412 lines
58 KiB
XML
412 lines
58 KiB
XML
<document id="38FBB007DF2BEB62AFF5FCCEF3D49DAB" ID-CLB-Dataset="22814" ID-DOI="10.11646/zootaxa.4766.4.1" ID-GBIF-Dataset="1e2996e2-e31f-45e3-9149-60ebce934708" ID-ISSN="1175-5334" ID-Zenodo-Dep="3765652" ID-ZooBank="urn:lsid:zoobank.org:pub:2F8FEDC1-6891-46D1-B372-858CDCBE4051" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587754498722" checkinUser="carolina" docAuthor="Kompier, Tom, Dow, Rory A. & Steinhoff, Philip O. M." docDate="2020" docId="AF6A87D5436B617DFF6ADFB64ACF3FB4" docLanguage="en" docName="zootaxa.4766.4.1.pdf.imf" docOrigin="Zootaxa 4766 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia pyriformis Laidlaw 1932" docType="treatment" docVersion="6" lastPageNumber="514" masterDocId="5353FFAD436C6170FFFDDB7048083A53" masterDocTitle="Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus" masterLastPageNumber="538" masterPageNumber="501" pageNumber="508" updateTime="1698835840532" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FED86CC19CC80320A0777F865F07E613">Five new species of Coeliccia Kirby, 1890 from Vietnam (Odonata: Platycnemididae), and information on several other species of the genus</mods:title>
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<mods:name id="EEDD46A96DC609378BA84B1376A24055" type="personal">
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<mods:namePart id="4A94795EE07C160F12DFF926AC764494">Kompier, Tom</mods:namePart>
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<mods:namePart id="017E06CB38343EAA45742F9D6A78C530">Dow, Rory A.</mods:namePart>
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<mods:namePart id="812C0E878A331B8F560429A0D560411C">Steinhoff, Philip O. M.</mods:namePart>
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<mods:typeOfResource id="A029DA836B2081F4ED0AAA348306DA3B">text</mods:typeOfResource>
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<mods:title id="30FA4C87E6F61A4F3BCBC49E46455AFE">Zootaxa</mods:title>
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<mods:part id="EB5A3F31725157FD27E227D8D931F7D0">
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<mods:date id="E047B812683018A2CF6209EDDAF5AB38">2020</mods:date>
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<mods:number id="39A3C7C4DB532F4F1EB4F4723DC992EB">2020-04-22</mods:number>
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<mods:number id="528EF2464844B6DAF3DCE2E89E492257">4766</mods:number>
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<treatment id="AF6A87D5436B617DFF6ADFB64ACF3FB4" ID-DOI="http://doi.org/10.5281/zenodo.3803424" ID-GBIF-Taxon="163739038" ID-Zenodo-Dep="3803424" LSID="urn:lsid:plazi:treatment:AF6A87D5436B617DFF6ADFB64ACF3FB4" httpUri="http://treatment.plazi.org/id/AF6A87D5436B617DFF6ADFB64ACF3FB4" lastPageId="13" lastPageNumber="514" pageId="7" pageNumber="508">
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<subSubSection id="6FD96548436B6177FF6ADFB64A683EB3" box="[151,608,1222,1249]" pageId="7" pageNumber="508" type="nomenclature">
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<paragraph id="277C36C3436B6177FF6ADFB64A683EB3" blockId="7.[151,608,1222,1249]" box="[151,608,1222,1249]" pageId="7" pageNumber="508">
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<heading id="7C3481AF436B6177FF6ADFB64A683EB3" bold="true" box="[151,608,1222,1249]" fontSize="10" level="2" pageId="7" pageNumber="508" reason="2">
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<emphasis id="15B7EAD1436B6177FF6ADFB64A683EB3" bold="true" box="[151,608,1222,1249]" pageId="7" pageNumber="508">
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3.
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<taxonomicName id="E0C34D40436B6177FF48DFB64A683EB3" ID-CoL="WQD3" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[181,608,1222,1249]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="pyriformis">
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<emphasis id="15B7EAD1436B6177FF48DFB649A13EB3" bold="true" box="[181,425,1222,1249]" italics="true" pageId="7" pageNumber="508">Coeliccia pyriformis</emphasis>
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<bibRefCitation id="43524B32436B6177FE4CDFB64A683EB3" author="Laidlaw, F. F." box="[433,608,1222,1248]" pageId="7" pageNumber="508" pagination="7 - 43" refId="ref18880" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 43." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6FD96548436B6177FF6ADE7B4A9B3F71" box="[151,659,1291,1315]" pageId="7" pageNumber="508" type="synonymic_list">
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<paragraph id="277C36C3436B6177FF6ADE7B4A9B3F71" blockId="7.[151,659,1291,1315]" box="[151,659,1291,1315]" pageId="7" pageNumber="508">
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Syn
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<taxonomicName id="E0C34D40436B6177FF38DE7B4A033F70" authority="Fraser, 1933" authorityName="sensu Fraser" authorityYear="1933" box="[197,523,1291,1315]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="15B7EAD1436B6177FF38DE7B498D3F71" box="[197,389,1291,1315]" italics="true" pageId="7" pageNumber="508">Coeliccia montana</emphasis>
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<bibRefCitation id="43524B32436B6177FE76DE7B4A033F70" author="Fraser, F. C." box="[395,523,1291,1315]" pageId="7" pageNumber="508" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser, 1933</bibRefCitation>
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</taxonomicName>
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, syn. novum.
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</paragraph>
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</subSubSection>
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<subSubSection id="6FD96548436B617DFF6ADE214ACF3FB4" lastPageId="13" lastPageNumber="514" pageId="7" pageNumber="508" type="discussion">
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<paragraph id="277C36C3436B6177FF6ADE2148E13C34" blockId="7.[151,1437,1361,2035]" pageId="7" pageNumber="508">
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Fraser described
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<taxonomicName id="E0C34D40436B6177FEABDE214A223F38" authorityName="sensu Fraser" authorityYear="1933" box="[342,554,1361,1387]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="15B7EAD1436B6177FEABDE214A223F38" box="[342,554,1361,1387]" italics="true" pageId="7" pageNumber="508">Coeliccia montana</emphasis>
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</taxonomicName>
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in 1933 on the basis of a male specimen collected in 1932 by Dr. A. Kerr from Muang Huang in Lao DPR (
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<bibRefCitation id="43524B32436B6177FE24DE054A6D3FDC" author="Fraser, F. C." box="[473,613,1397,1423]" pageId="7" pageNumber="508" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser 1933</bibRefCitation>
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). The description is quite extensive, although not accompanied by drawings. It mentions, amongst other characters, a black prothorax with the sides of middle pronotal lobe pale blue, changing to creamy yellow below. The synthorax is dorsally black with narrow irregular antehumeral stripes, “the inner border of which is quite straight, but the outer very irregular; laterally pale blue with a narrow black stripe on the postero-lateral suture.” The abdomen segments S9–10 are “broadly pale blue on dorsum and sides, or pale creamy yellow in the young state, the lower parts of sides black.” The anal appendages are described as creamy yellow.
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</paragraph>
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<paragraph id="277C36C3436B6177FF3ADD014C903CFC" blockId="7.[151,1437,1361,2035]" pageId="7" pageNumber="508">The statement about the color of S9–10 is surprising. Fraser described a single specimen. It is difficult to understand how he could infer a color difference between the adult insect and its immature stage.</paragraph>
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<paragraph id="277C36C3436B6177FF3ADDC9491B3DA0" blockId="7.[151,1437,1361,2035]" pageId="7" pageNumber="508">
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<bibRefCitation id="43524B32436B6177FF3ADDC9497F3C87" author="Asahina, S." box="[199,375,1721,1748]" pageId="7" pageNumber="508" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">Asahina (1969)</bibRefCitation>
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recorded a male of
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<taxonomicName id="E0C34D40436B6177FD9CDDC94B3F3C80" authorityName="sensu Fraser" authorityYear="1933" box="[609,823,1721,1747]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="15B7EAD1436B6177FD9CDDC94B3F3C80" box="[609,823,1721,1747]" italics="true" pageId="7" pageNumber="508">Coeliccia montana</emphasis>
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</taxonomicName>
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from “
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<quantity id="E03B9B26436B6177FC76DDC94C073C80" box="[907,1039,1721,1747]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="508" unit="km" value="7.0">7 kilometer</quantity>
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east, Postal, de M. dark,
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<collectingCountry id="5FD47653436B6177FACBDDC94D903C80" box="[1334,1432,1721,1747]" name="Vietnam" pageId="7" pageNumber="508">Vietnam</collectingCountry>
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,
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<quantity id="E03B9B26436B6177FF6ADDAC48E83CA4" box="[151,224,1756,1783]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="508" unit="m" value="500.0">500 m</quantity>
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” collected by Mr. Yoshimoto and preserved in the Bernice P. Bishop Museum in Honolulu. Asahina provides three drawings and a short description. What stands out is that this is “a slender pale yellow and black species” and that S2–9 are “black on the dorsal side; end of 9 and whole the 10 […] orange yellow.”
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<figureCitation id="BFF82A46436B6177FBA7DC554CB23D6C" box="[1114,1210,1829,1855]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,250,1615,1640]" captionStartId-1="6.[151,250,1882,1907]" captionTargetBox-0="[151,1436,633,1590]" captionTargetBox-1="[151,1436,417,1857]" captionTargetId-0="figure@4.[151,1436,633,1590]" captionTargetId-1="figure@6.[151,1436,417,1857]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 2. Prothorax of male Coeliccia spp. in lateral view. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype." captionText-1="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi-0="http://doi.org/10.5281/zenodo.3765656" figureDoi-1="http://doi.org/10.5281/zenodo.3765658" httpUri-0="https://zenodo.org/record/3765656/files/figure.png" httpUri-1="https://zenodo.org/record/3765658/files/figure.png" pageId="7" pageNumber="508">Figs 2–3</figureCitation>
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(
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<bibRefCitation id="43524B32436B6177FB3ADC554D6E3D6C" author="Asahina, S." box="[1223,1382,1829,1855]" pageId="7" pageNumber="508" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">Asahina 1969</bibRefCitation>
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, p.4) confirm this: S9 is almost completely dark, apart from the posterior margin. It is therefore remarkable that Asahina states: “This single male specimen agrees rather well with the original description and the
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<typeStatus id="F8788861436B6177FB75DC1D4CB03DD4" box="[1160,1208,1901,1927]" pageId="7" pageNumber="508">type</typeStatus>
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specimen preserved in the British Museum (Nat. Hist.). The differences are: 1) the length of abdomen which is stated as “
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<quantity id="E03B9B26436B6177FAE2DCE14D7C3DF8" box="[1311,1396,1937,1963]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="508" unit="mm" value="42.0">42 mm</quantity>
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” in the original description is shorter […], 2) the coloration of abdomen 9 and 10 is said to be “pale dorsally and dark laterally”.”
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</paragraph>
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<caption id="73BC664B43646178FF6AD8FE49EE3998" ID-DOI="http://doi.org/10.5281/zenodo.3765662" ID-Zenodo-Dep="3765662" httpUri="https://zenodo.org/record/3765662/files/figure.png" pageId="8" pageNumber="509" startId="8.[151,250,910,935]" targetBox="[151,1436,181,885]" targetPageId="8">
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<paragraph id="277C36C343646178FF6AD8FE49EE3998" blockId="8.[151,1436,910,971]" pageId="8" pageNumber="509">
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<emphasis id="15B7EAD143646178FF6AD8FE491A39F4" bold="true" box="[151,274,910,935]" pageId="8" pageNumber="509">FIGURE 5.</emphasis>
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Thorax of male
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<taxonomicName id="E0C34D4043646178FE4AD8FF4A1039F4" authorityName="Kirby" authorityYear="1890" box="[439,536,911,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="509" phylum="Arthropoda" rank="genus">
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<emphasis id="15B7EAD143646178FE4AD8FF4A1039F4" box="[439,536,911,935]" italics="true" pageId="8" pageNumber="509">Coeliccia</emphasis>
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</taxonomicName>
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spp. in dorsal view. (a)
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<taxonomicName id="E0C34D4043646178FCF8D8FF4B7339F4" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[773,891,911,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="caerulea">
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<emphasis id="15B7EAD143646178FCF8D8FF4B7339F4" box="[773,891,911,935]" italics="true" pageId="8" pageNumber="509">C. caerulea</emphasis>
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</taxonomicName>
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, holotype; (b)
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<taxonomicName id="E0C34D4043646178FBF3D8FF4C8D39F4" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[1038,1157,911,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="coronata">
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<emphasis id="15B7EAD143646178FBF3D8FF4C8D39F4" box="[1038,1157,911,935]" italics="true" pageId="8" pageNumber="509">C. coronata</emphasis>
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</taxonomicName>
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, holotype; (c)
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<taxonomicName id="E0C34D4043646178FAEAD8FF4D9D39F4" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[1303,1429,911,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="diomedea">
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<emphasis id="15B7EAD143646178FAEAD8FF4D9D39F4" box="[1303,1429,911,935]" italics="true" pageId="8" pageNumber="509">C. diomedea</emphasis>
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</taxonomicName>
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, holotype; (d)
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<taxonomicName id="E0C34D4043646178FEDCD8C349723998" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[289,378,947,971]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="curua">
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<emphasis id="15B7EAD143646178FEDCD8C349723998" box="[289,378,947,971]" italics="true" pageId="8" pageNumber="509">C. curua</emphasis>
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</taxonomicName>
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, holotype.
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</paragraph>
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</caption>
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<caption id="73BC664B43646178FF6ADCF349B23D93" ID-DOI="http://doi.org/10.5281/zenodo.3765664" ID-Zenodo-Dep="3765664" httpUri="https://zenodo.org/record/3765664/files/figure.png" pageId="8" pageNumber="509" startId="8.[151,250,1923,1948]" targetBox="[151,1436,993,1898]" targetPageId="8">
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<paragraph id="277C36C343646178FF6ADCF349B23D93" blockId="8.[151,1436,1923,1984]" pageId="8" pageNumber="509">
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<emphasis id="15B7EAD143646178FF6ADCF3491B3DCF" bold="true" box="[151,275,1923,1948]" pageId="8" pageNumber="509">FIGURE 6.</emphasis>
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Holotype of
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<taxonomicName id="E0C34D4043646178FE64DCF44A563DCF" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[409,606,1924,1948]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="pulchella">
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<emphasis id="15B7EAD143646178FE64DCF44A563DCF" box="[409,606,1924,1948]" italics="true" pageId="8" pageNumber="509">Coeliccia pulchella</emphasis>
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</taxonomicName>
|
||
(a) thorax in lateral view; (b) thorax in dorsal view; (c) head in dorsolateral view; (d) prothorax in lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="73BC664B43656179FF6ADC654D923D01" ID-DOI="http://doi.org/10.5281/zenodo.3765666" ID-Zenodo-Dep="3765666" httpUri="https://zenodo.org/record/3765666/files/figure.png" pageId="9" pageNumber="510" startId="9.[151,250,1813,1838]" targetBox="[151,1436,181,1786]" targetPageId="9">
|
||
<paragraph id="277C36C343656179FF6ADC654D923D01" blockId="9.[151,1437,1813,1874]" pageId="9" pageNumber="510">
|
||
<emphasis id="15B7EAD143656179FF6ADC65491A3D7D" bold="true" box="[151,274,1813,1838]" pageId="9" pageNumber="510">FIGURE 7.</emphasis>
|
||
Abdomen tip of male
|
||
<taxonomicName id="E0C34D4043656179FE0EDC654A5C3D7E" authorityName="Kirby" authorityYear="1890" box="[499,596,1813,1837]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="15B7EAD143656179FE0EDC654A5C3D7E" box="[499,596,1813,1837]" italics="true" pageId="9" pageNumber="510">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
spp. [a,e,g,i] lateral view, [c] dorsolateral view, [b,d,f,h,j] dorsal view. (a,b)
|
||
<taxonomicName id="E0C34D4043656179FAA9DC6648C43D02" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="caerulea">
|
||
<emphasis id="15B7EAD143656179FAA9DC6648C43D02" italics="true" pageId="9" pageNumber="510">C. caerulea</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (c,d)
|
||
<taxonomicName id="E0C34D4043656179FE88DC4A49F83D02" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[373,496,1849,1873]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="15B7EAD143656179FE88DC4A49F83D02" box="[373,496,1849,1873]" italics="true" pageId="9" pageNumber="510">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (e,f)
|
||
<taxonomicName id="E0C34D4043656179FD68DC4A4AE53D02" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[661,749,1850,1873]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="curua">
|
||
<emphasis id="15B7EAD143656179FD68DC4A4AE53D02" box="[661,749,1850,1873]" italics="true" pageId="9" pageNumber="510">C. curua</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (g,h)
|
||
<taxonomicName id="E0C34D4043656179FC6ADC4A4C073D02" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[919,1039,1850,1873]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="coronata">
|
||
<emphasis id="15B7EAD143656179FC6ADC4A4C073D02" box="[919,1039,1850,1873]" italics="true" pageId="9" pageNumber="510">C. coronata</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (i,j)
|
||
<taxonomicName id="E0C34D4043656179FB50DC4A4D253D02" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[1197,1325,1849,1873]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="diomedea">
|
||
<emphasis id="15B7EAD143656179FB50DC4A4D253D02" box="[1197,1325,1849,1873]" italics="true" pageId="9" pageNumber="510">C. diomedea</emphasis>
|
||
</taxonomicName>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="73BC664B4366617AFF6ADC404D113D3E" ID-DOI="http://doi.org/10.5281/zenodo.3765668" ID-Zenodo-Dep="3765668" httpUri="https://zenodo.org/record/3765668/files/figure.png" pageId="10" pageNumber="511" startId="10.[151,250,1840,1865]" targetBox="[166,1421,181,1813]" targetPageId="10">
|
||
<paragraph id="277C36C34366617AFF6ADC404D113D3E" blockId="10.[151,1437,1840,1901]" pageId="10" pageNumber="511">
|
||
<emphasis id="15B7EAD14366617AFF6ADC40491B3D1A" bold="true" box="[151,275,1840,1865]" pageId="10" pageNumber="511">FIGURE 8.</emphasis>
|
||
S10 and caudal appendages of male
|
||
<taxonomicName id="E0C34D404366617AFD75DC414AE13D1A" authorityName="Kirby" authorityYear="1890" box="[648,745,1841,1865]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="511" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="15B7EAD14366617AFD75DC414AE13D1A" box="[648,745,1841,1865]" italics="true" pageId="10" pageNumber="511">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
spp. [a,c,e,] lateral view, [g] dorsolateral view, [b,d,f,h] dorsal view. (a,b)
|
||
<taxonomicName id="E0C34D404366617AFF30DC25494D3D3E" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[205,325,1877,1901]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="coronata">
|
||
<emphasis id="15B7EAD14366617AFF30DC25494D3D3E" box="[205,325,1877,1901]" italics="true" pageId="10" pageNumber="511">C. coronata</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (c,d)
|
||
<taxonomicName id="E0C34D404366617AFE10DC254A643D3E" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[493,620,1877,1901]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="diomedea">
|
||
<emphasis id="15B7EAD14366617AFE10DC254A643D3E" box="[493,620,1877,1901]" italics="true" pageId="10" pageNumber="511">C. diomedea</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (e,f)
|
||
<taxonomicName id="E0C34D404366617AFCECDC254B8F3D3E" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[785,903,1877,1901]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="caerulea">
|
||
<emphasis id="15B7EAD14366617AFCECDC254B8F3D3E" box="[785,903,1877,1901]" italics="true" pageId="10" pageNumber="511">C. caerulea</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (g,h)
|
||
<taxonomicName id="E0C34D404366617AFBCCDC254CA43D3E" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[1073,1196,1877,1901]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="15B7EAD14366617AFBCCDC254CA43D3E" box="[1073,1196,1877,1901]" italics="true" pageId="10" pageNumber="511">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="73BC664B4367617BFF6ADD634C133C03" ID-DOI="http://doi.org/10.5281/zenodo.3765670" ID-Zenodo-Dep="3765670" httpUri="https://zenodo.org/record/3765670/files/figure.png" pageId="11" pageNumber="512" startId="11.[151,250,1555,1580]" targetBox="[151,1436,181,1529]" targetPageId="11">
|
||
<paragraph id="277C36C34367617BFF6ADD634C133C03" blockId="11.[151,1437,1555,1616]" pageId="11" pageNumber="512">
|
||
<emphasis id="15B7EAD14367617BFF6ADD63491C3C7F" bold="true" box="[151,276,1555,1580]" pageId="11" pageNumber="512">FIGURE 9.</emphasis>
|
||
S10 and caudal appendages of male
|
||
<taxonomicName id="E0C34D404367617BFD6FDD634B3D3C78" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[658,821,1555,1579]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="curua">
|
||
<emphasis id="15B7EAD14367617BFD6FDD634B3D3C78" box="[658,821,1555,1579]" italics="true" pageId="11" pageNumber="512">Coeliccia curua</emphasis>
|
||
</taxonomicName>
|
||
from Xuan Son NP. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b) paratype, 23 vii 2017; (c,d) paratype, 9 vi 2018; (e,f) second paratype, 23 vii 2017.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="277C36C34367617BFF3ADD0E4D8F3D7B" blockId="11.[151,1436,1662,2013]" pageId="11" pageNumber="512">
|
||
This is remarkable, because 1) Fraser was clear about the blue coloration on head, prothorax and thorax. Nothing in his description suggests that
|
||
<taxonomicName id="E0C34D404367617BFDD5DDD24AA43CEF" authorityName="sensu Fraser" authorityYear="1933" box="[552,684,1698,1724]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14367617BFDD5DDD24AA43CEF" box="[552,684,1698,1724]" italics="true" pageId="11" pageNumber="512">C. montana</emphasis>
|
||
</taxonomicName>
|
||
is a ‘pale yellow and black species’ and 2) Fraser was clear about the fact that S9–10 were dorsally completely pale blue (or creamy yellow). He does not state pale dorsally and dark laterally as such, but the pattern of S9 surely was different from that illustrated by
|
||
<bibRefCitation id="43524B324367617BFBCBDD9A4CEF3D56" author="Asahina, S." box="[1078,1255,1770,1797]" pageId="11" pageNumber="512" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">Asahina (1969)</bibRefCitation>
|
||
. On the basis of these facts there seems to be no clear reason to identify the specimen collected by Mr. Yoshimoto as
|
||
<taxonomicName id="E0C34D404367617BFB00DC7E4D883D7B" authorityName="sensu Fraser" authorityYear="1933" box="[1277,1408,1806,1832]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14367617BFB00DC7E4D883D7B" box="[1277,1408,1806,1832]" italics="true" pageId="11" pageNumber="512">C. montana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="277C36C34367617CFF3ADC424C743B10" blockId="11.[151,1436,1662,2013]" lastBlockId="12.[151,1437,153,323]" lastPageId="12" lastPageNumber="513" pageId="11" pageNumber="512">
|
||
Subsequently,
|
||
<bibRefCitation id="43524B324367617BFE97DC424A623D1F" author="Steinhoff, P. O. M. & Do, C. M." box="[362,618,1842,1868]" pageId="11" pageNumber="512" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
followed the interpretation by
|
||
<bibRefCitation id="43524B324367617BFC39DC424C793D1E" author="Asahina, S." box="[964,1137,1842,1869]" pageId="11" pageNumber="512" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">Asahina (1969)</bibRefCitation>
|
||
and provided more details of a
|
||
<taxonomicName id="E0C34D404367617BFF30DC2649303D23" authorityName="Kirby" authorityYear="1890" box="[205,312,1878,1904]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="15B7EAD14367617BFF30DC2649303D23" box="[205,312,1878,1904]" italics="true" pageId="11" pageNumber="512">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
species that shares characteristics with both the illustrations provided by Asahina and his description. In their article,
|
||
<bibRefCitation id="43524B324367617BFE7DDC0A4A893DC7" author="Steinhoff, P. O. M. & Do, C. M." box="[384,641,1914,1940]" pageId="11" pageNumber="512" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
note that: “Here we compare the characters of our specimens with the characters described by Fraser.” They write that: “Thorax markings and coloration almost identical as described by
|
||
<bibRefCitation id="43524B324367617BFF6ADCB2495B3D8F" author="Fraser, F. C." box="[151,339,1986,2012]" pageId="11" pageNumber="512" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">FRASER (1933)</bibRefCitation>
|
||
and drawn by
|
||
<bibRefCitation id="43524B324367617BFE05DCB24AC13D8E" author="Asahina, S." box="[504,713,1986,2013]" pageId="11" pageNumber="512" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">ASAHINA (1969)</bibRefCitation>
|
||
. Regarding the mesepisternum,
|
||
<bibRefCitation id="43524B324367617BFBCCDCB24D2C3D8E" author="Fraser, F. C." box="[1073,1316,1986,2013]" pageId="11" pageNumber="512" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">FRASER (1933: 116)</bibRefCitation>
|
||
wrote that “... the outer border of the antehumeral stripes is very irregular”. We found the antehumeral stripes resembling a pair of paddles, with indeed the outer (proximal) border less straight than the distal border, forming the “blade of the paddle” (
|
||
<figureCitation id="BFF82A464360617CFED4DB9149763AAF" box="[297,382,225,252]" captionStart="FIGURE 3" captionStartId="6.[151,250,1882,1907]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure@6.[151,1436,417,1857]" captionTargetPageId="6" captionText="FIGURE 3. Prothorax of Coeliccia spp. in dorsal view. [a-e] male, [f] female. (a) C. caerulea, holotype; (b) C. coronata, holotype; (c) C. diomedea, holotype; (d) C. curua, holotype; (e) C. pulchella sp. nov., holotype; (f) C. caerulea, paratype used for description." figureDoi="http://doi.org/10.5281/zenodo.3765658" httpUri="https://zenodo.org/record/3765658/files/figure.png" pageId="12" pageNumber="513">Fig. 3b</figureCitation>
|
||
). This specific marking pattern is also visible in Asahina’s drawing.” In addition
|
||
<bibRefCitation id="43524B324360617CFAEADB9149033B4C" author="Steinhoff, P. O. M. & Do, C. M." pageId="12" pageNumber="513" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
note that the abdomen and anal appendages are as described by
|
||
<bibRefCitation id="43524B324360617CFC1CDA754C743B4C" author="Fraser, F. C." box="[993,1148,261,287]" pageId="12" pageNumber="513" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser (1933)</bibRefCitation>
|
||
. However, their drawings show S9 with only limited yellow towards the posterior margin, as in Asahina’s drawing.
|
||
</paragraph>
|
||
<caption id="73BC664B4360617CFF6ADD8C4C623D6A" ID-DOI="http://doi.org/10.5281/zenodo.3765672" ID-Zenodo-Dep="3765672" httpUri="https://zenodo.org/record/3765672/files/figure.png" pageId="12" pageNumber="513" startId="12.[151,250,1788,1813]" targetBox="[151,1436,345,1763]" targetPageId="12">
|
||
<paragraph id="277C36C34360617CFF6ADD8C4C623D6A" blockId="12.[151,1436,1788,1849]" pageId="12" pageNumber="513">
|
||
<emphasis id="15B7EAD14360617CFF6ADD8C49293D46" bold="true" box="[151,289,1788,1813]" pageId="12" pageNumber="513">FIGURE 10.</emphasis>
|
||
S10 and caudal appendages of male
|
||
<taxonomicName id="E0C34D404360617CFD63DD8C4AF73D47" authorityName="Kirby" authorityYear="1890" box="[670,767,1788,1812]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="513" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="15B7EAD14360617CFD63DD8C4AF73D47" box="[670,767,1788,1812]" italics="true" pageId="12" pageNumber="513">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
spp. [a,c,e] lateral view, [b,d,f] dorsal view. (a,b)
|
||
<taxonomicName id="E0C34D404360617CFAFDDD8D4D513D47" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[1280,1369,1789,1812]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="curua">
|
||
<emphasis id="15B7EAD14360617CFAFDDD8D4D513D47" box="[1280,1369,1789,1812]" italics="true" pageId="12" pageNumber="513">C. curua</emphasis>
|
||
</taxonomicName>
|
||
, holotype; (c,d)
|
||
<taxonomicName id="E0C34D404360617CFEFBDC5149563D6B" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[262,350,1825,1848]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="curua">
|
||
<emphasis id="15B7EAD14360617CFEFBDC5149563D6B" box="[262,350,1825,1848]" italics="true" pageId="12" pageNumber="513">C. curua</emphasis>
|
||
</taxonomicName>
|
||
, paratype, 2 vii 2017; (e,f)
|
||
<taxonomicName id="E0C34D404360617CFD89DC514B3D3D6A" authority=", Xuan Son" authorityName="Xuan Son" box="[628,821,1825,1849]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="uenoi">
|
||
<emphasis id="15B7EAD14360617CFD89DC514ACC3D6B" box="[628,708,1825,1848]" italics="true" pageId="12" pageNumber="513">C.uenoi</emphasis>
|
||
, Xuan Son
|
||
</taxonomicName>
|
||
NP, Phu Tho Prov., 9 vi 2018.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="277C36C34360617DFF3ADC174C203BFC" blockId="12.[151,1437,1895,2029]" lastBlockId="13.[151,1437,153,1511]" lastPageId="13" lastPageNumber="514" pageId="12" pageNumber="513">
|
||
Although
|
||
<bibRefCitation id="43524B324360617CFEC6DC174A4C3DD2" author="Steinhoff, P. O. M. & Do, C. M." box="[315,580,1895,1921]" pageId="12" pageNumber="513" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
stated that they compared their specimens with the description by Fraser, clearly they are more in line with Asahina’s specimen. The coloration of the specimens studied by
|
||
<bibRefCitation id="43524B324360617CFAFEDCFB49063D9A" author="Steinhoff, P. O. M. & Do, C. M." pageId="12" pageNumber="513" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff and Do (2013)</bibRefCitation>
|
||
was whitish-yellow and black, but because information regarding the age of their specimens and the coloration in life was missing, the authors did not take the coloration into account (Philip Steinhoff, pers. comm.). Colors often fade in preserved specimens, and immature individuals of
|
||
<taxonomicName id="E0C34D404361617DFC4BDBE94C293AE0" authorityName="Kirby" authorityYear="1890" box="[950,1057,153,179]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="15B7EAD14361617DFC4BDBE94C293AE0" box="[950,1057,153,179]" italics="true" pageId="13" pageNumber="514">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
species often display other colors than the adults (
|
||
<bibRefCitation id="43524B324361617DFEB3DBCD49FB3A84" author="Kosterin, O. E." box="[334,499,189,215]" pageId="13" pageNumber="514" pagination="1 - 108" refId="ref18741" refString="Kosterin, O. E. (2011) Odonata of the Cambodian coastal regions revisited: beginning of dry season in 2010. International Dragonfly Fund - Report, 40, 1 - 108." type="journal article" year="2011">Kosterin 2011</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="43524B324361617DFDFDDBCD4A973A84" author="Laidlaw, F. F." box="[512,671,189,215]" pageId="13" pageNumber="514" pagination="7 - 43" refId="ref18880" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 43." type="journal article" year="1932">Laidlaw 1932</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="43524B324361617DFD51DBCD4BA23A84" author="Steinhoff, P. O. M. & Uhl, G." box="[684,938,188,215]" pageId="13" pageNumber="514" pagination="257 - 276" refId="ref19435" refString="Steinhoff, P. O. M. & Uhl, G. (2015) Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis. Zootaxa, 4059 (2), 257 - 276. https: // doi. org / 10.11646 / zootaxa. 4059.2.2" type="journal article" year="2015">Steinhoff & Uhl 2015</bibRefCitation>
|
||
). Thus, in their comparison
|
||
<bibRefCitation id="43524B324361617DFB16DBCD48EA3AA8" author="Steinhoff, P. O. M. & Do, C. M." pageId="13" pageNumber="514" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
did not take into consideration that 1) Fraser had described the sides of the thorax as pale blue, 2) that he indicated that S9 was completely pale blue or yellow dorsally, 3) although they called the proximal border of the antehumeral stripe the outer border, this must be the opposite of what Fraser meant. Clearly he referred to the outer (=distal) border as more irregular. Especially when taking the shape of the superior appendages into consideration, it seems reasonable that the specimens described by
|
||
<bibRefCitation id="43524B324361617DFD1ADA014BE03BD8" author="Steinhoff, P. O. M. & Do, C. M." box="[743,1000,369,395]" pageId="13" pageNumber="514" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
concern the same species as described by
|
||
<bibRefCitation id="43524B324361617DFF44DAE549603BE3" author="Asahina, S." box="[185,360,405,432]" pageId="13" pageNumber="514" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">Asahina (1969)</bibRefCitation>
|
||
, but it is clearly different from
|
||
<taxonomicName id="E0C34D404361617DFD3ADAE54C2B3BFC" authority="sensu Fraser, 1933" authorityName="sensu Fraser" authorityYear="1933" box="[711,1059,405,431]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFD3ADAE54B413BFC" box="[711,841,405,431]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
sensu
|
||
<bibRefCitation id="43524B324361617DFC68DAE54C2B3BFC" author="Fraser, F. C." box="[917,1059,405,431]" pageId="13" pageNumber="514" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="277C36C34361617DFF3ADAC94A0F38A0" blockId="13.[151,1437,153,1511]" pageId="13" pageNumber="514">
|
||
In order to verify the characteristics of
|
||
<taxonomicName id="E0C34D404361617DFD71DAC94B193B80" authorityName="sensu Fraser" authorityYear="1933" box="[652,785,441,467]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFD71DAC94B193B80" box="[652,785,441,467]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
</taxonomicName>
|
||
we obtained photos of the
|
||
<typeStatus id="F87888614361617DFBB0DAC94CA73B80" box="[1101,1199,441,467]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
|
||
stored in the British Museum of Natural History (
|
||
<figureCitation id="BFF82A464361617DFE1BDAAD4A323BA4" box="[486,570,477,503]" captionStart="FIGURE 23" captionStartId="28.[151,250,1893,1918]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure@28.[151,1436,181,1868]" captionTargetPageId="28" captionText="FIGURE 23. Holotype of Coeliccia montana Fraser, 1932. Photos courtesy of Dr. Ben Price, Natural History Museum, London. (a) thorax and head in lateral view; (b) abdomen tip and caudal appendages in lateral view; (c) thorax in dorsal view; (d) type on card with labels." figureDoi="http://doi.org/10.5281/zenodo.3765698" httpUri="https://zenodo.org/record/3765698/files/figure.png" pageId="13" pageNumber="514">Fig. 23</figureCitation>
|
||
) through kind help from Dr. Ben Price. We could ascertain that (
|
||
<figureCitation id="BFF82A464361617DFAD7DAAD4D983BAB" box="[1322,1424,477,504]" captionStart="FIGURE 23" captionStartId="28.[151,250,1893,1918]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure@28.[151,1436,181,1868]" captionTargetPageId="28" captionText="FIGURE 23. Holotype of Coeliccia montana Fraser, 1932. Photos courtesy of Dr. Ben Price, Natural History Museum, London. (a) thorax and head in lateral view; (b) abdomen tip and caudal appendages in lateral view; (c) thorax in dorsal view; (d) type on card with labels." figureDoi="http://doi.org/10.5281/zenodo.3765698" httpUri="https://zenodo.org/record/3765698/files/figure.png" pageId="13" pageNumber="514">Fig. 23b</figureCitation>
|
||
): 1) S9–10 are indeed dorsally completely pale, with only the lower sides black; 2) the color is a mix of paler blue and ochre, although ochre is the overriding impression; 3) the superior appendages have an almost square ventral expansion well proximal of the apex of the appendages, (
|
||
<figureCitation id="BFF82A464361617DFCE8D9394B7D3830" box="[789,885,585,611]" captionStart="FIGURE 23" captionStartId="28.[151,250,1893,1918]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure@28.[151,1436,181,1868]" captionTargetPageId="28" captionText="FIGURE 23. Holotype of Coeliccia montana Fraser, 1932. Photos courtesy of Dr. Ben Price, Natural History Museum, London. (a) thorax and head in lateral view; (b) abdomen tip and caudal appendages in lateral view; (c) thorax in dorsal view; (d) type on card with labels." figureDoi="http://doi.org/10.5281/zenodo.3765698" httpUri="https://zenodo.org/record/3765698/files/figure.png" pageId="13" pageNumber="514">Fig. 23a</figureCitation>
|
||
); 4) the sides of the thorax are clearly blue with a thin black line over the metapleural suture, (
|
||
<figureCitation id="BFF82A464361617DFD79D91D4AF538D4" box="[644,765,621,647]" captionStart="FIGURE 23" captionStartId="28.[151,250,1893,1918]" captionTargetBox="[151,1436,181,1868]" captionTargetId="figure@28.[151,1436,181,1868]" captionTargetPageId="28" captionText="FIGURE 23. Holotype of Coeliccia montana Fraser, 1932. Photos courtesy of Dr. Ben Price, Natural History Museum, London. (a) thorax and head in lateral view; (b) abdomen tip and caudal appendages in lateral view; (c) thorax in dorsal view; (d) type on card with labels." figureDoi="http://doi.org/10.5281/zenodo.3765698" httpUri="https://zenodo.org/record/3765698/files/figure.png" pageId="13" pageNumber="514">Fig. 23a, c</figureCitation>
|
||
); 5) the mesepisternum has a long antehumeral stripe that in dorsal view appears somewhat curved; 6) the middle pronotal lobe of the prothorax has large yellow areas laterally and the propleuron is largely yellowish, the anterior and posterior pronotal lobes connected by black over the middle of the middle pronotal lobe only.
|
||
</paragraph>
|
||
<paragraph id="277C36C34361617DFF3AD98D49B939D0" blockId="13.[151,1437,153,1511]" pageId="13" pageNumber="514">
|
||
These points confirm that the
|
||
<taxonomicName id="E0C34D404361617DFDE2D98D4AAB3944" authorityName="sensu Fraser" authorityYear="1933" box="[543,675,765,791]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFDE2D98D4AAB3944" box="[543,675,765,791]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
</taxonomicName>
|
||
described in
|
||
<bibRefCitation id="43524B324361617DFCC1D98D4BE3394B" author="Asahina, S." box="[828,1003,765,792]" pageId="13" pageNumber="514" pagination="1 - 19" refId="ref18315" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16, 1 - 19." type="journal article" year="1969">Asahina (1969)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="43524B324361617DFBD8D98D4D223944" author="Steinhoff, P. O. M. & Do, C. M." box="[1061,1322,765,791]" pageId="13" pageNumber="514" pagination="347 - 357" refId="ref19396" refString="Steinhoff, P. O. M. & Do, C. M. (2013) Notes on some Coeliccia species from Vietnam (Zygoptera: Platycnemididae). Odonatologica, 42, 347 - 357." type="journal article" year="2013">Steinhoff & Do (2013)</bibRefCitation>
|
||
is not the same species as
|
||
<taxonomicName id="E0C34D404361617DFEB2D8514AA53968" authority="sensu Fraser, 1933" authorityName="sensu Fraser" authorityYear="1933" box="[335,685,801,827]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFEB2D85149DA3968" box="[335,466,801,827]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
sensu
|
||
<bibRefCitation id="43524B324361617DFDE2D8514AA53968" author="Fraser, F. C." box="[543,685,801,827]" pageId="13" pageNumber="514" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
. Most likely their specimens refer to
|
||
<taxonomicName id="E0C34D404361617DFBACD8514CDE3968" authority="Kompier & Dow & Steinhoff, 2020" authorityName="Kompier & Dow & Steinhoff" authorityYear="2020" box="[1105,1238,801,827]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="coronata" status="sp. nov.">
|
||
<emphasis id="15B7EAD14361617DFBACD8514CDE3968" box="[1105,1238,801,827]" italics="true" pageId="13" pageNumber="514">C. coronata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15B7EAD14361617DFB20D8514D3F3968" bold="true" box="[1245,1335,801,827]" pageId="13" pageNumber="514">
|
||
<taxonomicNameLabel id="0E8457AA4361617DFB20D8514D3F3968" box="[1245,1335,801,827]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The mix of the apparent color of S9–10 may have led Fraser to infer that they would be blue in the adult and yellow in the immature of
|
||
<taxonomicName id="E0C34D404361617DFED5D81949A239D0" authorityName="sensu Fraser" authorityYear="1933" box="[296,426,873,899]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFED5D81949A239D0" box="[296,426,873,899]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="277C36C34361617DFF3AD8FD4ACF3FB4" blockId="13.[151,1437,153,1511]" pageId="13" pageNumber="514">
|
||
The characteristics of the
|
||
<typeStatus id="F87888614361617DFE19D8FD4A4E39F4" box="[484,582,909,935]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E0C34D404361617DFD94D8FD4AE339F4" authorityName="sensu Fraser" authorityYear="1933" box="[617,747,909,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFD94D8FD4AE339F4" box="[617,747,909,935]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
</taxonomicName>
|
||
in fact exactly match those of
|
||
<taxonomicName id="E0C34D404361617DFBC6D8FD4CD839F4" authorityName="Laidlaw" authorityYear="1932" box="[1083,1232,909,935]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="15B7EAD14361617DFBC6D8FD4CD839F4" box="[1083,1232,909,935]" italics="true" pageId="13" pageNumber="514">C. pyriformis</emphasis>
|
||
</taxonomicName>
|
||
, the type of which was collected in 1924 from Bao Ha in
|
||
<collectingCountry id="5FD476534361617DFDAAD8C14AAE3998" box="[599,678,945,971]" name="Vietnam" pageId="13" pageNumber="514">Tonkin</collectingCountry>
|
||
(present day Bao Ha Commune, Bao Yen District,
|
||
<collectingRegion id="E507F8214361617DFB0AD8C14D913998" box="[1271,1433,945,971]" country="Vietnam" name="Lao Cai" pageId="13" pageNumber="514">Lao Cai Prov.</collectingRegion>
|
||
, northern
|
||
<collectingCountry id="5FD476534361617DFF01D8A5496839A3" box="[252,352,981,1008]" name="Vietnam" pageId="13" pageNumber="514">Vietnam</collectingCountry>
|
||
). However, this is not immediately clear from the description.
|
||
<bibRefCitation id="43524B324361617DFBE8D8A54CCA39BC" author="Laidlaw, F. F." box="[1045,1218,981,1007]" pageId="13" pageNumber="514" pagination="7 - 43" refId="ref18880" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 43." type="journal article" year="1932">Laidlaw (1932)</bibRefCitation>
|
||
states: “
|
||
<emphasis id="15B7EAD14361617DFADCD8A54D9439BC" box="[1313,1436,981,1007]" italics="true" pageId="13" pageNumber="514">Synthorax:</emphasis>
|
||
black, the black extending on to the sides and fading gradually to a creamy white below. A pyriform mark of pale blue on either side on the dorsum with its pointed end upwards. (This mark has faded whilst the specimen was undergoing relaxation for setting, and is now not evident in the type, the whole dorsum being uniformly black).” He continues: “
|
||
<emphasis id="15B7EAD14361617DFEE3DF1549E13E2C" box="[286,489,1125,1151]" italics="true" pageId="13" pageNumber="514">Anal appendages:</emphasis>
|
||
ochreous, lower pair much longer than upper, angularly forcipate. Upper pair rather club-shaped, no marked projection.” Even if Fraser was familiar with this description, he still might have considered it clear that his specimen with bluish sides to thorax and ventral projection of the superior appendages had to be a different species. However, the reexamination of the
|
||
<typeStatus id="F87888614361617DFCE2DFA14B893EB8" box="[799,897,1233,1259]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E0C34D404361617DFC5ADFA14C343EB8" authorityName="Laidlaw" authorityYear="1932" box="[935,1084,1233,1259]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="15B7EAD14361617DFC5ADFA14C343EB8" box="[935,1084,1233,1259]" italics="true" pageId="13" pageNumber="514">C. pyriformis</emphasis>
|
||
</taxonomicName>
|
||
carried out by
|
||
<bibRefCitation id="43524B324361617DFB1BDFA148EA3F5C" author="Steinhoff, P. O. M. & Uhl, G." pageId="13" pageNumber="514" pagination="257 - 276" refId="ref19435" refString="Steinhoff, P. O. M. & Uhl, G. (2015) Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis. Zootaxa, 4059 (2), 257 - 276. https: // doi. org / 10.11646 / zootaxa. 4059.2.2" type="journal article" year="2015">Steinhoff & Uhl (2015)</bibRefCitation>
|
||
showed that Laidlaw’s description of the appendages was incorrect. They are in fact identical to those of
|
||
<taxonomicName id="E0C34D404361617DFF6ADE6949FD3F60" authority="sensu Fraser, 1933" authorityName="sensu Fraser" authorityYear="1933" box="[151,501,1305,1331]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFF6ADE6949123F60" box="[151,282,1305,1331]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
sensu
|
||
<bibRefCitation id="43524B324361617DFE9ADE6949FD3F60" author="Fraser, F. C." box="[359,501,1305,1331]" pageId="13" pageNumber="514" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
. The coloration of the thorax appears to have blackened in the preservation process not just in respect to the dorsum, but also laterally. Evidently, based on the combination of patterning and identical shape of appendages,
|
||
<taxonomicName id="E0C34D404361617DFE6DDE114AE53F28" authority="sensu Fraser, 1933" authorityName="sensu Fraser" authorityYear="1933" box="[400,749,1377,1403]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="montana">
|
||
<emphasis id="15B7EAD14361617DFE6DDE114A1B3F28" box="[400,531,1377,1403]" italics="true" pageId="13" pageNumber="514">C. montana</emphasis>
|
||
sensu
|
||
<bibRefCitation id="43524B324361617DFD9CDE114AE53F28" author="Fraser, F. C." box="[609,749,1377,1403]" pageId="13" pageNumber="514" pagination="109 - 141" refId="ref18596" refString="Fraser, F. C. (1933) Dragonflies from the Laos country. Journal of the Siam Society, Natural History, Supplement, 9, 109 - 141." type="journal article" year="1933">Fraser, 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="E0C34D404361617DFC01DE114D363F28" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[1020,1342,1377,1403]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="15B7EAD14361617DFC01DE114C9B3F28" box="[1020,1171,1377,1403]" italics="true" pageId="13" pageNumber="514">C. pyriformis</emphasis>
|
||
<bibRefCitation id="43524B324361617DFB67DE114D363F28" author="Laidlaw, F. F." box="[1178,1342,1377,1403]" pageId="13" pageNumber="514" pagination="7 - 43" refId="ref18880" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 43." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="E0C34D404361617DFAB7DE11493E3FCC" authorityName="Laidlaw" authorityYear="1932" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pyriformis">
|
||
<emphasis id="15B7EAD14361617DFAB7DE11493E3FCC" italics="true" pageId="13" pageNumber="514">Coeliccia pyriformis</emphasis>
|
||
</taxonomicName>
|
||
is in fact a common species in northern
|
||
<collectingCountry id="5FD476534361617DFD03DEF54B553FCC" box="[766,861,1413,1439]" name="Vietnam" pageId="13" pageNumber="514">Vietnam</collectingCountry>
|
||
as far south as Dong Hoi (
|
||
<bibRefCitation id="43524B324361617DFB77DEF54D993FCC" author="Kompier, T. & Phan, Q. T." box="[1162,1425,1413,1439]" pageId="13" pageNumber="514" pagination="131 - 140" refId="ref18689" refString="Kompier, T. & Phan, Q. T. (2017) Coeliccia mientrung spec. nov. from Central Vietnam (Odonata: Platycnemididae). Zootaxa, 4247 (2), 131 - 140. https: // doi. org / 10.11646 / zootaxa. 4247.2.4" type="journal article" year="2017">Kompier & Phan 2017</bibRefCitation>
|
||
). Muang Huang, the location where Fraser’s specimen was collected, is situated in Khetphiset Xaisomboun, in
|
||
<collectingCountry id="5FD476534361617DFAA2DED94D903F90" box="[1375,1432,1449,1475]" name="Laos" pageId="13" pageNumber="514">Laos</collectingCountry>
|
||
, well north of the southernmost record in
|
||
<collectingCountry id="5FD476534361617DFD9CDEBD4ACC3FB4" box="[609,708,1485,1511]" name="Vietnam" pageId="13" pageNumber="514">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |