241 lines
31 KiB
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241 lines
31 KiB
XML
<document id="75006C68545AD62391690D994C44E369" ID="10.11646/zootaxa.4184.2.1" ID-DOI="10.11646/zootaxa.4184.2.1" ID-GBIF-Dataset="9aad7478-023f-4cf9-9aa7-4e954b607673" ID-ISSN="1175-5326" ID-Zenodo-Dep="164718" ID-ZooBank="5A692B15-5A1F-45C7-AC36-FF155F5A4FB0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478243422585" checkinUser="plazi" docAuthor="Grogan, Willliam L., Díaz, Florentina, Spinelli, Gustavo R. & Ronderos, Maria M." docDate="2016" docId="6E4ABE2FFFA1FF9E30C372DD27BDA355" docLanguage="en" docName="zootaxa.4184.2.1.pdf" docOrigin="Zootaxa 4184 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea spatula Grogan, Díaz, Spinelli & Ronderos, 2016, new species" docType="treatment" docVersion="9" lastPageNumber="219" masterDocId="9273C657FFAEFF8C3054776D2752A640" masterDocTitle="The biting and predaceous midges of Guadeloupe (Diptera: Ceratopogonidae). II. Species of the subfamily Dasyheleinae" masterLastPageNumber="254" masterPageNumber="201" pageNumber="216" updateTime="1698391953395" updateUser="plazi">
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<mods:title id="0735E7F0D2B9AA6F74ACA9F6B89A0BAA">The biting and predaceous midges of Guadeloupe (Diptera: Ceratopogonidae). II. Species of the subfamily Dasyheleinae</mods:title>
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<mods:name id="1A19F5E84CD35CA0E7F16679A9306D57" type="personal">
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<mods:roleTerm id="605894EC38A94E00BC2662B1E5A940A1">Author</mods:roleTerm>
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<mods:namePart id="E3EF0E38FB4A1B278FB62673BB01A835">Grogan, Willliam L.</mods:namePart>
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<mods:namePart id="1FE44356DD3E3DF697C55C1CE3F13817">Díaz, Florentina</mods:namePart>
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<mods:name id="BCB7A0A13521457CF4648E3C615DF86D" type="personal">
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<mods:roleTerm id="E5C686C8CE78E06635D0215176E91193">Author</mods:roleTerm>
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<mods:namePart id="1EC44E468759D0BC4F27156FA904BCFC">Spinelli, Gustavo R.</mods:namePart>
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<mods:name id="B3E69AB3F6CFC72DB1850BE47B467786" type="personal">
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<mods:roleTerm id="28FC907B745A47F9843041AF53C87294">Author</mods:roleTerm>
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<mods:namePart id="0588AD4AA24EE1CB95231E2BA1F99253">Ronderos, Maria M.</mods:namePart>
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<mods:typeOfResource id="724956F67635BEB96A2598E7222611EC">text</mods:typeOfResource>
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<mods:title id="C92BF719BF0EA16B66564599319E94F3">Zootaxa</mods:title>
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<mods:date id="D978604BE552C3A02B93E68C40BD1FA1">2016</mods:date>
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<mods:number id="2264BD1292121C5C1206F1E6EBBBC3BE">4184</mods:number>
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<mods:classification id="7DFF9FFB5DFF764C7AAD3539BAECECED">journal article</mods:classification>
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<mods:identifier id="E315922A152EC7B38ACDFF738EBBE4B1" type="DOI">10.11646/zootaxa.4184.2.1</mods:identifier>
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<treatment id="6E4ABE2FFFA1FF9E30C372DD27BDA355" ID-DOI="http://doi.org/10.5281/zenodo.5457178" ID-GBIF-Taxon="124562864" ID-Zenodo-Dep="5457178" LSID="urn:lsid:plazi:treatment:6E4ABE2FFFA1FF9E30C372DD27BDA355" httpUri="http://treatment.plazi.org/id/6E4ABE2FFFA1FF9E30C372DD27BDA355" lastPageId="18" lastPageNumber="219" pageId="15" pageNumber="216">
