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<document id="5A205716940CF9854AD8DBA2DAF9125A" ID-DOI="10.59893/bjc.23(2).005" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203378" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657783974" checkinUser="felipe" docAuthor="Minkina, Łukasz" docDate="2023" docId="65588792FF98FFECCE5EE838FE8F717D" docLanguage="en" docName="BalticJColeopterol.23.2.209-214.pdf" docOrigin="Baltic Journal of Coleopterology (Athens, Greece) 23 (2)" docSource="http://dx.doi.org/10.1073/pnas.2404108121" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Rhyparus anichtchenkoi Minkina, 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="213" masterDocId="9961FFEAFF99FFE8CE2CEA06FF89755A" masterDocTitle="A new species of the genusRhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) from Papua New Guinea" masterLastPageNumber="214" masterPageNumber="209" pageNumber="210" updateTime="1722697355684" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="DD7E0B3A00D35C1A9E6B4A3A461B2702" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3089D66454BCC26A03024E9EE507DB66">
<mods:title id="ECFC05C031A403F40FB03008E025325E">A new species of the genusRhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) from Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="81DD1856806243B4222734B97CEB1FEA" type="personal">
<mods:role id="417B848DA560F690268CEC8E1E3FAAC4">
<mods:roleTerm id="4086C9FDDB383FCE05972CE91F575858">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41A8BD733BF15C8E175E6080B9EDF1B0">Minkina, Łukasz</mods:namePart>
</mods:name>
<mods:typeOfResource id="4118105BB3C53C85958CD4DABEBD2937">text</mods:typeOfResource>
<mods:relatedItem id="54134FFAD28F336BEBF75E8A059A0A6A" type="host">
<mods:titleInfo id="7835FC11333EE3E4F2BC0461208B65CB">
<mods:title id="9F18DFF1FAF521C8915A613E0BCA9CBB">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="BAB98C08C3C18F22068313244A8D4A0F">
<mods:date id="CF89A96D20F0243E1481DDBD1EF24572">2023</mods:date>
<mods:detail id="1925E406CD86A8A29E841D7E517C6905" type="series">
<mods:title id="1F1D53367F5E5FCCD721C80A4AF351B9">Athens, Greece</mods:title>
</mods:detail>
<mods:detail id="2F4D9C3F4BE4D20AFCD67D8B19A30C59" type="volume">
<mods:number id="13A184B88586A6D4C5771E1A59344246">23</mods:number>
</mods:detail>
<mods:detail id="01524ED5FB40FB6C05AEED74DAB10255" type="issue">
<mods:number id="AFF104548DDE7DB5D6188E935867B358">2</mods:number>
</mods:detail>
<mods:extent id="3ED5C84D20E4D887739307E677B21731" unit="page">
<mods:start id="0FB16339072CABEC5CC79040D8EF8691">209</mods:start>
<mods:end id="05777FF92DB16111BB111BC0FC1EB6AD">214</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A6CA7BA17111A6D2F9467F3A4EE96B98">
<mods:url id="7B983D54BCBF492886F887F7CF1E7305">http://dx.doi.org/10.1073/pnas.2404108121</mods:url>
</mods:location>
<mods:classification id="9DB04688CCF662EF2B3A94C44ADB249D">journal article</mods:classification>
<mods:identifier id="DC5D0F56AED89EC1F61114AD6DE76E39" type="DOI">10.59893/bjc.23(2).005</mods:identifier>
<mods:identifier id="0366CDDC645B58D9D402CEDC7A0523A3" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="CDF7FAB663DEF93E0369FE78BE3DE2A8" type="Zenodo-Dep">13203378</mods:identifier>
</mods:mods>
<treatment id="65588792FF98FFECCE5EE838FE8F717D" LSID="urn:lsid:plazi:treatment:65588792FF98FFECCE5EE838FE8F717D" httpUri="http://treatment.plazi.org/id/65588792FF98FFECCE5EE838FE8F717D" lastPageId="4" lastPageNumber="213" pageId="1" pageNumber="210">
<subSubSection id="A5EB650FFF98FFE9CE5EE838FE52770F" box="[114,475,574,598]" pageId="1" pageNumber="210" type="nomenclature">
<paragraph id="ED4E3684FF98FFE9CE5EE838FE52770F" blockId="1.[114,475,512,598]" box="[114,475,574,598]" pageId="1" pageNumber="210">
<heading id="B60681E8FF98FFE9CE5EE838FE52770F" box="[114,475,574,598]" fontSize="10" level="2" pageId="1" pageNumber="210" reason="7">
<taxonomicName id="2AF14D07FF98FFE9CE5EE838FE09770C" ID-CoL="C399V" authorityName="Minkina" authorityYear="2023" box="[114,384,574,598]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">
<emphasis id="DF85EA96FF98FFE9CE5EE838FE09770C" bold="true" box="[114,384,574,598]" italics="true" pageId="1" pageNumber="210">Rhyparus anichtchenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4B657EDFF98FFE9CFABE838FE52770F" box="[391,475,574,597]" pageId="1" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A5EB650FFF98FFE9CE5EE87DFF5277C9" box="[114,219,635,659]" pageId="1" pageNumber="210" type="description">
<paragraph id="ED4E3684FF98FFE9CE5EE87DFF5277C9" blockId="1.[114,593,635,750]" box="[114,219,635,659]" pageId="1" pageNumber="210">
(
<figureCitation id="75CA2A01FF98FFE9CE55E87DFF5B77C9" box="[121,210,635,659]" captionStart-0="Figs 1-3" captionStart-1="Figs 4-6" captionStartId-0="1.[114,160,1439,1463]" captionStartId-1="1.[624,670,810,834]" captionTargetBox-0="[120,593,788,1398]" captionTargetBox-1="[632,1098,187,786]" captionTargetId-0="figure-242@1.[114,598,781,1402]" captionTargetId-1="figure-244@1.[624,1106,171,804]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="Figs 1-3.Rhyparus anichtchenkoi sp. nov., ♂, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Scale lines: 1.0 mm." captionText-1="Figs 4-6.Rhyparus anichtchenkoisp. nov.: 4- head, ♂, holotype; 5- last abdominal ventrite and pygidium, ♀, paratype; 6- aedeagus, holotype. Scale lines: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13203382" figureDoi-1="http://doi.org/10.5281/zenodo.13203380" httpUri-0="https://zenodo.org/record/13203382/files/figure.png" httpUri-1="https://zenodo.org/record/13203380/files/figure.png" pageId="1" pageNumber="210">Figs 1-6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A5EB650FFF98FFE9CE5EE8BEFF1373E4" pageId="1" pageNumber="210" type="materials_examined">
<paragraph id="ED4E3684FF98FFE9CE5EE8BEFEA677B4" blockId="1.[114,593,635,750]" pageId="1" pageNumber="210">
<materialsCitation id="5D993CD9FF98FFE9CE5EE8BEFEA277B4" country="Papua New Guinea" location="Batteta" pageId="1" pageNumber="210" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="DF85EA96FF98FFE9CE5EE8BEFE87778A" bold="true" box="[114,270,696,720]" pageId="1" pageNumber="210">
<typeStatus id="324A8826FF98FFE9CE5EE8BEFF25778A" box="[114,172,696,720]" pageId="1" pageNumber="210">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="95E67614FF98FFE9CF39E8BEFE61778A" box="[277,488,696,720]" name="Papua New Guinea" pageId="1" pageNumber="210">Papua New Guinea</collectingCountry>
,
<collectingRegion id="2F35F866FF98FFE9CFDFE8BEFF5B77B4" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="210">Madang province</collectingRegion>
,
<location id="E82E605FFF98FFE9CEF0E8D1FEA277B4" LSID="urn:lsid:plazi:treatment:65588792FF98FFECCE5EE838FE8F717D:E82E605FFF98FFE9CEF0E8D1FEA277B4" box="[220,299,727,750]" country="Papua New Guinea" name="Batteta" pageId="1" pageNumber="210" stateProvince="Madang">Batteta</location>
</materialsCitation>
.
