov.</collectingRegion>
,
<date id="537D10034376616AFD40DED94B313F97" box="[701,825,1449,1476]" pageId="26" pageNumber="527" value="2018-06-25">25 vi 2018</date>
, QTP leg.;
<specimenCount id="31C5FD4A4376616AFC44DEDA4BEE3F97" box="[953,998,1450,1476]" pageId="26" pageNumber="527" type="male">1 ♂</specimenCount>
,
<specimenCount id="31C5FD4A4376616AFC0EDEDA4C153F97" box="[1011,1053,1450,1476]" pageId="26" pageNumber="527" type="female">1 ♀</specimenCount>
, Son Kim Commune, Huong Son District,
<collectingRegion id="E507F8214376616AFF04DEBE49913FBB" box="[249,409,1485,1512]" country="Vietnam" name="Ha Tinh" pageId="26" pageNumber="527">Ha Tinh Prov.</collectingRegion>
,
<date id="537D10034376616AFE5EDEBD4A2E3FBB" box="[419,550,1485,1512]" pageId="26" pageNumber="527" value="2018-07-05">05 vii 2018</date>
, QTP leg.;
<specimenCount id="31C5FD4A4376616AFD59DEBE4AED3FBB" box="[676,741,1486,1512]" pageId="26" pageNumber="527" type="male">2 ♂♂</specimenCount>
, Phu Gia Commune, Huong Khe District,
<collectingRegion id="E507F8214376616AFB3CDEBE4D683FBB" box="[1217,1376,1485,1512]" country="Vietnam" name="Ha Tinh" pageId="26" pageNumber="527">Ha Tinh Prov.</collectingRegion>
,
<date id="537D10034376616AFA96DEBE48DA3C5F" pageId="26" pageNumber="527" value="2019-05-10">10 v 2019</date>
, QTP leg.;
<specimenCount id="31C5FD4A4376616AFEADDE8249743C5F" box="[336,380,1522,1548]" pageId="26" pageNumber="527" type="male">1 ♂</specimenCount>
, Truing Xuan,
<collectingRegion id="E507F8214376616AFDD9DE824AF93C5F" box="[548,753,1521,1548]" country="Vietnam" name="Quang Binh" pageId="26" pageNumber="527">Quang Binh Prov.</collectingRegion>
,
<date id="537D10034376616AFD01DE824B763C5F" box="[764,894,1521,1548]" pageId="26" pageNumber="527" value="2019-07-30">30 vii 2019</date>
, Do Manh Cuong leg.
</paragraph>
</subSubSection>
<subSubSection id="6FD965484376616AFF3ADD6649B83C00" pageId="26" pageNumber="527" type="etymology">
<paragraph id="277C36C34376616AFF3ADD6649B83C00" blockId="26.[151,1437,1269,2016]" pageId="26" pageNumber="527">
<emphasis id="15B7EAD14376616AFF3ADD6649403C63" bold="true" box="[199,328,1558,1584]" pageId="26" pageNumber="527">Etymology</emphasis>
. The adjective
<taxonomicName id="E0C34D404376616AFE0BDD664A563C63" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[502,606,1558,1584]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="15B7EAD14376616AFE0BDD664A563C63" box="[502,606,1558,1584]" italics="true" pageId="26" pageNumber="527">pulchella</emphasis>
</taxonomicName>
means “beautiful little” in Latin and refers to the small size and beautiful coloration of the species.
</paragraph>
</subSubSection>
<subSubSection id="6FD965484376616FFF3ADD2D4BAA3B34" lastPageId="31" lastPageNumber="532" pageId="26" pageNumber="527" type="description">
<paragraph id="277C36C34376616AFF3ADD2D4C9B3D78" blockId="26.[151,1437,1269,2016]" pageId="26" pageNumber="527">
<emphasis id="15B7EAD14376616AFF3ADD2D49D43C2B" bold="true" box="[199,476,1629,1656]" pageId="26" pageNumber="527">
Description of
<typeStatus id="F87888614376616AFE88DD2E49D43C2B" box="[373,476,1630,1656]" pageId="26" pageNumber="527" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="BFF82A464376616AFE17DD2D4A393C24" box="[490,561,1629,1655]" captionStart="FIGURE 6" captionStartId="8.[151,250,1923,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure@8.[151,1436,993,1898]" captionTargetPageId="8" captionText="FIGURE 6. Holotype of Coeliccia pulchella (a) thorax in lateral view; (b) thorax in dorsal view; (c) head in dorsolateral view; (d) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765664" httpUri="https://zenodo.org/record/3765664/files/figure.png" pageId="26" pageNumber="527">Figs 6</figureCitation>
,
<figureCitation id="BFF82A464376616AFDC0DD2D4A423C24" box="[573,586,1629,1655]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="26" pageNumber="527">7</figureCitation>
cd, 8gh, 11cd).
<emphasis id="15B7EAD14376616AFCE2DD2E4B543C2B" box="[799,860,1630,1656]" italics="true" pageId="26" pageNumber="527">Head</emphasis>
(
<figureCitation id="BFF82A464376616AFC97DD2D4BB33C2B" box="[874,955,1629,1656]" captionStart="FIGURE 6" captionStartId="8.[151,250,1923,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure@8.[151,1436,993,1898]" captionTargetPageId="8" captionText="FIGURE 6. Holotype of Coeliccia pulchella (a) thorax in lateral view; (b) thorax in dorsal view; (c) head in dorsolateral view; (d) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765664" httpUri="https://zenodo.org/record/3765664/files/figure.png" pageId="26" pageNumber="527">Fig. 6c</figureCitation>
). Labrum and postclypeus shiny black. Anteclypeus yellow, with two darker areas left and right on lower half. Genae and base of mandibles yellow, the yellow narrowly continuing over the antefrons above the postclypeus for about one-fourth in of the width of the antefrons, remainder of frons matte black. Dorsal surface of head matte black, apart from two small yellow spots between base of antennae and lateral ocelli and two oblong pale yellow postocular spots. Antennae black, top of first segment and base of second segment pale yellow. Compound eyes in life black dorsally, green ventrally.
</paragraph>
<paragraph id="277C36C34376616AFF3ADC464CB43D8C" blockId="26.[151,1437,1269,2016]" pageId="26" pageNumber="527">
<emphasis id="15B7EAD14376616AFF3ADC4649103D03" box="[199,280,1846,1872]" italics="true" pageId="26" pageNumber="527">Thorax</emphasis>
. Prothorax (
<figureCitation id="BFF82A464376616AFE55DC4549F73D03" box="[424,511,1845,1872]" captionStart="FIGURE 6" captionStartId="8.[151,250,1923,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure@8.[151,1436,993,1898]" captionTargetPageId="8" captionText="FIGURE 6. Holotype of Coeliccia pulchella (a) thorax in lateral view; (b) thorax in dorsal view; (c) head in dorsolateral view; (d) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765664" httpUri="https://zenodo.org/record/3765664/files/figure.png" pageId="26" pageNumber="527">Fig. 6d</figureCitation>
): matte black, marked with brownish yellow lines, propleuron bright yellow, but posterior third brownish black. Synthorax (
<figureCitation id="BFF82A464376616AFD8ADC294AB03D27" box="[631,696,1881,1908]" captionStart="FIGURE 6" captionStartId="8.[151,250,1923,1948]" captionTargetBox="[151,1436,993,1898]" captionTargetId="figure@8.[151,1436,993,1898]" captionTargetPageId="8" captionText="FIGURE 6. Holotype of Coeliccia pulchella (a) thorax in lateral view; (b) thorax in dorsal view; (c) head in dorsolateral view; (d) prothorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3765664" httpUri="https://zenodo.org/record/3765664/files/figure.png" pageId="26" pageNumber="527">Fig. 6</figureCitation>
ab): mesepisternum black, but with a yellow, almost square, spot, on one side bordered by mesopleural suture, starting at about one-seventh length of mesepisternum and extending to about three-sevenths. Mesepimeron and mesinfraepisternum black. Metepisternum and metepimeron yellow. Legs: coxae and trochanters yellow. Femur pale yellow with black lines. Claws and spines brownish.
</paragraph>
<paragraph id="277C36C34377616BFF3ADBEA4C1D3AE0" blockId="27.[151,1437,153,324]" box="[199,1045,153,179]" pageId="27" pageNumber="528">
<emphasis id="15B7EAD14377616BFF3ADBEA49033AE0" box="[199,267,154,179]" italics="true" pageId="27" pageNumber="528">Wings</emphasis>
. Hyaline, 18 Px in FW,
<quantity id="E03B9B264377616BFDE7DBE94A5B3AE0" box="[538,595,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="27" pageNumber="528" unit="in" value="16.0">16 in</quantity>
HW. Pt dark brown, covering 12 cells.
</paragraph>
<paragraph id="277C36C34377616BFF3ADBCD4AFD3B17" blockId="27.[151,1437,153,324]" pageId="27" pageNumber="528">
<emphasis id="15B7EAD14377616BFF3ADBCD49313A84" box="[199,313,189,215]" italics="true" pageId="27" pageNumber="528">Abdomen.</emphasis>
S1 yellow with posterior margin black; S2 black dorsally and yellow below, black along anterior and posterior margin; S36 black dorsally and somewhat paler below, and with prominent subapical annulus; S7 black shading to blackish brown posteriorly and with yellow apical lateral spot; S8 anterior two-thirds brownish, posterior one-third brownish yellow; S910 yellow (
<figureCitation id="BFF82A464377616BFD84DA594AB43B17" box="[633,700,297,324]" captionStart="FIGURE 7" captionStartId="9.[151,250,1813,1838]" captionTargetBox="[151,1436,181,1786]" captionTargetId="figure@9.[151,1436,181,1787]" captionTargetPageId="9" captionText="FIGURE 7. Abdomen tip of male Coeliccia spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b) C. caerulea, holotype; (c,d) C. pulchella, holotype; (e,f) C. curua, holotype; (g,h) C. coronata, holotype; (i,j) C. diomedea, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="27" pageNumber="528">Fig. 7</figureCitation>
cd).
</paragraph>
<caption id="73BC664B4377616BFF6ADE924C1C3C4C" ID-DOI="http://doi.org/10.5281/zenodo.3765696" ID-Zenodo-Dep="3765696" httpUri="https://zenodo.org/record/3765696/files/figure.png" pageId="27" pageNumber="528" startId="27.[151,250,1506,1531]" targetBox="[157,1419,345,1481]" targetPageId="27">
<paragraph id="277C36C34377616BFF6ADE924C1C3C4C" blockId="27.[151,1436,1506,1567]" pageId="27" pageNumber="528">
<emphasis id="15B7EAD14377616BFF6ADE9249293FA8" bold="true" box="[151,289,1506,1531]" pageId="27" pageNumber="528">FIGURE 22.</emphasis>
Abdomen tip of female
<taxonomicName id="E0C34D404377616BFDE1DE934A753FA8" authorityName="Kirby" authorityYear="1890" box="[540,637,1507,1531]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD14377616BFDE1DE934A753FA8" box="[540,637,1507,1531]" italics="true" pageId="27" pageNumber="528">Coeliccia</emphasis>
</taxonomicName>
spp. used for description in lateral view. (a)
<taxonomicName id="E0C34D404377616BFBBBDE934CB73FA8" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1094,1215,1507,1531]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="coronata">
<emphasis id="15B7EAD14377616BFBBBDE934CB73FA8" box="[1094,1215,1507,1531]" italics="true" pageId="27" pageNumber="528">C. coronata</emphasis>
</taxonomicName>
; (b)
<taxonomicName id="E0C34D404377616BFB0CDE934D7A3FA8" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1265,1394,1507,1531]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="diomedea">
<emphasis id="15B7EAD14377616BFB0CDE934D7A3FA8" box="[1265,1394,1507,1531]" italics="true" pageId="27" pageNumber="528">C. diomedea</emphasis>
</taxonomicName>
; (c)
<taxonomicName id="E0C34D404377616BFF6ADD7749033C4C" authorityName="Phan &amp; Kompier" authorityYear="2016" box="[151,267,1543,1567]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="15B7EAD14377616BFF6ADD7749033C4C" box="[151,267,1543,1567]" italics="true" pageId="27" pageNumber="528">C. hayashii</emphasis>
</taxonomicName>
; (d)
<taxonomicName id="E0C34D404377616BFEC1DD7749963C4D" authorityName="Dow" authorityYear="2016" box="[316,414,1543,1566]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="suoitia">
<emphasis id="15B7EAD14377616BFEC1DD7749963C4D" box="[316,414,1543,1566]" italics="true" pageId="27" pageNumber="528">C. suoitia</emphasis>
</taxonomicName>
, covered in dirt; (e)
<taxonomicName id="E0C34D404377616BFD92DD774AD13C4C" authorityName="Wilson &amp; Reels" authorityYear="2003" box="[623,729,1543,1567]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="galbina">
<emphasis id="15B7EAD14377616BFD92DD774AD13C4C" box="[623,729,1543,1567]" italics="true" pageId="27" pageNumber="528">C. galbina</emphasis>
</taxonomicName>
; (f)
<taxonomicName id="E0C34D404377616BFCFADD774B753C4C" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[775,893,1543,1567]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD14377616BFCFADD774B753C4C" box="[775,893,1543,1567]" italics="true" pageId="27" pageNumber="528">C. caerulea</emphasis>
</taxonomicName>
, alive in hand.
</paragraph>
</caption>
<paragraph id="277C36C34377616BFF3ADD3D4B843C87" blockId="27.[151,1437,1613,2036]" pageId="27" pageNumber="528">
<emphasis id="15B7EAD14377616BFF3ADD3D49823C34" box="[199,394,1613,1639]" italics="true" pageId="27" pageNumber="528">Anal appendages</emphasis>
(
<figureCitation id="BFF82A464377616BFE64DD3D49D33C3B" box="[409,475,1613,1640]" captionStart="FIGURE 8" captionStartId="10.[151,250,1840,1865]" captionTargetBox="[166,1421,181,1813]" captionTargetId="figure@10.[166,1421,181,1814]" captionTargetPageId="10" captionText="FIGURE 8. S10 and caudal appendages of male Coeliccia spp. [a,c,e,] lateral view, [g] dorsolateral view, [b,d,f,h] dorsal view. (a,b) C. coronata, holotype; (c,d) C. diomedea, holotype; (e,f) C. caerulea, holotype; (g,h) C. pulchella, holotype." figureDoi="http://doi.org/10.5281/zenodo.3765668" httpUri="https://zenodo.org/record/3765668/files/figure.png" pageId="27" pageNumber="528">Fig. 8</figureCitation>
gh). Yellow. In lateral view the cerci somewhat club-shaped, thinner at the proximal end, and only a little shorter than paraprocts, and with subapical ventral expansion, tipped with black tooth. In dorsal view a small tooth at one-third of length of cerci, from where the subapical ventral expansion starts. The paraprocts of usual
<typeStatus id="F87888614377616BFF05DDCA49233C87" box="[248,299,1722,1748]" pageId="27" pageNumber="528">type</typeStatus>
, apically curved inwards and with black tooth at apex.
</paragraph>
<paragraph id="277C36C34377616BFF3ADDAD4BD93D4F" blockId="27.[151,1437,1613,2036]" pageId="27" pageNumber="528">
<emphasis id="15B7EAD14377616BFF3ADDAD496C3CA4" box="[199,356,1757,1783]" italics="true" pageId="27" pageNumber="528">Genital ligula</emphasis>
(
<figureCitation id="BFF82A464377616BFE88DDAD49CC3CAB" box="[373,452,1757,1784]" captionStart="FIGURE 11" captionStartId="16.[151,250,1577,1602]" captionTargetBox="[158,1416,166,1551]" captionTargetId="figure@16.[158,1416,166,1551]" captionTargetPageId="16" captionText="FIGURE 11. Genital ligula of Coeliccia spp. [a,c,e,g] ventral view, [b,d,f,h] lateral view. (a,b) C. diomedea, holotype; (c,d) C. pulchella, holotype; (e,f) C. coronata, paratype, Da Lat, Lam Dong Prov., 16 iv 2016; (g,h) C. caerulea, paratype, flagellum missing on one side." figureDoi="http://doi.org/10.5281/zenodo.3765674" httpUri="https://zenodo.org/record/3765674/files/figure.png" pageId="27" pageNumber="528">Fig. 11</figureCitation>
cd). The apical segment thin and flat, forming a tongue, with two thin curved flagella originating laterally close to its base. Internal fold and apical flap present.
</paragraph>
<paragraph id="277C36C34377616BFF3ADC564B853D6C" blockId="27.[151,1437,1613,2036]" box="[199,909,1829,1856]" pageId="27" pageNumber="528">Measurements (in mm). HW 23; abdomen incl. appendages 36.</paragraph>
<paragraph id="277C36C34377616BFF3ADC3848EB3D83" blockId="27.[151,1437,1613,2036]" pageId="27" pageNumber="528">
<emphasis id="15B7EAD14377616BFF3ADC384A103D30" bold="true" box="[199,536,1864,1891]" pageId="27" pageNumber="528">
Variation in male
<typeStatus id="F87888614377616BFE67DC394A1A3D30" box="[410,530,1865,1891]" pageId="27" pageNumber="528" type="paratype">paratypes</typeStatus>
.
</emphasis>
Some variability in extent of yellow on middle pronotal lobe of prothorax, sometimes almost completely yellow. Px in FW 1718, in HW 1516. Yellow spots on mesepisternum sometimes slightly larger, covering about one-third of length. One male with thin black line over posterior two-thirds of metapleural suture.
</paragraph>
<paragraph id="277C36C34377616BFF3ADCAA4BCC3DA0" blockId="27.[151,1437,1613,2036]" box="[199,964,2008,2036]" pageId="27" pageNumber="528">Measurements (mm). HW 2225; abdomen incl. appendages 3639.</paragraph>
<caption id="73BC664B4370616CFF6ADC1549563D95" ID-DOI="http://doi.org/10.5281/zenodo.3765698" ID-Zenodo-Dep="3765698" httpUri="https://zenodo.org/record/3765698/files/figure.png" pageId="28" pageNumber="529" startId="28.[151,250,1893,1918]" targetBox="[151,1436,181,1868]" targetPageId="28">
<paragraph id="277C36C34370616CFF6ADC1549563D95" blockId="28.[151,1437,1893,1990]" pageId="28" pageNumber="529">
<emphasis id="15B7EAD14370616CFF6ADC1549173D2D" bold="true" box="[151,287,1893,1918]" pageId="28" pageNumber="529">FIGURE 23.</emphasis>
Holotype of
<taxonomicName id="E0C34D404370616CFE5FDC164AEE3D2D" authority="Fraser, 1932" authorityName="Fraser" authorityYear="1932" box="[418,742,1894,1918]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="15B7EAD14370616CFE5FDC164A0B3D2D" box="[418,515,1894,1918]" italics="true" pageId="28" pageNumber="529">Coeliccia</emphasis>
<emphasis id="15B7EAD14370616CFDF5DC164A693D2E" box="[520,609,1894,1917]" italics="true" pageId="28" pageNumber="529">montana</emphasis>
Fraser, 1932
</taxonomicName>
. Photos courtesy of Dr. Ben Price, Natural History Museum, London. (a) thorax and head in lateral view; (b) abdomen tip and caudal appendages in lateral view; (c) thorax in dorsal view; (d) type on card with labels.
</paragraph>
</caption>
<caption id="73BC664B4371616DFF6ADEA54BEA3C09" ID-DOI="http://doi.org/10.5281/zenodo.3765700" ID-Zenodo-Dep="3765700" httpUri="https://zenodo.org/record/3765700/files/figure.png" pageId="29" pageNumber="530" startId="29.[151,250,1493,1518]" targetBox="[151,1436,181,1468]" targetPageId="29">
<paragraph id="277C36C34371616DFF6ADEA54BEA3C09" blockId="29.[151,1437,1493,1626]" pageId="29" pageNumber="530">
<emphasis id="15B7EAD14371616DFF6ADEA549293FBD" bold="true" box="[151,289,1493,1518]" pageId="29" pageNumber="530">FIGURE 24.</emphasis>
Thorax and head alive in hand of
<taxonomicName id="E0C34D404371616DFD7ADEA64AE03FBD" authorityName="Kirby" authorityYear="1890" box="[647,744,1494,1518]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD14371616DFD7ADEA64AE03FBD" box="[647,744,1494,1518]" italics="true" pageId="29" pageNumber="530">Coeliccia</emphasis>
</taxonomicName>
spp. [a,b]
<taxonomicName id="E0C34D404371616DFCA5DEA64BC73FBD" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[856,975,1494,1518]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD14371616DFCA5DEA64BC73FBD" box="[856,975,1494,1518]" italics="true" pageId="29" pageNumber="530">C. caerulea</emphasis>
</taxonomicName>
, [c,d]
<taxonomicName id="E0C34D404371616DFBE9DEA64C863FBD" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1044,1166,1494,1518]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="coronata">
<emphasis id="15B7EAD14371616DFBE9DEA64C863FBD" box="[1044,1166,1494,1518]" italics="true" pageId="29" pageNumber="530">C. coronata</emphasis>
</taxonomicName>
, [e,f]
<taxonomicName id="E0C34D404371616DFB33DEA64D463FBD" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1230,1358,1494,1518]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="530" phylum="Arthropoda" rank="species" species="diomedea">
<emphasis id="15B7EAD14371616DFB33DEA64D463FBD" box="[1230,1358,1494,1518]" italics="true" pageId="29" pageNumber="530">C. diomedea</emphasis>
</taxonomicName>
, [a,c,e] male, [b,d,f] female. (a) holotype in lateral view; (b) female used for description in dorsolateral view; (c) paratype in dorsolateral view, Da Lat, Lam Dong Prov., 16 iv 2016; (d) female used for description in lateral view; (e) dorsolateral view, Quang Nam Prov., along QL14, 26 ix 2015; (f) female used for description in dorsolateral view.
</paragraph>
</caption>
<paragraph id="277C36C34371616DFF3ADDF74C9B3D05" blockId="29.[151,1437,1671,2023]" pageId="29" pageNumber="530">
<emphasis id="15B7EAD14371616DFF3ADDF74A333CF1" bold="true" box="[199,571,1671,1698]" pageId="29" pageNumber="530">
Description of female
<typeStatus id="F87888614371616DFE2DDDF84A333CF1" box="[464,571,1672,1698]" pageId="29" pageNumber="530" type="paratype">paratype</typeStatus>
</emphasis>
(
<figureCitation id="BFF82A464371616DFDB1DDF84AB93CF1" box="[588,689,1672,1698]" captionStart="FIGURE 14" captionStartId="19.[151,250,1649,1674]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure@19.[151,1436,181,1623]" captionTargetPageId="19" captionText="FIGURE 14. Heads of female Coeliccia ssp. used for description in dorsolateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765680" httpUri="https://zenodo.org/record/3765680/files/figure.png" pageId="29" pageNumber="530">Figs 14e</figureCitation>
,
<figureCitation id="BFF82A464371616DFD40DDF84AE03CF1" box="[701,744,1672,1698]" captionStart="FIGURE 16" captionStartId="21.[151,250,1653,1678]" captionTargetBox="[151,1435,181,1627]" captionTargetId="figure@21.[151,1435,181,1627]" captionTargetPageId="21" captionText="FIGURE 16. Prothorax of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765684" httpUri="https://zenodo.org/record/3765684/files/figure.png" pageId="29" pageNumber="530">16e</figureCitation>
,
<figureCitation id="BFF82A464371616DFD09DDF84B173CF1" box="[756,799,1672,1698]" captionStart="FIGURE 17" captionStartId="22.[151,250,1779,1804]" captionTargetBox="[151,1436,309,1754]" captionTargetId="figure@22.[151,1436,309,1754]" captionTargetPageId="22" captionText="FIGURE 17. Prothorax of female Coeliccia spp. used for description in dorsal view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765686" httpUri="https://zenodo.org/record/3765686/files/figure.png" pageId="29" pageNumber="530">17e</figureCitation>
,
<figureCitation id="BFF82A464371616DFCD1DDF84B5F3CF1" box="[812,855,1672,1698]" captionStart="FIGURE 19" captionStartId="24.[151,250,1197,1222]" captionTargetBox="[182,1384,181,1171]" captionTargetId="figure@24.[182,1384,181,1171]" captionTargetPageId="24" captionText="FIGURE 19. Thorax of female Coeliccia spp. used for description in lateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765690" httpUri="https://zenodo.org/record/3765690/files/figure.png" pageId="29" pageNumber="530">19e</figureCitation>
,
<figureCitation id="BFF82A464371616DFC9EDDF84B993CF0" box="[867,913,1672,1699]" captionStart="FIGURE 21" captionStartId="26.[151,250,1018,1043]" captionTargetBox="[151,1436,181,993]" captionTargetId="figure@26.[151,1436,181,993]" captionTargetPageId="26" captionText="FIGURE 21. Posterior pronotal lobe of Coeliccia spp. used for description in oblique dorsal view. (a) C. suoitia; (b) C. pulchella; (c) C. galbina; (d) C. caerulea." figureDoi="http://doi.org/10.5281/zenodo.3765694" httpUri="https://zenodo.org/record/3765694/files/figure.png" pageId="29" pageNumber="530">21b</figureCitation>
).
<emphasis id="15B7EAD14371616DFC5BDDF84BEB3CF1" box="[934,995,1672,1698]" italics="true" pageId="29" pageNumber="530">Head</emphasis>
(
<figureCitation id="BFF82A464371616DFC0EDDF84C5C3CF1" box="[1011,1108,1672,1698]" captionStart="FIGURE 14" captionStartId="19.[151,250,1649,1674]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure@19.[151,1436,181,1623]" captionTargetPageId="19" captionText="FIGURE 14. Heads of female Coeliccia ssp. used for description in dorsolateral view. (a) C. coronata; (b) C. diomedea; (c) C. hayashii; (d) C. galbina; (e) C. pulchella; (f) C. suoitia." figureDoi="http://doi.org/10.5281/zenodo.3765680" httpUri="https://zenodo.org/record/3765680/files/figure.png" pageId="29" pageNumber="530">Fig. 14e</figureCitation>
). Labrum yellow with dorsal margin dark brown. Anteclypeus yellow, with two darker areas left and right on lower half. Postclypeus shiny black. Genae and base of mandibles yellow, the yellow connected over the antefrons above the postclypeus, remainder of frons matte black. Dorsal surface of head matte black, apart from two small yellow spots between base of antennae and lateral ocelli and two oblong pale yellow postocular spots. Antennae blackish brown, top of first segment and base of second segment pale yellow. Compound eyes in life black dorsally, green ventrally.
</paragraph>
<paragraph id="277C36C34371616EFF3ADC104A933B4C" blockId="29.[151,1437,1671,2023]" lastBlockId="30.[151,1436,153,539]" lastPageId="30" lastPageNumber="531" pageId="29" pageNumber="530">
<emphasis id="15B7EAD14371616DFF3ADC1049103D29" box="[199,280,1888,1914]" italics="true" pageId="29" pageNumber="530">Thorax</emphasis>
. Prothorax (16e, 17e, 21b): yellow with frontal part of anterior pronotal lobe and with posterior pronotal lobe blackish brown. Posterior pronotal lobe with central part tapering from base to rounded apex, noticeably inflated (
<figureCitation id="BFF82A464371616DFF1BDCD8494D3D90" box="[230,325,1960,1987]" captionStart="FIGURE 21" captionStartId="26.[151,250,1018,1043]" captionTargetBox="[151,1436,181,993]" captionTargetId="figure@26.[151,1436,181,993]" captionTargetPageId="26" captionText="FIGURE 21. Posterior pronotal lobe of Coeliccia spp. used for description in oblique dorsal view. (a) C. suoitia; (b) C. pulchella; (c) C. galbina; (d) C. caerulea." figureDoi="http://doi.org/10.5281/zenodo.3765694" httpUri="https://zenodo.org/record/3765694/files/figure.png" pageId="29" pageNumber="530">Fig. 21b</figureCitation>
) in dorsal view, in lateral view raised, inflated until near apex where thin and flattened. Lapels well developed, rounded. Synthorax: mesepisternum blackish brown, with two yellow, almost square, spots, on one side bordered by mesopleural suture, the first on the anterior half of the mesepisternum a little larger than the second on the middle of the posterior half (
<figureCitation id="BFF82A464372616EFDF0DBCD4A683A84" box="[525,608,189,215]" captionStart="FIGURE 27" captionStartId="32.[151,250,1908,1933]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure@32.[151,1436,597,1883]" captionTargetPageId="32" captionText="FIGURE 27. Habitus of Coeliccia spp. [a-c] C. diomedea, [d-f] C. pulchella from Huu Lien NR, Lang Son Prov. (a) male, Thua Thien-Hue Prov., along QL14, 14 v 2017; (b) male, Quang Nam Prov., along QL14, 27 ix 2015; (c) female, Quang Nam Prov., along QL14, 26 ix 2015; (d) female, 26 vii 2014; (e) copula, 11 x 2015; (f) male, 14 vii 2014." figureDoi="http://doi.org/10.5281/zenodo.3765706" httpUri="https://zenodo.org/record/3765706/files/figure.png" pageId="30" pageNumber="531">Fig. 27</figureCitation>
de, which show live specimens). Mesepimeron and mesinfraepisternum brown. Metepisternum and metepimeron pale yellow. Legs: coxae and trochanters pale yellow. Femur pale yellow with brown lines. Claws and spines brownish.
</paragraph>
<paragraph id="277C36C34372616EFF3ADA5A4C223B10" blockId="30.[151,1436,153,539]" box="[199,1066,296,323]" pageId="30" pageNumber="531">
<emphasis id="15B7EAD14372616EFF3ADA5A49033B10" box="[199,267,298,323]" italics="true" pageId="30" pageNumber="531">Wings</emphasis>
. Hyaline, 19 Px in FW,
<quantity id="E03B9B264372616EFDE7DA594A5B3B10" box="[538,595,297,323]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="30" pageNumber="531" unit="in" value="18.0">18 in</quantity>
HW. Pt dark brown, covering 11.5 cells.
</paragraph>
<paragraph id="277C36C34372616EFF3ADA3D4AC03BFC" blockId="30.[151,1436,153,539]" pageId="30" pageNumber="531">
<emphasis id="15B7EAD14372616EFF3ADA3D49313B34" box="[199,313,333,359]" italics="true" pageId="30" pageNumber="531">Abdomen.</emphasis>
S1 yellow with posterior margin brown; S2 brown dorsally and yellow below; S37 brown dorsally and paler yellow below, darker at the basal and apical margins and with prominent subapical whitish yellow annulus, S7 without dark apical margin; S810 yellow.
</paragraph>
<paragraph id="277C36C34372616EFF3ADAC949273BA4" blockId="30.[151,1436,153,539]" pageId="30" pageNumber="531">
<emphasis id="15B7EAD14372616EFF3ADAC949803B80" box="[199,392,441,467]" italics="true" pageId="30" pageNumber="531">Anal appendages</emphasis>
. Cerci brown, half as long as S10, ovipositor yellow, extending almost 1.5 times length of S10 beyond cerci.
</paragraph>
<paragraph id="277C36C34372616EFF3AD9714B853848" blockId="30.[151,1436,153,539]" box="[199,909,513,539]" pageId="30" pageNumber="531">Measurements (in mm). HW 23; abdomen incl. appendages 34.</paragraph>
<caption id="73BC664B4372616EFF6ADC3E4D5A3DFC" ID-DOI="http://doi.org/10.5281/zenodo.3765702" ID-Zenodo-Dep="3765702" httpUri="https://zenodo.org/record/3765702/files/figure.png" pageId="30" pageNumber="531" startId="30.[151,250,1870,1895]" targetBox="[151,1436,561,1845]" targetPageId="30">
<paragraph id="277C36C34372616EFF6ADC3E4D5A3DFC" blockId="30.[151,1437,1870,1967]" pageId="30" pageNumber="531">
<emphasis id="15B7EAD14372616EFF6ADC3E49283D34" bold="true" box="[151,288,1870,1895]" pageId="30" pageNumber="531">FIGURE 25.</emphasis>
Thorax and head of alive male
<taxonomicName id="E0C34D404372616EFD99DC3F4ACD3D34" authorityName="Kirby" authorityYear="1890" box="[612,709,1871,1895]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD14372616EFD99DC3F4ACD3D34" box="[612,709,1871,1895]" italics="true" pageId="30" pageNumber="531">Coeliccia</emphasis>
</taxonomicName>
spp. [a-d]
<taxonomicName id="E0C34D404372616EFCC9DC3F4B853D34" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[820,909,1871,1895]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="531" phylum="Arthropoda" rank="species" species="curua">
<emphasis id="15B7EAD14372616EFCC9DC3F4B853D34" box="[820,909,1871,1895]" italics="true" pageId="30" pageNumber="531">C. curua</emphasis>
</taxonomicName>
, [e,f]
<taxonomicName id="E0C34D404372616EFC31DC3F4C143D35" box="[972,1052,1871,1894]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="531" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD14372616EFC31DC3F4C143D35" box="[972,1052,1871,1894]" italics="true" pageId="30" pageNumber="531">C.uenoi</emphasis>
</taxonomicName>
. (a,b) paratype in dorsolateral (a) and lateral (b) view, Xuan Son NP, Phu Tho Prov., 23 vii 2017; (c) holotype in dorsolateral view; (d) holotype in lateral view; (e) dorsolateral view, Xuan Son NP, Phu Tho Prov., 9 vi 2018; (f) lateral view, Cuc Phuong NP, Ninh Binh Prov., 3 v 2014.
</paragraph>
</caption>
<paragraph id="277C36C34373616FFF3ADBE84A1D3B10" blockId="31.[151,1437,152,611]" pageId="31" pageNumber="532">
<emphasis id="15B7EAD14373616FFF3ADBE84A2C3AE0" bold="true" box="[199,548,152,179]" pageId="31" pageNumber="532">
Variation in female
<typeStatus id="F87888614373616FFE4CDBE94A163AE0" box="[433,542,153,179]" pageId="31" pageNumber="532" type="paratype">paratype</typeStatus>
.
</emphasis>
The central part of the posterior pronotal lobe of the prothorax is not noticeably inflated. Coloration generally blacker. Mesepisternum, mesepimeron and mesinfraepisternum black, the posterior spot on mesepisternum distinct, but clearly smaller. Some black over posterior end of metapleural suture. Dorsum of abdominal segments blackish brown. S9 mostly black, with anterodorsal yellow spot. S10 and cerci black, ovipositor black. 16 Px in FW,
<quantity id="E03B9B264373616FFE5FDA5849D33B10" box="[418,475,296,323]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="31" pageNumber="532" unit="in" value="15.0">15 in</quantity>
HW.
</paragraph>
<paragraph id="277C36C34373616FFF3ADA3D4BAA3B34" blockId="31.[151,1437,152,611]" box="[199,930,332,359]" pageId="31" pageNumber="532">Measurements (in mm). HW 26.5; abdomen incl. appendages 39.</paragraph>
</subSubSection>
<subSubSection id="6FD9654843736150FF3ADA004C443B10" lastPageId="32" lastPageNumber="533" pageId="31" pageNumber="532" type="diagnosis">
<paragraph id="277C36C34373616FFF3ADA004A2F3830" blockId="31.[151,1437,152,611]" pageId="31" pageNumber="532">
<emphasis id="15B7EAD14373616FFF3ADA0049C53BD8" bold="true" box="[199,461,368,395]" pageId="31" pageNumber="532">Differential diagnosis.</emphasis>
The male shares the combination of small squarish yellow spots on the otherwise black dorsum of the thorax in
<collectingCountry id="5FD476534373616FFE5EDAE54A0A3BFC" box="[419,514,405,431]" name="Vietnam" pageId="31" pageNumber="532">Vietnam</collectingCountry>
with
<taxonomicName id="E0C34D404373616FFDBCDAE54AC73BFC" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[577,719,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="diomedea">
<emphasis id="15B7EAD14373616FFDBCDAE54AC73BFC" box="[577,719,405,431]" italics="true" pageId="31" pageNumber="532">C. diomedea</emphasis>
</taxonomicName>
(
<figureCitation id="BFF82A464373616FFD23DAE54B273BFC" box="[734,815,405,431]" captionStart="FIGURE 27" captionStartId="32.[151,250,1908,1933]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure@32.[151,1436,597,1883]" captionTargetPageId="32" captionText="FIGURE 27. Habitus of Coeliccia spp. [a-c] C. diomedea, [d-f] C. pulchella from Huu Lien NR, Lang Son Prov. (a) male, Thua Thien-Hue Prov., along QL14, 14 v 2017; (b) male, Quang Nam Prov., along QL14, 27 ix 2015; (c) female, Quang Nam Prov., along QL14, 26 ix 2015; (d) female, 26 vii 2014; (e) copula, 11 x 2015; (f) male, 14 vii 2014." figureDoi="http://doi.org/10.5281/zenodo.3765706" httpUri="https://zenodo.org/record/3765706/files/figure.png" pageId="31" pageNumber="532">Fig. 27</figureCitation>
) and
<emphasis id="15B7EAD14373616FFC90DAE54C0F3BFC" box="[877,1031,405,431]" italics="true" pageId="31" pageNumber="532">
<taxonomicName id="E0C34D404373616FFC90DAE54C0B3BFC" authorityName="Asahina" authorityYear="1984" box="[877,1027,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="yamasakii">C. yamasakii</taxonomicName>
.
</emphasis>
Lack of pruinosity and the structure of its genital ligula exclude
<emphasis id="15B7EAD14373616FFE30DAC94A6A3B80" box="[461,610,441,467]" italics="true" pageId="31" pageNumber="532">
<taxonomicName id="E0C34D404373616FFE30DAC94A573B80" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[461,607,441,467]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="diomedea">C. diomedea</taxonomicName>
.
</emphasis>
Both
<taxonomicName id="E0C34D404373616FFD58DAC94B303B80" authorityName="Asahina" authorityYear="1984" box="[677,824,441,467]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="yamasakii">
<emphasis id="15B7EAD14373616FFD58DAC94B303B80" box="[677,824,441,467]" italics="true" pageId="31" pageNumber="532">C. yamasakii</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D404373616FFC90DAC94BF23B80" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[877,1018,441,467]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="diomedea">
<emphasis id="15B7EAD14373616FFC90DAC94BF23B80" box="[877,1018,441,467]" italics="true" pageId="31" pageNumber="532">C. diomedea</emphasis>
</taxonomicName>
differ by,
<emphasis id="15B7EAD14373616FFB90DACA4CDD3B80" box="[1133,1237,441,467]" italics="true" pageId="31" pageNumber="532">inter alia</emphasis>
, the lack of white rings on the abdomen, much less yellow on the abdominal tip (no yellow on S8, reduced or no yellow on S9) and both these species also have a much more pronounced black line over the metapleural suture. The genital ligula of
<emphasis id="15B7EAD14373616FFF6AD9554938386C" box="[151,304,549,575]" italics="true" pageId="31" pageNumber="532">
<taxonomicName id="E0C34D404373616FFF6AD9554923386C" authorityName="Asahina" authorityYear="1984" box="[151,299,549,575]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="yamasakii">C. yamasakii</taxonomicName>
,
</emphasis>
although similar in structure, has the flagella placed further distally from the base of the terminal segment (see Fig.
<quantity id="E03B9B264373616FFEC3D938497F3830" box="[318,375,584,611]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="31" pageNumber="532" unit="in" value="50.0">50 in</quantity>
<bibRefCitation id="43524B324373616FFE81D9394A133830" author="Asahina, S." box="[380,539,585,611]" pageId="31" pageNumber="532" pagination="2 - 20" refId="ref18344" refString="Asahina, S. (1984) A list of the Odonata recorded from Thailand Part VI. Platycnemididae-Genus Coeliccia. Tombo, 27, 2 - 20." type="journal article" year="1984">Asahina 1984</bibRefCitation>
).
</paragraph>
<caption id="73BC664B4373616FFF6ADCE64CEE3D80" ID-DOI="http://doi.org/10.5281/zenodo.3765704" ID-Zenodo-Dep="3765704" httpUri="https://zenodo.org/record/3765704/files/figure.png" pageId="31" pageNumber="532" startId="31.[151,250,1942,1967]" targetBox="[151,1436,633,1917]" targetPageId="31">
<paragraph id="277C36C34373616FFF6ADCE64CEE3D80" blockId="31.[151,1436,1942,2003]" pageId="31" pageNumber="532">
<emphasis id="15B7EAD14373616FFF6ADCE649173DFC" bold="true" box="[151,287,1942,1967]" pageId="31" pageNumber="532">FIGURE 26.</emphasis>
Habitus of
<taxonomicName id="E0C34D404373616FFE6FDCE749FB3DFC" authorityName="Kirby" authorityYear="1890" box="[402,499,1943,1967]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD14373616FFE6FDCE749FB3DFC" box="[402,499,1943,1967]" italics="true" pageId="31" pageNumber="532">Coeliccia</emphasis>
</taxonomicName>
spp. from Bao Loc, Lam Dong Prov. [a-c]
<taxonomicName id="E0C34D404373616FFC60DCE74C1B3DFC" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[925,1043,1943,1967]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis id="15B7EAD14373616FFC60DCE74C1B3DFC" box="[925,1043,1943,1967]" italics="true" pageId="31" pageNumber="532">C. caerulea</emphasis>
</taxonomicName>
, [d-f]
<taxonomicName id="E0C34D404373616FFBACDCE74CC13DFC" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1105,1225,1943,1967]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="532" phylum="Arthropoda" rank="species" species="coronata">
<emphasis id="15B7EAD14373616FFBACDCE74CC13DFC" box="[1105,1225,1943,1967]" italics="true" pageId="31" pageNumber="532">C. coronata</emphasis>
</taxonomicName>
(a) male, 13 vi 2016; (b) male, 18 v 2016; (c) female, 16 vi 2016; (d) female, 13 vi 2016; (e) male, 16 v 2016; (f) male, 16 v 2016.
</paragraph>
</caption>
<paragraph id="277C36C3434C6150FF3ADBE94C443B10" blockId="32.[151,1437,153,575]" pageId="32" pageNumber="533">
The female has double spots on the mesepisternum (
<figureCitation id="BFF82A46434C6150FCE4DBE94B633AE7" box="[793,875,153,180]" captionStart="FIGURE 27" captionStartId="32.[151,250,1908,1933]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure@32.[151,1436,597,1883]" captionTargetPageId="32" captionText="FIGURE 27. Habitus of Coeliccia spp. [a-c] C. diomedea, [d-f] C. pulchella from Huu Lien NR, Lang Son Prov. (a) male, Thua Thien-Hue Prov., along QL14, 14 v 2017; (b) male, Quang Nam Prov., along QL14, 27 ix 2015; (c) female, Quang Nam Prov., along QL14, 26 ix 2015; (d) female, 26 vii 2014; (e) copula, 11 x 2015; (f) male, 14 vii 2014." figureDoi="http://doi.org/10.5281/zenodo.3765706" httpUri="https://zenodo.org/record/3765706/files/figure.png" pageId="32" pageNumber="533">Fig. 27</figureCitation>
de), which in
<collectingCountry id="5FD47653434C6150FBE9DBE94C7B3AE0" box="[1044,1139,153,179]" name="Vietnam" pageId="32" pageNumber="533">Vietnam</collectingCountry>
it shares only with the female of
<taxonomicName id="E0C34D40434C6150FF08DBCD4A443A84" authority="Do, 2011" authorityName="Do" authorityYear="2011" box="[245,588,189,215]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="sasamotoi">
<emphasis id="15B7EAD1434C6150FF08DBCD49D23A84" box="[245,474,189,215]" italics="true" pageId="32" pageNumber="533">Coeliccia sasamotoi</emphasis>
Do, 2011
</taxonomicName>
. The anterior spot in that species is clearly elongated and generally at least twice as long as the posterior spot.
<taxonomicName id="E0C34D40434C6150FDEADB914AF23AA8" authorityName="Do" authorityYear="2011" box="[535,762,225,251]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="sasamotoi">
<emphasis id="15B7EAD1434C6150FDEADB914AF23AA8" box="[535,762,225,251]" italics="true" pageId="32" pageNumber="533">Coeliccia sasamotoi</emphasis>
</taxonomicName>
has a shiny black labrum and has a different pattern on S89, with distinct white dorsal half-rings on the posterior half of the segments. The elongated central part of the posterior pronotal lobe of the prothorax of
<taxonomicName id="E0C34D40434C6150FDF1DA594A973B10" authorityName="Do" authorityYear="2011" box="[524,671,297,323]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="sasamotoi">
<emphasis id="15B7EAD1434C6150FDF1DA594A973B10" box="[524,671,297,323]" italics="true" pageId="32" pageNumber="533">C. sasamotoi</emphasis>
</taxonomicName>
is smaller and not noticeably inflated.
</paragraph>
</subSubSection>
<subSubSection id="6FD96548434C6151FF3ADA3D4D2E3B73" lastPageId="33" lastPageNumber="534" pageId="32" pageNumber="533" type="discussion">
<paragraph id="277C36C3434C6150FF3ADA3D4BD33B80" blockId="32.[151,1437,153,575]" pageId="32" pageNumber="533">
<taxonomicName id="E0C34D40434C6150FF3ADA3D4A3C3B34" authority="Asahina, 1970" authorityName="Asahina" authorityYear="1970" box="[199,564,333,359]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="svihleri">
<emphasis id="15B7EAD1434C6150FF3ADA3D49853B34" box="[199,397,333,359]" italics="true" pageId="32" pageNumber="533">Coeliccia svihleri</emphasis>
Asahina, 1970
</taxonomicName>
occurs in
<collectingRegion id="E507F821434C6150FD5BDA3D4AFA3B34" box="[678,754,333,359]" country="India" name="Assam" pageId="32" pageNumber="533">Assam</collectingRegion>
(
<collectingCountry id="5FD47653434C6150FCFCDA3D4B483B3B" box="[769,832,333,360]" name="India" pageId="32" pageNumber="533">India</collectingCountry>
),
<collectingCountry id="5FD47653434C6150FCA9DA3D4BA03B34" box="[852,936,333,359]" name="Bhutan" pageId="32" pageNumber="533">Bhutan</collectingCountry>
,
<collectingCountry id="5FD47653434C6150FC4FDA3D4C173B34" box="[946,1055,333,359]" name="Myanmar" pageId="32" pageNumber="533">Myanmar</collectingCountry>
and
<collectingRegion id="E507F821434C6150FBA9DA3D4CA23B34" box="[1108,1194,333,359]" country="China" name="Yunnan" pageId="32" pageNumber="533">Yunnan</collectingRegion>
(
<collectingCountry id="5FD47653434C6150FB45DA3D4CF63B3B" box="[1208,1278,333,360]" name="China" pageId="32" pageNumber="533">China</collectingCountry>
) (
<bibRefCitation id="43524B32434C6150FAE8DA3D4D993B34" author="Dow, R. A." box="[1301,1425,333,359]" pageId="32" pageNumber="533" pagination="169163" refId="ref18519" refString="Dow, R. A. (2019) Coeliccia svihleri. The IUCN Red List of Threatened Species, 2019, e. T 169163 A 123029000. Availabel from: https: // www. iucnredlist. org / species / 169163 / 6574947 (accessed 13 February 2019) https: // doi. org / 10.2305 / IUCN. UK. 2019 - 2. RLTS. T 169163 A 123029000. en" type="journal article" year="2019">Dow 2019</bibRefCitation>
). The shape of its ligula (
<bibRefCitation id="43524B32434C6150FE63DA014A373BD8" author="Asahina, S." box="[414,575,368,395]" pageId="32" pageNumber="533" pagination="2 - 13" refId="ref18373" refString="Asahina, S. (1985) Further contributions to the taxonomy of Southasiatic Coeliccia species (Odonata, Platycnemididae). Cho Cho, 8 (2), 2 - 13." type="journal article" year="1985">Asahina 1985</bibRefCitation>
: figs 4243) appears to be similar to that of
<taxonomicName id="E0C34D40434C6150FBD1DA014CBD3BD8" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1068,1205,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="15B7EAD1434C6150FBD1DA014CBD3BD8" box="[1068,1205,369,395]" italics="true" pageId="32" pageNumber="533">C. pulchella</emphasis>
</taxonomicName>
, but it differs by, for instance, black markings on the metepisternum and metepimeron (all yellow in
<taxonomicName id="E0C34D40434C6150FBF1DAE54C9C3BFC" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1036,1172,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="15B7EAD1434C6150FBF1DAE54C9C3BFC" box="[1036,1172,405,431]" italics="true" pageId="32" pageNumber="533">C. pulchella</emphasis>
</taxonomicName>
), by the markings of its abdomen and by the shape of the cerci (
<bibRefCitation id="43524B32434C6150FDA9DAC94AFC3B80" author="Asahina, S." box="[596,756,440,467]" pageId="32" pageNumber="533" pagination="2 - 13" refId="ref18373" refString="Asahina, S. (1985) Further contributions to the taxonomy of Southasiatic Coeliccia species (Odonata, Platycnemididae). Cho Cho, 8 (2), 2 - 13." type="journal article" year="1985">Asahina 1985</bibRefCitation>
: figs 3941, 4546).
</paragraph>
<paragraph id="277C36C3434C6150FF3ADAAD4996386C" blockId="32.[151,1437,153,575]" pageId="32" pageNumber="533">
<emphasis id="15B7EAD1434C6150FF3ADAAD49333BA4" bold="true" box="[199,315,477,503]" pageId="32" pageNumber="533">Remarks.</emphasis>
The difference between the general darkness of the female
<typeStatus id="F8788861434C6150FC09DAAD4C573BA4" box="[1012,1119,477,503]" pageId="32" pageNumber="533" type="paratype">paratypes</typeStatus>
likely results from the fact that the
<typeStatus id="F8788861434C6150FF0ED971495B3848" box="[243,339,513,539]" pageId="32" pageNumber="533" type="paratype">paratype</typeStatus>
used for description is not fully mature. This is supported by the photos of additional specimens provided in
<figureCitation id="BFF82A46434C6150FEE2D9554967386C" box="[287,367,549,575]" captionStart="FIGURE 27" captionStartId="32.[151,250,1908,1933]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure@32.[151,1436,597,1883]" captionTargetPageId="32" captionText="FIGURE 27. Habitus of Coeliccia spp. [a-c] C. diomedea, [d-f] C. pulchella from Huu Lien NR, Lang Son Prov. (a) male, Thua Thien-Hue Prov., along QL14, 14 v 2017; (b) male, Quang Nam Prov., along QL14, 27 ix 2015; (c) female, Quang Nam Prov., along QL14, 26 ix 2015; (d) female, 26 vii 2014; (e) copula, 11 x 2015; (f) male, 14 vii 2014." figureDoi="http://doi.org/10.5281/zenodo.3765706" httpUri="https://zenodo.org/record/3765706/files/figure.png" pageId="32" pageNumber="533">Fig. 27</figureCitation>
de.
</paragraph>
<caption id="73BC664B434C6150FF6ADC044C423D87" ID-DOI="http://doi.org/10.5281/zenodo.3765706" ID-Zenodo-Dep="3765706" httpUri="https://zenodo.org/record/3765706/files/figure.png" pageId="32" pageNumber="533" startId="32.[151,250,1908,1933]" targetBox="[151,1436,597,1883]" targetPageId="32">
<paragraph id="277C36C3434C6150FF6ADC044C423D87" blockId="32.[151,1436,1908,2005]" pageId="32" pageNumber="533">
<emphasis id="15B7EAD1434C6150FF6ADC0449173DDE" bold="true" box="[151,287,1908,1933]" pageId="32" pageNumber="533">FIGURE 27.</emphasis>
Habitus of
<taxonomicName id="E0C34D40434C6150FE6EDC0449FC3DDF" authorityName="Kirby" authorityYear="1890" box="[403,500,1908,1932]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD1434C6150FE6EDC0449FC3DDF" box="[403,500,1908,1932]" italics="true" pageId="32" pageNumber="533">Coeliccia</emphasis>
</taxonomicName>
spp. [a-c]
<taxonomicName id="E0C34D40434C6150FDA2DC054AD53DDF" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[607,733,1908,1932]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="diomedea">
<emphasis id="15B7EAD1434C6150FDA2DC054AD53DDF" box="[607,733,1908,1932]" italics="true" pageId="32" pageNumber="533">C. diomedea</emphasis>
</taxonomicName>
, [d-f]
<taxonomicName id="E0C34D40434C6150FCE3DC054B903DDF" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[798,920,1908,1932]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="533" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="15B7EAD1434C6150FCE3DC054B903DDF" box="[798,920,1908,1932]" italics="true" pageId="32" pageNumber="533">C. pulchella</emphasis>
</taxonomicName>
from Huu Lien NR, Lang Son Prov. (a) male, Thua Thien-Hue Prov., along QL14, 14 v 2017; (b) male, Quang Nam Prov., along QL14, 27 ix 2015; (c) female, Quang Nam Prov., along QL14, 26 ix 2015; (d) female, 26 vii 2014; (e) copula, 11 x 2015; (f) male, 14 vii 2014.
</paragraph>
</caption>
<paragraph id="277C36C3434D6151FF3ADBE94CB33A84" blockId="33.[151,1437,153,431]" pageId="33" pageNumber="534">
The male of
<taxonomicName id="E0C34D40434D6151FEA5DBE94ADA3AE0" authority="Asahina, 1984" authorityName="Asahina" authorityYear="1984" box="[344,722,153,179]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="schmidti">
<emphasis id="15B7EAD1434D6151FEA5DBE94A213AE0" box="[344,553,153,179]" italics="true" pageId="33" pageNumber="534">Coeliccia schmidti</emphasis>
<bibRefCitation id="43524B32434D6151FDCDDBE94ADA3AE0" author="Asahina, S." box="[560,722,153,179]" pageId="33" pageNumber="534" pagination="2 - 20" refId="ref18344" refString="Asahina, S. (1984) A list of the Odonata recorded from Thailand Part VI. Platycnemididae-Genus Coeliccia. Tombo, 27, 2 - 20." type="journal article" year="1984">Asahina, 1984</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="5FD47653434D6151FCE5DBE94B5A3AE0" box="[792,850,153,179]" name="India" pageId="33" pageNumber="534">India</collectingCountry>
looks rather similar to the male of
<taxonomicName id="E0C34D40434D6151FB1DDBE94D623AE0" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[1248,1386,153,179]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="15B7EAD1434D6151FB1DDBE94D623AE0" box="[1248,1386,153,179]" italics="true" pageId="33" pageNumber="534">C. pulchella</emphasis>
</taxonomicName>
, but the female has an elongated antehumeral stripe, not spots, on the thorax (
<bibRefCitation id="43524B32434D6151FC35DBCD4CA73A84" author="Joshi, S. &amp; Kunte, K." box="[968,1199,189,215]" pageId="33" pageNumber="534" pagination="6458 - 6472" refId="ref18628" refString="Joshi, S. &amp; Kunte, K. (2014) Dragonflies and damselflies (Insecta: Odonata) of Nagaland, with an addition to the Indian odonate fauna. Journal of Threatened Taxa, 6 (11), 6458 - 6472. https: // doi. org / 10.11609 / JoTT. o 3911.6458 - 72" type="journal article" year="2014">Joshi &amp; Kunte 2014</bibRefCitation>
).
</paragraph>
<paragraph id="277C36C3434D6151FF3ADB914D2E3B73" blockId="33.[151,1437,153,431]" pageId="33" pageNumber="534">
The genital ligulae of both subspecies of
<taxonomicName id="E0C34D40434D6151FD69DB914B073AA8" authorityName="Fraser" authorityYear="1924" box="[660,783,225,251]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="poungyi">
<emphasis id="15B7EAD1434D6151FD69DB914B073AA8" box="[660,783,225,251]" italics="true" pageId="33" pageNumber="534">C. poungyi</emphasis>
</taxonomicName>
look somewhat similar, but have the flagella further distal from the base of the terminal segment and a much longer central tongue (
<bibRefCitation id="43524B32434D6151FC36DA754C633B4C" author="Asahina, S." box="[971,1131,261,287]" pageId="33" pageNumber="534" pagination="2 - 20" refId="ref18344" refString="Asahina, S. (1984) A list of the Odonata recorded from Thailand Part VI. Platycnemididae-Genus Coeliccia. Tombo, 27, 2 - 20." type="journal article" year="1984">Asahina 1984</bibRefCitation>
,
<bibRefCitation id="43524B32434D6151FB8BDA754D123B73" author="Kosterin, O. E." box="[1142,1306,261,288]" pageId="33" pageNumber="534" pagination="1 - 16" refId="ref18777" refString="Kosterin, O. E. (2016) Coeliccia poungyi dasha subsp. nov. (Odonata, Platycnemididae, Calicnemiinae) from eastern Cambodia. International Dragonfly Fund-Report, 97, 1 - 16." type="journal article" year="2016">Kosterin 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6FD96548434D6151FF3ADA584B0C3BFC" pageId="33" pageNumber="534" type="biology_ecology">
<paragraph id="277C36C3434D6151FF3ADA584B0C3BFC" blockId="33.[151,1437,153,431]" pageId="33" pageNumber="534">
<emphasis id="15B7EAD1434D6151FF3ADA5849B73B10" bold="true" box="[199,447,296,323]" pageId="33" pageNumber="534">Habitat and ecology.</emphasis>
In Huu Lien NR and Cuc Phuong NP
<taxonomicName id="E0C34D40434D6151FC86DA594C5E3B10" authorityName="Kompier &amp; Dow &amp; Steinhoff" authorityYear="2020" box="[891,1110,297,323]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="pulchella">
<emphasis id="15B7EAD1434D6151FC86DA594C5E3B10" box="[891,1110,297,323]" italics="true" pageId="33" pageNumber="534">Coeliccia pulchella</emphasis>
</taxonomicName>
is found in karst mountains, where it can be quite common in the vicinity of small streams. In Huu Lien NR it was found together with the rare
<taxonomicName id="E0C34D40434D6151FA7EDA3D48E53BD8" authorityName="Wilson &amp; Reels" authorityYear="2003" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="galbina">
<emphasis id="15B7EAD1434D6151FA7EDA3D48E53BD8" italics="true" pageId="33" pageNumber="534">C. galbina</emphasis>
</taxonomicName>
at a particularly rocky hill side, but otherwise no other
<taxonomicName id="E0C34D40434D6151FCA6DA014BCE3BD8" authorityName="Kirby" authorityYear="1890" box="[859,966,369,395]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="15B7EAD1434D6151FCA6DA014BCE3BD8" box="[859,966,369,395]" italics="true" pageId="33" pageNumber="534">Coeliccia</emphasis>
</taxonomicName>
species were seen nearby. In Cuc Phuong NP it shares its habitat with
<taxonomicName id="E0C34D40434D6151FE2FDAE54A603BFC" authorityName="Laidlaw" authorityYear="1932" box="[466,616,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="pyriformis">
<emphasis id="15B7EAD1434D6151FE2FDAE54A603BFC" box="[466,616,405,431]" italics="true" pageId="33" pageNumber="534">C. pyriformis</emphasis>
</taxonomicName>
and
<taxonomicName id="E0C34D40434D6151FD63DAE54AF43BFC" box="[670,764,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="534" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="15B7EAD1434D6151FD63DAE54AF43BFC" box="[670,764,405,431]" italics="true" pageId="33" pageNumber="534">C. uenoi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>