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<subSubSection id="AEF95CB2FFA1FF8330C372DD2643A3AB" pageId="15" pageNumber="216" type="nomenclature">
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<paragraph id="E65C0F39FFA1FF8330C372DD2559A38A" blockId="15.[151,523,1456,1516]" box="[151,523,1456,1482]" pageId="15" pageNumber="216">
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<heading id="BD14B855FFA1FF8330C372DD2559A38A" bold="true" box="[151,523,1456,1482]" fontSize="11" level="1" pageId="15" pageNumber="216" reason="1">
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<emphasis id="D497D32BFFA1FF8330C372DD2559A38A" bold="true" box="[151,523,1456,1482]" pageId="15" pageNumber="216">
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<taxonomicName id="21E374BAFFA1FF8330C372DD2626A38A" ID-CoL="8BQX2" box="[151,372,1456,1482]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
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<emphasis id="D497D32BFFA1FF8330C372DD2626A38A" bold="true" box="[151,372,1456,1482]" italics="true" pageId="15" pageNumber="216">Dasyhelea spatula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CFA46E50FFA1FF83312F72DD2559A38A" box="[379,523,1456,1482]" pageId="15" pageNumber="216" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E65C0F39FFA1FF8330C372BE2643A3AB" blockId="15.[151,523,1456,1516]" box="[151,273,1491,1516]" pageId="15" pageNumber="216">
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(
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<figureCitation id="7ED813BCFFA1FF8330CB72BE2658A3AB" box="[159,266,1491,1516]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="16.[151,250,1985,2007]" captionStartId-1="17.[151,250,1794,1816]" captionTargetBox-0="[156,1429,363,1956]" captionTargetBox-1="[169,1421,206,1759]" captionTargetId-0="figure@16.[151,1435,355,1966]" captionTargetId-1="figure@17.[156,1431,193,1773]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." captionText-1="FIGURE 8. Dasyhelea spatula n. sp. Holotype male. A. genitalia. B. sternite 9. C. gonocoxal apodemes and paramere. D. aedeagus. Scale bar: 0.05 mm." httpUri-0="https://zenodo.org/record/164725/files/figure.png" httpUri-1="https://zenodo.org/record/164726/files/figure.png" pageId="15" pageNumber="216">Figs. 7–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="AEF95CB2FFA1FF8330C371762313A084" pageId="15" pageNumber="216" type="diagnosis">
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<paragraph id="E65C0F39FFA1FF8330C371762313A084" blockId="15.[151,1437,1563,2028]" pageId="15" pageNumber="216">
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<emphasis id="D497D32BFFA1FF8330C371762658A074" bold="true" box="[151,266,1563,1588]" pageId="15" pageNumber="216">Diagnosis</emphasis>
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. The only Neotropical species in the
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<taxonomicName id="21E374BAFFA1FF8332E7717625A4A074" box="[691,758,1563,1588]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="grisea">grisea</taxonomicName>
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group with male tergite 9 having an elongate, quadrangular distal portion that extends well beyond apices of gonocoxites and short, triangular, slightly divergent apicolateral processes; and distal portion of the paramere flat, triangular, greatly expanded, spatula-shaped with recurved tip. Females with slender, elongate anterior portion of subgenital plate, and posterior portion with U-shaped excavation; and an ovoid spermatheca with a stout, elongate, greatly recurved neck.