</paragraph>
<caption id="B98E660CFF98FFE9CC5CE92CFBAD76C4" ID-DOI="http://doi.org/10.5281/zenodo.13203380" ID-Zenodo-Dep="13203380" httpUri="https://zenodo.org/record/13203380/files/figure.png" pageId="1" pageNumber="210" startId="1.[624,670,810,834]" targetBox="[632,1098,187,786]" targetPageId="1" targetType="figure">
<paragraph id="ED4E3684FF98FFE9CC5CE92CFBAD76C4" blockId="1.[624,1103,810,926]" pageId="1" pageNumber="210">
Figs 4-6.
<taxonomicName id="2AF14D07FF98FFE9CCF7E92DFC6E7619" authorityName="Minkina" authorityYear="2023" box="[731,999,811,835]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">
<emphasis id="DF85EA96FF98FFE9CCF7E92DFC6E7619" box="[731,999,811,835]" italics="true" pageId="1" pageNumber="210">Rhyparus anichtchenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4B657EDFF98FFE9CDC6E92DFBC27618" box="[1002,1099,811,834]" pageId="1" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
: 4- head, ♂, holotype; 5- last abdominal ventrite and pygidium, ♀, paratype; 6- aedeagus, holotype. Scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="ED4E3684FF98FFE9CC5CE9C5FB8170D3" blockId="1.[624,1103,963,1417]" pageId="1" pageNumber="210">
<emphasis id="DF85EA96FF98FFE9CC5CE9C5FCA97681" bold="true" box="[624,800,963,987]" pageId="1" pageNumber="210">Type material.</emphasis>
<materialsCitation id="5D993CD9FF98FFE9CD03E9C5FB8170D3" collectingDate="1993-03-23" collectingDateMax="1996-07-02" collectingDateMin="1993-03-23" collectionCode="IRSN, LMCN, DUBC" collectorName="Coll. R. Isc. N. B. &amp; Coll. R. Isc. &amp; Light T &amp; Olivier Missa" location="Batteta" municipality="Canopy" pageId="1" pageNumber="210" specimenCount="5" specimenCount-male="1" stateProvince="Madang" typeStatus="holotype">
<emphasis id="DF85EA96FF98FFE9CD03E9C5FC1E7681" bold="true" box="[815,919,963,987]" pageId="1" pageNumber="210">
<typeStatus id="324A8826FF98FFE9CD03E9C5FC1E7681" box="[815,919,963,987]" pageId="1" pageNumber="210" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="FBF7FD0DFF98FFE9CD82E9C2FC4B7681" box="[942,962,964,987]" count="1" pageId="1" pageNumber="210" type="male">♂</specimenCount>
): “
<collectorName id="40045352FF98FFE9CDC7E9C2FD4676A0" pageId="1" pageNumber="210">Coll. R. Isc. N.B.</collectorName>
|
<collectingMunicipality id="0D2AACFEFF98FFE9CCCDE9E4FCBD76A0" box="[737,820,994,1018]" pageId="1" pageNumber="210">Canopy</collectingMunicipality>
mission P.N.G. |
<collectingRegion id="2F35F866FF98FFE9CDDDE9E4FD447143" country="Papua New Guinea" name="Madang" pageId="1" pageNumber="210">Madang province</collectingRegion>
|
<location id="E82E605FFF98FFE9CCC7EE04FCBF7143" LSID="urn:lsid:plazi:treatment:65588792FF98FFECCE5EE838FE8F717D:E82E605FFF98FFE9CCC7EE04FCBF7143" box="[747,822,1026,1049]" municipality="Canopy" name="Batteta" pageId="1" pageNumber="210" stateProvince="Madang">Batteta</location>
<date id="994F1044FF98FFE9CD6FEE07FC3C7143" box="[835,949,1025,1049]" pageId="1" pageNumber="210" value="1993-04-21">
<collectingDate id="890BE9ACFF98FFE9CD6FEE07FC3C7143" box="[835,949,1025,1049]" pageId="1" pageNumber="210" value="1993-04-21">21.iv.1993</collectingDate>
</date>
|
<collectorName id="40045352FF98FFE9CDFFEE07FBA37143" box="[979,1066,1025,1049]" pageId="1" pageNumber="210">Light T</collectorName>
2 | leg.
<collectorName id="40045352FF98FFE9CC80EE19FCC7716D" box="[684,846,1055,1079]" pageId="1" pageNumber="210">Olivier Missa</collectorName>
” (
<collectionCode id="8BE0AE41FF98FFE9CD58EE26FC3E7162" box="[884,951,1056,1080]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="210">IRSN</collectionCode>
).
<emphasis id="DF85EA96FF98FFE9CDFAEE19FBC1716D" bold="true" box="[982,1096,1055,1079]" pageId="1" pageNumber="210">
<typeStatus id="324A8826FF98FFE9CDFAEE19FBC1716D" box="[982,1096,1055,1079]" pageId="1" pageNumber="210" type="paratype">Paratypes</typeStatus>
</emphasis>
(8exx.):
<specimenCount id="FBF7FD0DFF98FFE9CCFDEE38FC88710C" box="[721,769,1086,1110]" count="1" pageId="1" pageNumber="210" type="generic">1ex.</specimenCount>
: “the same as
<typeStatus id="324A8826FF98FFE9CD98EE38FB98710C" box="[948,1041,1086,1110]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
but: “
<date id="994F1044FF98FFE9CC56EE5BFD66712F" box="[634,751,1117,1141]" pageId="1" pageNumber="210" value="1993-03-23">
<collectingDate id="890BE9ACFF98FFE9CC56EE5BFD66712F" box="[634,751,1117,1141]" pageId="1" pageNumber="210" value="1993-03-23">23.iii.1993</collectingDate>
</date>
|
<collectorName id="40045352FF98FFE9CD37EE5BFCF0712F" box="[795,889,1117,1141]" pageId="1" pageNumber="210">Light T</collectorName>
2””(
<collectionCode id="8BE0AE41FF98FFE9CD88EE5BFC6F712F" box="[932,998,1117,1141]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="210">IRSN</collectionCode>
); 2exx.: “the same as
<typeStatus id="324A8826FF98FFE9CD3AEE7AFCFD71CE" box="[790,884,1148,1172]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
but: “
<date id="994F1044FF98FFE9CDEAEE7DFBBD71C9" box="[966,1076,1147,1171]" pageId="1" pageNumber="210" value="1996-07-02">
<collectingDate id="890BE9ACFF98FFE9CDEAEE7DFBBD71C9" box="[966,1076,1147,1171]" pageId="1" pageNumber="210" value="1996-07-02">2.vii.1996</collectingDate>
</date>
| light AR60””(
<collectionCode id="8BE0AE41FF98FFE9CD3EEE9CFCDB71E8" box="[786,850,1178,1202]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="210">IRSN</collectionCode>
;
<collectionCode id="8BE0AE41FF98FFE9CD4BEE9CFC3071E8" box="[871,953,1178,1202]" pageId="1" pageNumber="210">ŁMCN</collectionCode>
);
<specimenCount id="FBF7FD0DFF98FFE9CDFAEE9CFB8F71E8" box="[982,1030,1178,1202]" count="1" pageId="1" pageNumber="210" type="generic">1ex.</specimenCount>
: “the same as
<typeStatus id="324A8826FF98FFE9CCF2EEBFFCB2718B" box="[734,827,1209,1233]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
but: “
<date id="994F1044FF98FFE9CDBEEEBFFC78718B" box="[914,1009,1209,1233]" pageId="1" pageNumber="210" value="1996-05-02">
<collectingDate id="890BE9ACFF98FFE9CDBEEEBFFC78718B" box="[914,1009,1209,1233]" pageId="1" pageNumber="210" value="1996-05-02">2.v.1996</collectingDate>
</date>
| light AR10””(
<collectionCode id="8BE0AE41FF98FFE9CCE3EEDEFC9671AA" box="[719,799,1240,1264]" pageId="1" pageNumber="210">DUBC</collectionCode>
); 2exx.: “the same as
<typeStatus id="324A8826FF98FFE9CC5CEEF0FD477054" box="[624,718,1270,1294]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
but: “
<date id="994F1044FF98FFE9CD7CEEF0FC4A7054" box="[848,963,1270,1294]" pageId="1" pageNumber="210" value="1996-04-22">
<collectingDate id="890BE9ACFF98FFE9CD7CEEF0FC4A7054" box="[848,963,1270,1294]" pageId="1" pageNumber="210" value="1996-04-22">22.iv.1996</collectingDate>
</date>
| light M8””(
<collectionCode id="8BE0AE41FF98FFE9CC9FEF13FD7F7077" box="[691,758,1301,1325]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="210">IRSN</collectionCode>
;
<collectionCode id="8BE0AE41FF98FFE9CD2BEF13FCD07077" box="[775,857,1301,1325]" pageId="1" pageNumber="210">ŁMCN</collectionCode>
);
<specimenCount id="FBF7FD0DFF98FFE9CD5DEF13FC2B7077" box="[881,930,1301,1325]" count="1" pageId="1" pageNumber="210" type="generic">1ex.</specimenCount>
: “the same as
<typeStatus id="324A8826FF98FFE9CC5CEF32FD477016" box="[624,718,1332,1356]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
but: “
<date id="994F1044FF98FFE9CD7CEF35FC4A7011" box="[848,963,1331,1355]" pageId="1" pageNumber="210" value="1996-04-15">
<collectingDate id="890BE9ACFF98FFE9CD7CEF35FC4A7011" box="[848,963,1331,1355]" pageId="1" pageNumber="210" value="1996-04-15">15.iv.1996</collectingDate>
</date>
| light AR9””(
<collectionCode id="8BE0AE41FF98FFE9CCEEEF54FC8D7030" box="[706,772,1362,1386]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="210">IRSN</collectionCode>
);
<specimenCount id="FBF7FD0DFF98FFE9CD34EF54FCC07030" box="[792,841,1362,1386]" count="1" pageId="1" pageNumber="210" type="generic">1ex.</specimenCount>
: “the same as
<typeStatus id="324A8826FF98FFE9CDC7EF54FBC67030" box="[1003,1103,1362,1386]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
but: “
<date id="994F1044FF98FFE9CC87EF77FC9870D3" box="[683,785,1393,1417]" pageId="1" pageNumber="210" value="1996-06-06">
<collectingDate id="890BE9ACFF98FFE9CC87EF77FC9870D3" box="[683,785,1393,1417]" pageId="1" pageNumber="210" value="1996-06-06">6.vi.1996</collectingDate>
</date>
| light AR16””(
<collectionCode id="8BE0AE41FF98FFE9CD96EF77FC7570D3" box="[954,1020,1393,1417]" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="210">IRSN</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="B98E660CFF98FFE9CE5EEF99FDBC70AE" ID-DOI="http://doi.org/10.5281/zenodo.13203382" ID-Zenodo-Dep="13203382" httpUri="https://zenodo.org/record/13203382/files/figure.png" pageId="1" pageNumber="210" startId="1.[114,160,1439,1463]" targetBox="[120,593,788,1398]" targetPageId="1" targetType="figure">
<paragraph id="ED4E3684FF98FFE9CE5EEF99FDBC70AE" blockId="1.[114,593,1439,1525]" pageId="1" pageNumber="210">
Figs 1-3.
<taxonomicName id="2AF14D07FF98FFE9CEF1EFA6FE6070E2" authorityName="Minkina" authorityYear="2023" box="[221,489,1440,1464]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">
<emphasis id="DF85EA96FF98FFE9CEF1EFA6FE6070E2" box="[221,489,1440,1464]" italics="true" pageId="1" pageNumber="210">Rhyparus anichtchenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4B657EDFF98FFE9CFC1EFA6FDC570ED" box="[493,588,1440,1463]" pageId="1" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Scale lines: 1.0 mm.
</paragraph>
</caption>
<paragraph id="ED4E3684FF98FFE9CE5EECA0FF1373E4" blockId="1.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="1" pageNumber="210">210</paragraph>
</subSubSection>
<subSubSection id="A5EB650FFF98FFECCC5CEFA8FD937701" lastPageId="4" lastPageNumber="213" pageId="1" pageNumber="210" type="description">
<paragraph id="ED4E3684FF98FFEACC5CEFA8FEFE747F" blockId="1.[624,1103,1454,1632]" lastBlockId="2.[189,668,178,293]" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210">
<emphasis id="DF85EA96FF98FFE9CC5CEFA8FC3F709C" bold="true" box="[624,950,1454,1478]" pageId="1" pageNumber="210">
Description of the
<typeStatus id="324A8826FF98FFE9CD6AEFA8FC39709C" box="[838,944,1454,1478]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
.
</emphasis>
Dorsum (
<figureCitation id="75CA2A01FF98FFE9CA32EFA8FDF470BF" captionStart="Figs 1-3" captionStartId="1.[114,160,1439,1463]" captionTargetBox="[120,593,788,1398]" captionTargetId="figure-242@1.[114,598,781,1402]" captionTargetPageId="1" captionText="Figs 1-3.Rhyparus anichtchenkoi sp. nov., ♂, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203382" httpUri="https://zenodo.org/record/13203382/files/figure.png" pageId="1" pageNumber="210">Fig. 1</figureCitation>
). Length:
<quantity id="2A099B61FF98FFE9CCDEEFCBFCC370BE" box="[754,842,1485,1509]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="210" unit="mm" value="6.2">6.2 mm</quantity>
; maximum width:
<quantity id="2A099B61FF98FFE9CA36EFCBFD147359" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="210" unit="mm" value="1.95">1.95 mm</quantity>
. Body medium-sized for members of this genus, moderately elongate, convex, shiny, apparently almost glabrous, though partly clothed with very small yellowish setae on head and all longitudinal costae on pronotum and elytra. Brownish to dark brown; antennae, tarsomeres and mouth parts pale brown.
</paragraph>
<paragraph id="ED4E3684FF9BFFEACE91EB4DFDB27674" blockId="2.[189,668,331,815]" pageId="2" pageNumber="211">
Head (
<figureCitation id="75CA2A01FF9BFFEACF3EEB4DFEE97439" box="[274,352,331,355]" captionStart="Figs 4-6" captionStartId="1.[624,670,810,834]" captionTargetBox="[632,1098,187,786]" captionTargetId="figure-244@1.[624,1106,171,804]" captionTargetPageId="1" captionText="Figs 4-6.Rhyparus anichtchenkoisp. nov.: 4- head, ♂, holotype; 5- last abdominal ventrite and pygidium, ♀, paratype; 6- aedeagus, holotype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203380" httpUri="https://zenodo.org/record/13203380/files/figure.png" pageId="2" pageNumber="211">Fig. 4</figureCitation>
) shiny, transversely subhexagonal, clypeus trapezoidal in outline, anteriorly very weakly sinuate, on sides weakly upturned as obtuse, weak tooth, and later sinuous on either side; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a not so deep groove; convexity with a pair of quite distinct, very short, convergent ridges, nearly on whole surface with distinct, moderately coarse punctures bearing small setae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by quite regularly spaced, quite dense, moderately large punctures bearing small setae.
</paragraph>
<paragraph id="ED4E3684FF9BFFEACE91E953FDF1702F" blockId="2.[189,669,852,1397]" pageId="2" pageNumber="211">Pronotum shiny, tops of costae distinctly shiny; with eight distinct costae and seven intercostae, with two lateral, rounded lobes on each side. Anterior and posteror lobes with similar high, anterior lobe somewhat narrower than posterior. Costae of middle, third and fourth pair not interrupted in basal part of apical half, very distinctly convergent in the middle; second pair of costae very distinctly and widely interrupted in the middle; costae on each side with very small punctures bearing very small setae. All intercostae in anterior part without additional short costae. Median intercostae with dense punctures, very distinctly concentrated around median part into longitudinal line, all intercostae in basal part with distinct, dense punctation.</paragraph>
<paragraph id="ED4E3684FF9BFFEACE91EF9DFD9170E9" blockId="2.[189,536,1435,1459]" box="[189,536,1435,1459]" pageId="2" pageNumber="211">Scutellum almost imperceptible.</paragraph>
<paragraph id="ED4E3684FF9BFFEACE91EFDEFC7C77EE" blockId="2.[189,668,1496,1613]" lastBlockId="2.[699,1179,178,692]" pageId="2" pageNumber="211">Elytra shiny. Each elytron with six elevated costae, and five flat intercostae. Costae on sides with very small punctures bearing very small setae. Preapical glandular area relatively small. Intercostae first, second and fourth with two distinct rows of punctures; third intercostae with three indistinct rows of punctures; additionally, on third intercostae there is very short additional costae with very short additional rows of punctures on its sides. Fifth intercosta with one row of punctures; in basal part of fourth intercosta there is region with extremally short additional costa, with some additional punctures. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and mediointernal caudal bulbs not divided; mediointernal caudal bulb shortened, transversally rounded, internally stretched.</paragraph>
<paragraph id="ED4E3684FF9BFFEACC97E8DCFC1F7617" blockId="2.[699,1178,730,845]" pageId="2" pageNumber="211">Pygidium with dense, irregularly spaced punctation, with weak longitudinal rib in the middle and deep excisionon its sides. Normaly developed.</paragraph>
<paragraph id="ED4E3684FF9BFFEACC97E975FC667186" blockId="2.[699,1179,883,1244]" pageId="2" pageNumber="211">
Venter (
<figureCitation id="75CA2A01FF9BFFEACD37E975FCEB76D1" box="[795,866,883,907]" captionStart="Figs 1-3" captionStartId="1.[114,160,1439,1463]" captionTargetBox="[120,593,788,1398]" captionTargetId="figure-242@1.[114,598,781,1402]" captionTargetPageId="1" captionText="Figs 1-3.Rhyparus anichtchenkoi sp. nov., ♂, holotype: 1- dorsal view; 2- ventral view; 3- lateral view. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203382" httpUri="https://zenodo.org/record/13203382/files/figure.png" pageId="2" pageNumber="211">Fig. 2</figureCitation>
) moderately shiny. Mesometaventral plate flattened in the middle, with distinct, narrow, quite deep longitudinal furrow; punctation of mesometaventral plate dense, quite regularly spaced, irregular in size; all punctures bearing short setae. Abdominal ventrites moderately shiny, very densely and coarsely punctate. Last abdominal ventrite normaly developed, with dense punctures, which are very coarse nearby its base; in the middle with longitudinal rib.
</paragraph>
<paragraph id="ED4E3684FF9BFFEACC97EF04FCB170CE" blockId="2.[699,1178,1281,1428]" pageId="2" pageNumber="211">Meso- and metafemora without tubercles on lower border; all femora shiny, with regular, very distinct, rather fine, very dense punctation; all punctures bearing small setae.</paragraph>
<paragraph id="ED4E3684FF9BFFEACC97EFBFFBBB7355" blockId="2.[699,1178,1465,1551]" pageId="2" pageNumber="211">
Aedeagus (
<figureCitation id="75CA2A01FF9BFFEACD18EFBFFCFF708B" box="[820,886,1465,1489]" captionStart="Figs 4-6" captionStartId="1.[624,670,810,834]" captionTargetBox="[632,1098,187,786]" captionTargetId="figure-244@1.[624,1106,171,804]" captionTargetPageId="1" captionText="Figs 4-6.Rhyparus anichtchenkoisp. nov.: 4- head, ♂, holotype; 5- last abdominal ventrite and pygidium, ♀, paratype; 6- aedeagus, holotype. Scale lines: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13203380" httpUri="https://zenodo.org/record/13203380/files/figure.png" pageId="2" pageNumber="211">Fig. 6</figureCitation>
) elongate, with parameres very short, with phallobase distinctly elongate, relatively weakly curved.
</paragraph>
<paragraph id="ED4E3684FF9BFFEACA40ECA0FB1D73E4" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="211">211</paragraph>
<caption id="B98E660CFF9AFFEBCE5EEAB4FF4F75B2" pageId="3" pageNumber="212" startId="3.[114,175,178,202]" targetType="table">
<paragraph id="ED4E3684FF9AFFEBCE5EEAB4FF4F75B2" blockId="3.[114,1103,178,232]" pageId="3" pageNumber="212">
Table 1. Differential characters of
<taxonomicName id="2AF14D07FF9AFFEBCFD8EAB4FD737590" authorityName="Minkina" authorityYear="2023" box="[500,762,178,202]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="anichtchenkoi" status="sp. nov.">
<emphasis id="DF85EA96FF9AFFEBCFD8EAB4FD737590" box="[500,762,178,202]" italics="true" pageId="3" pageNumber="212">Rhyparus anichtchenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C4B657EDFF9AFFEBCD28EAB4FCD57593" box="[772,860,178,201]" pageId="3" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
and the most similar species.
</paragraph>
</caption>
<paragraph id="ED4E3684FF9AFFEBCEAEEB13FB967305" pageId="3" pageNumber="212">
<table id="9FF1C424FF9A0017CEAEEB13FBB77337" box="[130,1086,277,1645]" gridcols="6" gridrows="33" pageId="3" pageNumber="212">
<tr id="53C134C6FF9A0017CEAEEB13FBB77471" box="[130,1086,277,299]" gridrow="0" pageId="3" pageNumber="212" rowspan-3="1">
<th id="10105DBAFF9A0017CEAEEB13FE8E7471" box="[130,263,277,299]" gridcol="0" gridrow="0" pageId="3" pageNumber="212">species</th>
<th id="10105DBAFF9A0017CF30EB13FEBE7471" box="[284,311,277,299]" gridcol="1" gridrow="0" pageId="3" pageNumber="212">/</th>
<th id="10105DBAFF9A0017CF61EB13FD897471" box="[333,512,277,299]" gridcol="2" gridrow="0" pageId="3" pageNumber="212">
<taxonomicName id="2AF14D07FF9AFFEBCF61EB13FE387470" authorityName="Arrow" authorityYear="1905" box="[333,433,277,298]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="DF85EA96FF9AFFEBCF61EB13FE387470" box="[333,433,277,298]" italics="true" pageId="3" pageNumber="212">R. gracilis</emphasis>
</taxonomicName>
</th>
<th id="10105DBAFF9A0017CC7AEB13FCBC7471" box="[598,821,277,299]" gridcol="4" gridrow="0" pageId="3" pageNumber="212">
<taxonomicName id="2AF14D07FF9AFFEBCC7AEB13FD7E7470" authorityName="Minkina" authorityYear="2023" box="[598,759,277,298]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="anichtchenkoi">
<emphasis id="DF85EA96FF9AFFEBCC7AEB13FD7E7470" box="[598,759,277,298]" italics="true" pageId="3" pageNumber="212">R. anichtchenkoi</emphasis>
</taxonomicName>
</th>
<th id="10105DBAFF9A0017CD60EB13FBB77471" box="[844,1086,277,299]" gridcol="5" gridrow="0" pageId="3" pageNumber="212">
<taxonomicName id="2AF14D07FF9AFFEBCD60EB13FC5B7470" authorityName="Stebnicka" authorityYear="1998" box="[844,978,277,298]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="212" phylum="Arthropoda" rank="species" species="mokaiensis">
<emphasis id="DF85EA96FF9AFFEBCD60EB13FC5B7470" box="[844,978,277,298]" italics="true" pageId="3" pageNumber="212">R. mokaiensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="53C134C6FF9A0017CEAEEB36FBB7741C" box="[130,1086,304,326]" gridrow="1" pageId="3" pageNumber="212" rowspan-1="1" rowspan-3="1">
<th id="10105DBAFF9A0017CEAEEB36FE8E741C" box="[130,263,304,326]" gridcol="0" gridrow="1" pageId="3" pageNumber="212">character</th>
<td id="10105DBAFF9A0017CF61EB36FD89741C" box="[333,512,304,326]" gridcol="2" gridrow="1" pageId="3" pageNumber="212">Arrow, 1905</td>
<td id="10105DBAFF9A0017CC7AEB36FCBC741C" box="[598,821,304,326]" gridcol="4" gridrow="1" pageId="3" pageNumber="212">sp. nov.</td>
<td id="10105DBAFF9A0017CD60EB36FBB7741C" box="[844,1086,304,326]" gridcol="5" gridrow="1" pageId="3" pageNumber="212">
<bibRefCitation id="89604B75FF9AFFEBCD60EB36FC6E741F" author="Stebnicka Z." box="[844,999,304,326]" pageId="3" pageNumber="212" pagination="833 - 895" refId="ref2810" refString="Stebnicka Z. 1998. The Aphodiinae of New Guinea, the Bismarck Archipelago and the Solomon Islands. Invertebrate Taxonomy 12 (5): 833 - 895." type="journal article" year="1998">Stebnicka, 1998</bibRefCitation>
</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEB57FBB774D8" box="[130,1086,337,386]" gridrow="2" pageId="3" pageNumber="212" rowspan-3="1">
<th id="10105DBAFF9A0017CEAEEB57FE8E74D8" box="[130,263,337,386]" gridcol="0" gridrow="2" pageId="3" pageNumber="212">proportions body</th>
<td id="10105DBAFF9A0017CF30EB57FEBE74D8" box="[284,311,337,386]" gridcol="1" gridrow="2" pageId="3" pageNumber="212">of</td>
<td id="10105DBAFF9A0017CF61EB57FD8974D8" box="[333,512,337,386]" gridcol="2" gridrow="2" pageId="3" pageNumber="212">distinctly elongate</td>
<td id="10105DBAFF9A0017CC7AEB57FCBC74D8" box="[598,821,337,386]" gridcol="4" gridrow="2" pageId="3" pageNumber="212">moderately elongate</td>
<td id="10105DBAFF9A0017CD60EB57FBB774D8" box="[844,1086,337,386]" gridcol="5" gridrow="2" pageId="3" pageNumber="212">moderately elongate</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEB8BFBB774F9" box="[130,1086,397,419]" gridrow="3" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="10105DBAFF9A0017CF61EB8BFDC974F9" box="[333,576,397,419]" colspan="2" colspanRight="1" gridcol="2" gridrow="3" pageId="3" pageNumber="212">all intercostae matt,</td>
<td id="10105DBAFF9A0017CD60EB8BFBB774F9" box="[844,1086,397,419]" gridcol="5" gridrow="3" pageId="3" pageNumber="212">all intercostae matt,</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEBAFFBB77480" box="[130,1086,425,474]" gridrow="4" pageId="3" pageNumber="212" rowspan-1="1">
<th id="10105DBAFF9A0017CEAEEBAFFE8E7480" box="[130,263,425,474]" gridcol="0" gridrow="4" pageId="3" pageNumber="212">shine</th>
<td id="10105DBAFF9A0017CF61EBAFFDC97480" box="[333,576,425,474]" colspan="2" colspanRight="1" gridcol="2" gridrow="4" pageId="3" pageNumber="212">rarely visible specimens with all intercostae</td>
<td id="10105DBAFF9A0017CC7AEBAFFCBC7480" box="[598,821,425,474]" gridcol="4" gridrow="4" pageId="3" pageNumber="212">all costae shiny</td>
<td id="10105DBAFF9A0017CD60EBAFFBB77480" box="[844,1086,425,474]" gridcol="5" gridrow="4" pageId="3" pageNumber="212">rarely visible specimens with all intercostae</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEBE6FBB774AC" box="[130,1086,480,502]" gridrow="5" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CF61EBE6FD8974AC" box="[333,512,480,502]" gridcol="2" gridrow="5" pageId="3" pageNumber="212">shiny</td>
<td id="10105DBAFF9A0017CD60EBE6FBB774AC" box="[844,1086,480,502]" gridcol="5" gridrow="5" pageId="3" pageNumber="212">shiny</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEBFBFBB77749" box="[130,1086,509,531]" gridrow="6" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="10105DBAFF9A0017CF61EBFBFDC97749" box="[333,576,509,531]" colspan="2" colspanRight="1" gridcol="2" gridrow="6" pageId="3" pageNumber="212">weakly do moderately</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE81EFBB77774" box="[130,1086,536,558]" gridrow="7" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="10105DBAFF9A0017CF61E81EFDC97774" box="[333,576,536,558]" colspan="2" colspanRight="1" gridcol="2" gridrow="7" pageId="3" pageNumber="212">distinctly developed;</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE832FBB777E2" box="[130,1086,564,696]" gridrow="8" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE832FE8E77E2" box="[130,263,564,696]" gridcol="0" gridrow="8" pageId="3" pageNumber="212">lateral lobes pronotum</th>
<td id="10105DBAFF9A0017CF30E832FEBE77E2" box="[284,311,564,696]" gridcol="1" gridrow="8" pageId="3" pageNumber="212">of</td>
<td id="10105DBAFF9A0017CF61E832FDC977E2" box="[333,576,564,696]" colspan="2" colspanRight="1" gridcol="2" gridrow="8" pageId="3" pageNumber="212">usually posterior lobes higher than anterior, rarely anterior and posterior lobes with similar high (especially</td>
<td id="10105DBAFF9A0017CC7AE832FCBC77E2" box="[598,821,564,696]" gridcol="4" gridrow="8" pageId="3" pageNumber="212">moderately distinctly developed; anterior and posteriori lobes with similar high</td>
<td id="10105DBAFF9A0017CD60E832FBB777E2" box="[844,1086,564,696]" gridcol="5" gridrow="8" pageId="3" pageNumber="212">moderately distinctly developed; anterior lobes somewhat higher than posterior</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE8B8FBB7778E" box="[130,1086,702,724]" gridrow="9" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="10105DBAFF9A0017CF61E8B8FDC9778E" box="[333,576,702,724]" colspan="2" colspanRight="1" gridcol="2" gridrow="9" pageId="3" pageNumber="212">when both are relatively</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE8DCFBB777AA" box="[130,1086,730,752]" gridrow="10" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="10105DBAFF9A0017CF61E8DCFDC977AA" box="[333,576,730,752]" colspan="2" colspanRight="1" gridcol="2" gridrow="10" pageId="3" pageNumber="212">weakly developed)</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE8F0FBB77672" box="[130,1086,758,808]" gridrow="11" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE8F0FE8E7672" box="[130,263,758,808]" gridcol="0" gridrow="11" pageId="3" pageNumber="212">costae pronotum</th>
<td id="10105DBAFF9A0017CF30E8F0FEBE7672" box="[284,311,758,808]" gridcol="1" gridrow="11" pageId="3" pageNumber="212">of</td>
<td id="10105DBAFF9A0017CF61E8F0FDC97672" box="[333,576,758,808]" colspan="2" colspanRight="1" gridcol="2" gridrow="11" pageId="3" pageNumber="212">relatively weakly to distinctly developed</td>
<td id="10105DBAFF9A0017CC7AE8F0FCBC7672" box="[598,821,758,808]" gridcol="4" gridrow="11" pageId="3" pageNumber="212">distinctly to very distinctly developed</td>
<td id="10105DBAFF9A0017CD60E8F0FBB77672" box="[844,1086,758,808]" gridcol="5" gridrow="11" pageId="3" pageNumber="212">distinctly developed</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE929FBB7761F" box="[130,1086,815,837]" gridrow="12" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE929FEBE761F" box="[130,311,815,837]" colspan="2" colspanRight="1" gridcol="0" gridrow="12" pageId="3" pageNumber="212">median costae of</th>
<td id="10105DBAFF9A0017CF61E929FD89761F" box="[333,512,815,837]" gridcol="2" gridrow="12" pageId="3" pageNumber="212">almost paralel</td>
<td id="10105DBAFF9A0017CC3BE929FDC9761F" box="[535,576,815,837]" gridcol="3" gridrow="12" pageId="3" pageNumber="212">to</td>
<td id="10105DBAFF9A0017CC7AE929FCBC761F" box="[598,821,815,837]" gridcol="4" gridrow="12" pageId="3" pageNumber="212">very distinctly</td>
<td id="10105DBAFF9A0017CD60E929FBB7761F" box="[844,1086,815,837]" gridcol="5" gridrow="12" pageId="3" pageNumber="212">more or less distinctly</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE94CFBB7763A" box="[130,1086,842,864]" gridrow="13" pageId="3" pageNumber="212" rowspan-1="1">
<th id="10105DBAFF9A0017CEAEE94CFE8E763A" box="[130,263,842,864]" gridcol="0" gridrow="13" pageId="3" pageNumber="212">pronotum</th>
<td id="10105DBAFF9A0017CF61E94CFDC9763A" box="[333,576,842,864]" colspan="2" colspanRight="1" gridcol="2" gridrow="13" pageId="3" pageNumber="212">distinctly convergent</td>
<td id="10105DBAFF9A0017CC7AE94CFCBC763A" box="[598,821,842,864]" gridcol="4" gridrow="13" pageId="3" pageNumber="212">convergent</td>
<td id="10105DBAFF9A0017CD60E94CFBB7763A" box="[844,1086,842,864]" gridcol="5" gridrow="13" pageId="3" pageNumber="212">convergent</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE96EFBB77624" box="[130,1086,872,894]" gridrow="14" pageId="3" pageNumber="212" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="10105DBAFF9A0017CEAEE96EFE8E7624" box="[130,263,872,894]" gridcol="0" gridrow="14" pageId="3" pageNumber="212">second pair</th>
<td id="10105DBAFF9A0017CF30E96EFEBE7624" box="[284,311,872,894]" gridcol="1" gridrow="14" pageId="3" pageNumber="212">of</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE985FBB776C3" box="[130,1086,899,921]" gridrow="15" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE985FE8E76C3" box="[130,263,899,921]" gridcol="0" gridrow="15" pageId="3" pageNumber="212">costae</th>
<td id="10105DBAFF9A0017CF30E985FEBE76C3" box="[284,311,899,921]" gridcol="1" gridrow="15" pageId="3" pageNumber="212">of</td>
<td id="10105DBAFF9A0017CF61E985FDC976C3" box="[333,576,899,921]" colspan="2" colspanRight="1" gridcol="2" gridrow="15" pageId="3" pageNumber="212">Zajmują about 1/3 of</td>
<td id="10105DBAFF9A0017CC7AE985FCBC76C3" box="[598,821,899,921]" gridcol="4" gridrow="15" pageId="3" pageNumber="212">no longer than 1/3 of</td>
<td id="10105DBAFF9A0017CD60E985FBB776C3" box="[844,1086,899,921]" gridcol="5" gridrow="15" pageId="3" pageNumber="212">Zajmują about 1/3 of</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE999FBB776EF" box="[130,1086,927,949]" gridrow="16" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE999FEBE76EF" box="[130,311,927,949]" colspan="2" colspanRight="1" gridcol="0" gridrow="16" pageId="3" pageNumber="212">pronotum in basal</th>
<td id="10105DBAFF9A0017CF61E999FDC976EF" box="[333,576,927,949]" colspan="2" colspanRight="1" gridcol="2" gridrow="16" pageId="3" pageNumber="212">length of pronotum</td>
<td id="10105DBAFF9A0017CC7AE999FCBC76EF" box="[598,821,927,949]" gridcol="4" gridrow="16" pageId="3" pageNumber="212">length of pronotum</td>
<td id="10105DBAFF9A0017CD60E999FBB776EF" box="[844,1086,927,949]" gridcol="5" gridrow="16" pageId="3" pageNumber="212">length of pronotum</td>
</tr>
<tr id="53C134C6FF9A0017CEAEE9BDFBB7768A" box="[130,1086,955,976]" gridrow="17" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE9BDFBB7768A" box="[130,1086,955,976]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="3" pageNumber="212">part</th>
</tr>
<tr id="53C134C6FF9A0017CEAEE9D1FBB77153" box="[130,1086,983,1033]" gridrow="18" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEE9D1FEBE7153" box="[130,311,983,1033]" colspan="2" colspanRight="1" gridcol="0" gridrow="18" pageId="3" pageNumber="212">Wgłębienia on pronotum</th>
<td id="10105DBAFF9A0017CF61E9D1FD897153" box="[333,512,983,1033]" gridcol="2" gridrow="18" pageId="3" pageNumber="212">moderately large deep</td>
<td id="10105DBAFF9A0017CC3BE9D1FDC97153" box="[535,576,983,1033]" gridcol="3" gridrow="18" pageId="3" pageNumber="212">and</td>
<td id="10105DBAFF9A0017CC7AE9D1FCBC7153" box="[598,821,983,1033]" gridcol="4" gridrow="18" pageId="3" pageNumber="212">very large and deep</td>
<td id="10105DBAFF9A0017CD60E9D1FBB77153" box="[844,1086,983,1033]" gridcol="5" gridrow="18" pageId="3" pageNumber="212">large and deep</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEE16FBB77122" box="[130,1086,1040,1144]" gridrow="19" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEEE16FE8E7122" box="[130,263,1040,1144]" gridcol="0" gridrow="19" pageId="3" pageNumber="212">punctation median intercostae pronotum</th>
<td id="10105DBAFF9A0017CF30EE16FEBE7122" box="[284,311,1040,1144]" gridcol="1" gridrow="19" pageId="3" pageNumber="212">of of</td>
<td id="10105DBAFF9A0017CF61EE16FDC97122" box="[333,576,1040,1144]" colspan="2" colspanRight="1" gridcol="2" gridrow="19" pageId="3" pageNumber="212">dense, more or less distinctly concentrated in the middle</td>
<td id="10105DBAFF9A0017CC7AEE16FCBC7122" box="[598,821,1040,1144]" gridcol="4" gridrow="19" pageId="3" pageNumber="212">very dense, very distinctly concentrated in the middle</td>
<td id="10105DBAFF9A0017CD60EE16FBB77122" box="[844,1086,1040,1144]" gridcol="5" gridrow="19" pageId="3" pageNumber="212">very dense, distinctly concentrated in the middle</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEE79FBB771CF" box="[130,1086,1151,1173]" gridrow="20" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EE79FBB771CF" box="[844,1086,1151,1173]" gridcol="5" gridrow="20" pageId="3" pageNumber="212">in part of specimens</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEE9DFBB771EB" box="[130,1086,1179,1201]" gridrow="21" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EE9DFBB771EB" box="[844,1086,1179,1201]" gridcol="5" gridrow="21" pageId="3" pageNumber="212">with three rows of</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEEB1FBB77197" box="[130,1086,1207,1229]" gridrow="22" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EEB1FBB77197" box="[844,1086,1207,1229]" gridcol="5" gridrow="22" pageId="3" pageNumber="212">punctures; in most of</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEED4FBB771B2" box="[130,1086,1234,1256]" gridrow="23" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EED4FBB771B2" box="[844,1086,1234,1256]" gridcol="5" gridrow="23" pageId="3" pageNumber="212">specimens with two</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEEE8FBB77061" box="[130,1086,1262,1339]" gridrow="24" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEEEE8FEBE7061" box="[130,311,1262,1339]" colspan="2" colspanRight="1" gridcol="0" gridrow="24" pageId="3" pageNumber="212">first and second intercostae of elytra</th>
<td id="10105DBAFF9A0017CF61EEE8FDC97061" box="[333,576,1262,1339]" colspan="2" colspanRight="1" gridcol="2" gridrow="24" pageId="3" pageNumber="212">always with two rows of punctures</td>
<td id="10105DBAFF9A0017CC7AEEE8FCBC7061" box="[598,821,1262,1339]" gridcol="4" gridrow="24" pageId="3" pageNumber="212">always with two rows of punctures</td>
<td id="10105DBAFF9A0017CD60EEE8FBB77061" box="[844,1086,1262,1339]" gridcol="5" gridrow="24" pageId="3" pageNumber="212">rows of punctures and some punctures between them; always:</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEF47FBB7700D" box="[130,1086,1345,1367]" gridrow="25" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EF47FBB7700D" box="[844,1086,1345,1367]" gridcol="5" gridrow="25" pageId="3" pageNumber="212">with two row of</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEF5AFBB77028" box="[130,1086,1372,1394]" gridrow="26" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EF5AFBB77028" box="[844,1086,1372,1394]" gridcol="5" gridrow="26" pageId="3" pageNumber="212">punctures with at least</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEF7EFBB770D4" box="[130,1086,1400,1422]" gridrow="27" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EF7EFBB770D4" box="[844,1086,1400,1422]" gridcol="5" gridrow="27" pageId="3" pageNumber="212">few punctures between</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEF95FBB770F3" box="[130,1086,1427,1449]" gridrow="28" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CD60EF95FBB770F3" box="[844,1086,1427,1449]" gridcol="5" gridrow="28" pageId="3" pageNumber="212">them</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEFB6FBB7709C" box="[130,1086,1456,1478]" gridrow="29" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-4="1">
<td id="10105DBAFF9A0017CF61EFB6FDC9709C" box="[333,576,1456,1478]" colspan="2" colspanRight="1" gridcol="2" gridrow="29" pageId="3" pageNumber="212">usually with three</td>
<td id="10105DBAFF9A0017CD60EFB6FBB7709C" box="[844,1086,1456,1478]" gridcol="5" gridrow="29" pageId="3" pageNumber="212">usually third (median)</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEFCAFBB77343" box="[130,1086,1484,1561]" gridrow="30" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEEFCAFEBE7343" box="[130,311,1484,1561]" colspan="2" colspanRight="1" gridcol="0" gridrow="30" pageId="3" pageNumber="212">third intercostae of elytra</th>
<td id="10105DBAFF9A0017CF61EFCAFDC97343" box="[333,576,1484,1561]" colspan="2" colspanRight="1" gridcol="2" gridrow="30" pageId="3" pageNumber="212">distinct row of punctures, rarely median row of</td>
<td id="10105DBAFF9A0017CC7AEFCAFCBC7343" box="[598,821,1484,1561]" gridcol="4" gridrow="30" pageId="3" pageNumber="212">usually third (median) row of punctures indistinct</td>
<td id="10105DBAFF9A0017CD60EFCAFBB77343" box="[844,1086,1484,1561]" gridcol="5" gridrow="30" pageId="3" pageNumber="212">row of punctures indistinct, not so rarely median row of</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEC19FBB7736F" box="[130,1086,1567,1589]" gridrow="31" pageId="3" pageNumber="212" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td id="10105DBAFF9A0017CF61EC19FD89736F" box="[333,512,1567,1589]" gridcol="2" gridrow="31" pageId="3" pageNumber="212">punctures absent</td>
<td id="10105DBAFF9A0017CD60EC19FBB7736F" box="[844,1086,1567,1589]" gridcol="5" gridrow="31" pageId="3" pageNumber="212">punctures absent</td>
</tr>
<tr id="53C134C6FF9A0017CEAEEC3AFBB77337" box="[130,1086,1596,1645]" gridrow="32" pageId="3" pageNumber="212">
<th id="10105DBAFF9A0017CEAEEC3AFEBE7337" box="[130,311,1596,1645]" colspan="2" colspanRight="1" gridcol="0" gridrow="32" pageId="3" pageNumber="212">medio-internal caudal bulbs</th>
<td id="10105DBAFF9A0017CF61EC3AFDC97337" box="[333,576,1596,1645]" colspan="2" colspanRight="1" gridcol="2" gridrow="32" pageId="3" pageNumber="212">distinctly transverse</td>
<td id="10105DBAFF9A0017CC7AEC3AFCBC7337" box="[598,821,1596,1645]" gridcol="4" gridrow="32" pageId="3" pageNumber="212">distinctly transverse</td>
<td id="10105DBAFF9A0017CD60EC3AFBB77337" box="[844,1086,1596,1645]" gridcol="5" gridrow="32" pageId="3" pageNumber="212">moderately transverse</td>
</tr>
</table>
</paragraph>
<paragraph id="ED4E3684FF9AFFEBCE5EECA0FF1373E4" blockId="3.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="3" pageNumber="212">212</paragraph>
<paragraph id="ED4E3684FF9DFFECCE91EAD5FE1574F9" blockId="4.[189,669,211,419]" pageId="4" pageNumber="213">
<emphasis id="DF85EA96FF9DFFECCE91EAD5FEBB75B1" bold="true" box="[189,306,211,235]" pageId="4" pageNumber="213">Variation.</emphasis>
Total body length
<quantity id="2A099B61FF9DFFECCC2FEAD5FDDE75B1" box="[515,599,211,235]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="4" pageNumber="213" unit="mm" value="6.1">6.1 mm</quantity>
–
<quantity id="2A099B61FF9DFFECCC58EAD5FF637453" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="213" unit="mm" value="6.8">6.8 mm</quantity>
. Punctation of pronotum can be more or less dense. Second pair of costae of pronotum more or less distinctly shortened basally. Median row of punctures on third intercostae of elytra may be more or less distinctly developed.
</paragraph>
<paragraph id="ED4E3684FF9DFFECCE91EBCEFD937701" blockId="4.[189,668,456,603]" pageId="4" pageNumber="213">
<emphasis id="DF85EA96FF9DFFECCE91EBCEFE3574BA" bold="true" box="[189,444,456,480]" pageId="4" pageNumber="213">Sexual dimorphism.</emphasis>
Typical for the members of the genus. Pygidium and last abdominal ventrite of male as in fig. 2. Pygidium and last abdominal ventrite of female not modified, as in fig. 5.
</paragraph>
</subSubSection>
<subSubSection id="A5EB650FFF9DFFECCE91E886FE8F717D" pageId="4" pageNumber="213" type="etymology">
<paragraph id="ED4E3684FF9DFFECCE91E886FDA57649" blockId="4.[189,668,640,787]" pageId="4" pageNumber="213">
<emphasis id="DF85EA96FF9DFFECCE91E886FEB677C2" bold="true" box="[189,319,640,664]" pageId="4" pageNumber="213">Etymology.</emphasis>
I dedicate the name of the new species to Alexander Anichtchenko – entomologist,
<taxonomicName id="2AF14D07FF9DFFECCF73E8B8FE42778C" authorityName="Latreille" authorityYear="1802" box="[351,459,702,726]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="213" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
specialist, friend who organized material for me with specimens of undescribed species.
</paragraph>
<paragraph id="ED4E3684FF9DFFECCE91E93EFE8F717D" blockId="4.[189,669,824,1063]" pageId="4" pageNumber="213">
<emphasis id="DF85EA96FF9DFFECCE91E93EFE95760A" bold="true" box="[189,284,824,848]" pageId="4" pageNumber="213">Affinity.</emphasis>
The newly described species appears to be intermediate between
<taxonomicName id="2AF14D07FF9DFFECCE91E970FDBB76D4" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[189,562,886,910]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="213" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="DF85EA96FF9DFFECCE91E970FE0176D4" box="[189,392,886,910]" italics="true" pageId="4" pageNumber="213">Rhyparus gracilis</emphasis>
Arrow, 1905
</taxonomicName>
and
<taxonomicName id="2AF14D07FF9DFFECCC53E970FE7276F7" authority="Stebnicka, 1998" authorityName="Stebnicka" authorityYear="1998" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="213" phylum="Arthropoda" rank="species" species="mokaiensis">
<emphasis id="DF85EA96FF9DFFECCC53E970FEBA76F7" italics="true" pageId="4" pageNumber="213">R. mokaiensis</emphasis>
<bibRefCitation id="89604B75FF9DFFECCF6CE992FE7276F7" author="Stebnicka Z." box="[320,507,916,941]" pageId="4" pageNumber="213" pagination="833 - 895" refId="ref2810" refString="Stebnicka Z. 1998. The Aphodiinae of New Guinea, the Bismarck Archipelago and the Solomon Islands. Invertebrate Taxonomy 12 (5): 833 - 895." type="journal article" year="1998">Stebnicka, 1998</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="2AF14D07FF9DFFECCC28E993FDF176F7" authorityName="Arrow" authorityYear="1905" box="[516,632,917,941]" class="Insecta" family="Scarabaeidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="213" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="DF85EA96FF9DFFECCC28E993FDF176F7" box="[516,632,917,941]" italics="true" pageId="4" pageNumber="213">R. gracilis</emphasis>
</taxonomicName>
is one of the most variable species within the genus and for this reason differentiating features are compared in detail in the table below.
</paragraph>
</subSubSection>
</treatment>
</document>
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