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</paragraph>
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</subSubSection>
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<subSubSection id="AEF95CB2FFA1FF9E309371A22300A525" lastPageId="18" lastPageNumber="219" pageId="15" pageNumber="216" type="description">
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<paragraph id="E65C0F39FFA1FF83309371A222CFA1AC" blockId="15.[151,1437,1563,2028]" pageId="15" pageNumber="216">
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<emphasis id="D497D32BFFA1FF83309371A22656A0A8" bold="true" box="[199,260,1743,1768]" pageId="15" pageNumber="216">Male</emphasis>
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. Head (
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<figureCitation id="7ED813BCFFA1FF83313071A226F9A0A7" box="[356,427,1743,1768]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="15" pageNumber="216">Fig. 7</figureCitation>
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A). Brown. Eyes contiguous for a distance equal to the width of 4 ommatidia. Frontal sclerite broad, slightly bilobed. Antennal flagellum dark brown; flagellomeres 2–9 rhomboidal, 10–13 elongate, 11 or 13 longest; flagellomeres 2–10 with whorl of long sensilla chaetica that form the dense plume that extends to base of flagellomere 13; antennal ratio 0.93. Clypeus (
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<figureCitation id="7ED813BCFFA1FF8332AE7056246FA113" box="[762,829,1851,1876]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="15" pageNumber="216">Fig. 7</figureCitation>
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B) with 6 pairs of setae. Palpus (
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<figureCitation id="7ED813BCFFA1FF8334F8705623BDA113" box="[1196,1263,1851,1876]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="15" pageNumber="216">Fig. 7</figureCitation>
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C) light brown; segment 3 moderately short, slightly bulbous with scattered capitate sensilla; palpal ratio 2.27. Thorax. Scutum dark brown; scutellum with 8 large, 3 smaller setae. Wing (
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<figureCitation id="7ED813BCFFA1FF83330570EE24C5A1DB" box="[849,919,1923,1948]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="15" pageNumber="216">Fig. 7</figureCitation>
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D) slender; membrane hyaline, covered with dense macrotrichia; 2nd radial cell narrow; apex of costa, radius at slight oblique angle; fork of CuA1 at level just anterior to base of 2nd radial cell; anal lobe well developed; wing length
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<quantity id="211BA2DCFFA1FF8333ED70B9234DA1AB" box="[953,1055,2004,2028]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" pageId="15" pageNumber="216" unit="mm" value="1.04">1.04 mm</quantity>
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, width
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<quantity id="211BA2DCFFA1FF83342570BE2385A1AB" box="[1137,1239,2003,2028]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="15" pageNumber="216" unit="mm" value="0.36">0.36 mm</quantity>
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; costal ratio 0.52.
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</paragraph>
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<paragraph id="E65C0F39FFBEFF9E30C377FA26A1A725" blockId="16.[151,1437,151,320]" lastBlockId="18.[151,1437,151,1301]" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217">
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Abdomen. Dark brown. Genitalia (
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<figureCitation id="7ED813BCFFBEFF9C324977FA2538A6EF" box="[541,618,151,176]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="16" pageNumber="217">Figs. 7</figureCitation>
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E, 8A). Tergite 9 tapering progressively distally to apices of gonocoxites, with an elongate quadrangular distal portion that extends considerably beyond gonocoxites, with short, triangular, slightly divergent apicolateral processes with 1 small apical seta; cercus small, located anterior to apicolateral process with several short setae. Sternite 9 (
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<figureCitation id="7ED813BCFFBEFF9C32C2766925B4A75C" box="[662,742,260,285]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="16" pageNumber="217">Figs. 7</figureCitation>
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E, 8B) 1/3 length of greatest width; anterior margin curved, convex; posteromedian portion slightly elongated with short cap-like process that extends onto posterior margin of basal arch of aedeagus. Gonocoxite moderately short, 1.50 x longer than greatest width; gonostylus slightly longer than gonocoxite, pilose, nearly straight, proximal 1/4 with short mesal lobe near terminus of tapered section, apex bluntly rounded. Gonocoxal apodemes and paramere (
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<figureCitation id="7ED813BCFFBFFF9D32AB70C6241CA183" box="[767,846,1963,1988]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="17" pageNumber="218">Figs. 7</figureCitation>
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E, 8C) forming an asymmetrical structure; proximal portion of gonocoxal apodemes broad, with recurved extreme base; right apodeme with tapered, slightly sinuate distal portion, the slender pointed apex of which narrowly contacts paramere; left apodeme with moderately slender distal portion, with truncate apex that does not contact paramere; paramere slender basally, broader mesally, distal portion recurved ventrolaterally, apex broad, spatulate with recurved tip. Aedeagus (
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<figureCitation id="7ED813BCFFBCFF9E355577B22202A6B7" box="[1281,1360,223,248]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="18" pageNumber="219">Figs. 7</figureCitation>
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E, 8D) heavily sclerotized, 0.60 length of greatest width; basal arch very low, extending only 0.10 of total length; basal arm broad, apex narrowed, recurved nearly 180º; posterolateral arms stout, slightly convergent with blunt apices, each with serrate inner margin.
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</paragraph>
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<caption id="B29C5FB1FFBEFF9C30C370AC2590A1B5" httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="16" pageNumber="217" targetBox="[156,1429,363,1956]" targetPageId="16">
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<paragraph id="E65C0F39FFBEFF9C30C370AC2590A1B5" blockId="16.[151,1436,1985,2037]" pageId="16" pageNumber="217">
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<emphasis id="D497D32BFFBEFF9C30C370AC2640A196" bold="true" box="[151,274,1985,2007]" pageId="16" pageNumber="217">FIGURE 7.</emphasis>
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<taxonomicName id="21E374BAFFBEFF9C314D70AC2687A197" box="[281,469,1985,2007]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
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<emphasis id="D497D32BFFBEFF9C314D70AC2687A197" box="[281,469,1985,2007]" italics="true" pageId="16" pageNumber="217">Dasyhelea spatula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CFA46E50FFBEFF9C318E70AC2541A196" box="[474,531,1985,2006]" pageId="16" pageNumber="217" rank="species">
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<emphasis id="D497D32BFFBEFF9C318E70AC2541A196" bold="true" box="[474,531,1985,2006]" pageId="16" pageNumber="217">n. sp.</emphasis>
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</taxonomicNameLabel>
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Holotype male, A–E, Allotype female, F–I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca.
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</paragraph>
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</caption>
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<caption id="B29C5FB1FFBFFF9D30C3706F269EA177" httpUri="https://zenodo.org/record/164726/files/figure.png" pageId="17" pageNumber="218" targetBox="[169,1421,206,1759]" targetPageId="17">
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<paragraph id="E65C0F39FFBFFF9D30C3706F269EA177" blockId="17.[151,1436,1794,1847]" pageId="17" pageNumber="218">
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<emphasis id="D497D32BFFBFFF9D30C3706F2647A157" bold="true" box="[151,277,1794,1816]" pageId="17" pageNumber="218">FIGURE 8.</emphasis>
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<taxonomicName id="21E374BAFFBFFF9D314A706F268FA158" box="[286,477,1794,1816]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="spatula" status="sp. nov.">
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<emphasis id="D497D32BFFBFFF9D314A706F268FA158" box="[286,477,1794,1816]" italics="true" pageId="17" pageNumber="218">Dasyhelea spatula</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="CFA46E50FFBFFF9D31B1706F2573A157" box="[485,545,1794,1815]" pageId="17" pageNumber="218" rank="species">
|
||
<emphasis id="D497D32BFFBFFF9D31B1706F2573A157" bold="true" box="[485,545,1794,1815]" pageId="17" pageNumber="218">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Holotype male. A. genitalia. B. sternite 9. C. gonocoxal apodemes and paramere. D. aedeagus. Scale bar: 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E65C0F39FFBCFF9E309376022300A525" blockId="18.[151,1437,151,1301]" pageId="18" pageNumber="219">
|
||
<emphasis id="D497D32BFFBCFF9E30937602264CA7C8" bold="true" box="[199,286,367,392]" pageId="18" pageNumber="219">Female</emphasis>
|
||
. Head (
|
||
<figureCitation id="7ED813BCFFBCFF9E312F76022693A7C8" box="[379,449,367,392]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="18" pageNumber="219">Fig. 7</figureCitation>
|
||
F). Brown. Eyes contiguous for a distance equal to the width of 4 ommatidia. Frontal sclerite broad, slightly bilobed, with long, slender ventral projection. Antennal flagellum dark brown; flagellomeres 2–8 vasiform, 9–13 becoming gradually more elongate, 13 longest; antennal ratio 0.87. Clypeus with 5 pairs of setae. Palpus (
|
||
<figureCitation id="7ED813BCFFBCFF9E31FC76B126B9A7B4" box="[424,491,476,501]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="18" pageNumber="219">Fig. 7</figureCitation>
|
||
G) light brown, short; segment 3 moderately short with 2 capitate sensillae on mesal surface; palpal ratio 2.09. Thorax. Scutum brown, humeral areas and prescutellar depression paler; scutellum slightly paler with 8 large, 2 smaller setae. Legs pale brown; hind femur with mesal dark spot; femorotibial joints dark; tarsomeres 5 infuscated; hind tibial comb with 6 spines; prothoracic tarsal ratio 2.36, mesothoracic tarsal ratio 2.07, metathoracic tarsal ratio 2.40. Wing (
|
||
<figureCitation id="7ED813BCFFBCFF9E32DA75012586A4C4" box="[654,724,620,645]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="18" pageNumber="219">Fig. 7</figureCitation>
|
||
H) moderately broad; membrane hyaline, covered with dense macrotrichia; 2nd radial reduced to a short suture; apex of costa, radius at oblique angle; fork of CuA1 at level anterior to base of 2nd radial cell; anal lobe well developed, broad; wing length
|
||
<quantity id="211BA2DCFFBCFF9E347275D023DCA494" box="[1062,1166,701,725]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="18" pageNumber="219" unit="mm" value="0.98">0.98 mm</quantity>
|
||
, width
|
||
<quantity id="211BA2DCFFBCFF9E34B075D0221FA494" box="[1252,1357,701,725]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="219" unit="mm" value="0.44">0.44 mm</quantity>
|
||
; costal ratio 0.51. Halter pale. Abdomen (
|
||
<figureCitation id="7ED813BCFFBCFF9E3241758D2501A4B9" box="[533,595,736,761]" captionStart="FIGURE 7" captionStartId="16.[151,250,1985,2007]" captionTargetBox="[156,1429,363,1956]" captionTargetId="figure@16.[151,1435,355,1966]" captionTargetPageId="16" captionText="FIGURE 7. Dasyhelea spatula n. sp. Holotype male, A – E, Allotype female, F – I. A, F. head. B, G. clypeus. C, G. palpus. D, H. wing. E. genitalia. I. subgenital plate and spermatheca." httpUri="https://zenodo.org/record/164725/files/figure.png" pageId="18" pageNumber="219">
|
||
Fig.
|
||
<date id="925D29F9FFBCFF9E3210758D2501A4B9" box="[580,595,736,761]" pageId="18" pageNumber="219">7</date>
|
||
</figureCitation>
|
||
I). Brown. Subgenital plate moderately sclerotized; anterior portion slender, elongate; posterior portion stout, with U-shaped median excavation; posterolateral arms more heavily sclerotized, bases connected by slender sclerite. Spermatheca ovoid, length/width
|
||
<date id="925D29F9FFBCFF9E33F37445248AA500" box="[935,984,808,833]" pageId="18" pageNumber="219" value="1968-10">68 x</date>
|
||
60 µm, with stout greatly elongate neck that is recurved nearly 180º and narrowly separated from spermatheca, length 50 µm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEF95CB2FFBCFF9E3093741D26A4A5C9" box="[199,502,880,905]" pageId="18" pageNumber="219" type="distribution">
|
||
<paragraph id="E65C0F39FFBCFF9E3093741D26A4A5C9" blockId="18.[151,1437,151,1301]" box="[199,502,880,905]" pageId="18" pageNumber="219">
|
||
<emphasis id="D497D32BFFBCFF9E3093741D260BA5C9" bold="true" box="[199,345,880,905]" pageId="18" pageNumber="219">Distribution</emphasis>
|
||
.
|
||
<collectingRegion id="2427C1DBFFBCFF9E3133741D26A0A5C9" box="[359,498,880,905]" country="France" name="Guadeloupe" pageId="18" pageNumber="219">
|
||
<collectingCountry id="9EF44FA9FFBCFF9E3133741D26A0A5C9" box="[359,498,880,905]" name="Guadeloupe" pageId="18" pageNumber="219">Guadeloupe</collectingCountry>
|
||
</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEF95CB2FFBCFF9E309374F926B6A258" pageId="18" pageNumber="219" type="materials_examined">
|
||
<paragraph id="E65C0F39FFBCFF9E309374F92479A591" blockId="18.[151,1437,151,1301]" pageId="18" pageNumber="219">
|
||
<emphasis id="D497D32BFFBCFF9E309374F9263DA5ED" bold="true" box="[199,367,916,941]" pageId="18" pageNumber="219">Type material</emphasis>
|
||
. Holotype male,
|
||
<materialsCitation id="568B0564FFBCFF9E326374F92475A591" ID-GBIF-Occurrence="1324639453" collectingDate="2012-05-22" collectionCode="FSCA" collectorName="R. H. Turnbow" country="Guadeloupe" county="Cretes" location="Basse Terre" municipality="Trace" pageId="18" pageNumber="219" specimenCount="1" typeStatus="allotype">
|
||
<typeStatus id="3958B19BFFBCFF9E326374F925C3A5ED" box="[567,657,916,941]" pageId="18" pageNumber="219" type="allotype">allotype</typeStatus>
|
||
female labeled “
|
||
<collectingCountry id="9EF44FA9FFBCFF9E330074F924B3A5ED" box="[852,993,916,941]" name="Guadeloupe" pageId="18" pageNumber="219">Guadeloupe</collectingCountry>
|
||
,
|
||
<location id="E33C59E2FFBCFF9E33B974F82324A5EC" LSID="urn:lsid:plazi:treatment:6E4ABE2FFFA1FF9E30C372DD27BDA355:E33C59E2FFBCFF9E33B974F82324A5EC" box="[1005,1142,917,940]" country="Guadeloupe" county="Cretes" municipality="Trace" name="Basse Terre" pageId="18" pageNumber="219">Basse Terre</location>
|
||
,
|
||
<collectingMunicipality id="06389543FFBCFF9E34D674F82393A5EC" box="[1154,1217,917,940]" pageId="18" pageNumber="219">Trace</collectingMunicipality>
|
||
des
|
||
<collectingCounty id="0F3D77B5FFBCFF9E34AD74F82212A5ED" box="[1273,1344,917,941]" pageId="18" pageNumber="219">Cretes</collectingCounty>
|
||
(D-14),
|
||
<collectingDate id="8219D011FFBCFF9E30C374D42640A591" box="[151,274,953,977]" pageId="18" pageNumber="219" value="2012-05-22">
|
||
<date id="925D29F9FFBCFF9E30C374D42640A591" box="[151,274,953,977]" pageId="18" pageNumber="219" value="2012-05-22">22-V-2012</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="4B166AEFFFBCFF9E314974D42695A591" box="[285,455,952,977]" pageId="18" pageNumber="219">R. H. Turnbow</collectorName>
|
||
,
|
||
<collectingMethod id="3FA2772EFFBCFF9E318674D42579A591" box="[466,555,953,977]" pageId="18" pageNumber="219">BL trap</collectingMethod>
|
||
” (deposited in
|
||
<collectionCode id="80F297FCFFBCFF9E328074D4244DA591" box="[724,799,953,977]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="18" pageNumber="219">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E65C0F39FFBCFF9E309374B126B6A258" blockId="18.[151,1437,151,1301]" pageId="18" pageNumber="219">
|
||
<emphasis id="D497D32BFFBCFF9E309374B12579A5B5" bold="true" box="[199,555,988,1013]" pageId="18" pageNumber="219">Derivation of specific epithet</emphasis>
|
||
. The species epithet is a reference to the broad, triangular, spatulate distal portion of the male paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AEF95CB2FFBCFF9E3093734927BDA355" pageId="18" pageNumber="219" type="discussion">
|
||
<paragraph id="E65C0F39FFBCFF9E3093734927BDA355" blockId="18.[151,1437,151,1301]" pageId="18" pageNumber="219">
|
||
<emphasis id="D497D32BFFBCFF9E309373492617A27D" bold="true" box="[199,325,1060,1085]" pageId="18" pageNumber="219">Discussion</emphasis>
|
||
. The aedeagus of the
|
||
<typeStatus id="3958B19BFFBCFF9E3211734925F5A27D" box="[581,679,1060,1085]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="21E374BAFFBCFF9E3344734824D5A27C" box="[784,903,1060,1084]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="spatula">
|
||
<emphasis id="D497D32BFFBCFF9E334473482476A27C" box="[784,804,1061,1084]" italics="true" pageId="18" pageNumber="219">D</emphasis>
|
||
.
|
||
<emphasis id="D497D32BFFBCFF9E3360734924D5A27C" box="[820,903,1060,1084]" italics="true" pageId="18" pageNumber="219">spatula</emphasis>
|
||
</taxonomicName>
|
||
has serrate inner margins of the posterolateral arms that are similar to those of
|
||
<taxonomicName id="21E374BAFFBCFF9E32557324252AA220" box="[513,632,1096,1120]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="hamula" status="sp. nov.">
|
||
<emphasis id="D497D32BFFBCFF9E325573242547A220" box="[513,533,1097,1120]" italics="true" pageId="18" pageNumber="219">D</emphasis>
|
||
.
|
||
<emphasis id="D497D32BFFBCFF9E32707325252AA220" box="[548,632,1096,1120]" italics="true" pageId="18" pageNumber="219">hamula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CFA46E50FFBCFF9E322B732525ECA221" box="[639,702,1096,1121]" pageId="18" pageNumber="219" rank="species">
|
||
<emphasis id="D497D32BFFBCFF9E322B732525ECA221" bold="true" box="[639,702,1096,1121]" pageId="18" pageNumber="219">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
However, the
|
||
<typeStatus id="3958B19BFFBCFF9E333673252496A221" box="[866,964,1096,1121]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
|
||
male of that species differs from this new species in having a slender distal portion of the paramere with a subapical sharply pointed tubercle and a short, recurved apex. The subgenital plate and spermatheca of the
|
||
<typeStatus id="3958B19BFFBCFF9E336573FD24D9A2E9" box="[817,907,1168,1193]" pageId="18" pageNumber="219" type="allotype">allotype</typeStatus>
|
||
female of
|
||
<taxonomicName id="21E374BAFFBCFF9E345673FC2324A2E8" box="[1026,1142,1168,1192]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="spatula">
|
||
<emphasis id="D497D32BFFBCFF9E345673FC2344A2E8" box="[1026,1046,1169,1192]" italics="true" pageId="18" pageNumber="219">D</emphasis>
|
||
.
|
||
<emphasis id="D497D32BFFBCFF9E347773FD2324A2E8" box="[1059,1142,1168,1192]" italics="true" pageId="18" pageNumber="219">spatula</emphasis>
|
||
</taxonomicName>
|
||
is similar to females of
|
||
<taxonomicName id="21E374BAFFBCFF9E35D573FC260AA28C" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="pseudoincisurata">
|
||
<emphasis id="D497D32BFFBCFF9E35D573FC22C7A2E8" box="[1409,1429,1169,1192]" italics="true" pageId="18" pageNumber="219">D</emphasis>
|
||
.
|
||
<emphasis id="D497D32BFFBCFF9E30C373D9260AA28C" box="[151,344,1204,1228]" italics="true" pageId="18" pageNumber="219">pseudoincisurata</emphasis>
|
||
</taxonomicName>
|
||
, but the latter species is larger (wing length
|
||
<quantity id="211BA2DCFFBCFF9E330373D82352A28D" box="[855,1024,1205,1229]" metricMagnitude="-3" metricUnit="m" metricValue="1.335" metricValueMax="1.47" metricValueMin="1.2" pageId="18" pageNumber="219" unit="mm" value="1.335" valueMax="1.47" valueMin="1.2">1.20–1.47 mm</quantity>
|
||
), the distal portion of the subgenital plate is extremely slender and the neck of its spermatheca is tapered and not nearly as recurved as in this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |