treatments-xml/data/0A/70/87/0A7087BBFFAAFFBE41897E0DFD46015C.xml
2024-06-21 12:29:01 +02:00

255 lines
28 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3699AADC4C4D3E46E1BEB3FCA47CE29D" ID-DOI="10.11646/zootaxa.5448.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="11232285" ID-ZooBank="B82C87CC-501B-4844-ACAB-21BD1377E2A3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716280072248" checkinUser="plazi" docAuthor="Kehlmaier, Christian, Kuchling, Gerald, Shea, Glenn, Georges, Arthur, Zhang, Xiuwen, Fritz, Uwe &amp; Thomson, Scott" docDate="2024" docId="0A7087BBFFAAFFBE41897E0DFD46015C" docLanguage="en" docName="zootaxa.5448.4.6.pdf" docOrigin="Zootaxa 5448 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5448.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chelodina rugosa Ogilby 1890" docType="treatment" docVersion="2" lastPageNumber="544" masterDocId="F649FFC3FFA1FFB0411E795CFF950041" masterDocTitle="History and historical DNA: Identity of Chelodina intergularis Fry, 1915 and type localities of C. intergularis and C. rugosa Ogilby, 1890" masterLastPageNumber="555" masterPageNumber="531" pageNumber="542" updateTime="1716281573474" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2E7A365D5E37D07F52CACF1A2D405226" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="20CC3666B46B31AB28C171DD78628AE5">
<mods:title id="9ED4493CBA113750FE8E73D17246F9D1">History and historical DNA: Identity of Chelodina intergularis Fry, 1915 and type localities of C. intergularis and C. rugosa Ogilby, 1890</mods:title>
</mods:titleInfo>
<mods:name id="1386E292A08B6380631CD541E1BD925C" type="personal">
<mods:role id="AC8BD47A57862E7137951740115889CF">
<mods:roleTerm id="D82A3AD1551DCC2CBEF688698773186F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB7C72B22023C68EA4B5D1D2AB07593E">Kehlmaier, Christian</mods:namePart>
<mods:affiliation id="4F5A2BE8164333EE5A30A69E57D5D11B">Museum of Zoology, Senckenberg Dresden, 01109 Dresden, Germany</mods:affiliation>
</mods:name>
<mods:name id="0625F76F93172CAE40F9200C9903ADB8" type="personal">
<mods:role id="CE511EAC4825ACC80704A7185D6BC1AD">
<mods:roleTerm id="608B094AB7AE0579509E6E96B3041328">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FF336DB0072673F3D24D07C2C84674F">Kuchling, Gerald</mods:namePart>
<mods:affiliation id="1D2400824D1254A0A0DC237E9A3CFFC7">School of Biological Sciences, The University of Western Australia, Crawley, WA 6009, Australia</mods:affiliation>
</mods:name>
<mods:name id="20028B1BC85D1CC765E7A9229028C8D6" type="personal">
<mods:role id="2C94D5687D8DF6CE1D6A4B1DA52D7BF9">
<mods:roleTerm id="07019B03D1F4515C79E7E614F3F3D5C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAA6ADCD935B875954C875480BA15380">Shea, Glenn</mods:namePart>
<mods:affiliation id="01C0FE009A20188CEADBB42DBCAE7125">Sydney School of Veterinary Science, B 01, University of Sydney, NSW 2006, Australia &amp; Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney, NSW 2010, Australia</mods:affiliation>
</mods:name>
<mods:name id="25FAF2EB1A3C0EFA53D20DA17407DDD2" type="personal">
<mods:role id="0AC46778A08C70434F709C25DA81FC2A">
<mods:roleTerm id="43424B82FE4B355B0843D0123C754B9A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66CCDB8DC8AEBECAA4DDCD6526E14CE4">Georges, Arthur</mods:namePart>
<mods:affiliation id="B81A601795D7E3361911E8D5CAFC9FAA">Institute for Applied Ecology, University of Canberra, Canberra, ACT 2601, Australia</mods:affiliation>
</mods:name>
<mods:name id="5F60553C87EE937AABE9B70BC4425F38" type="personal">
<mods:role id="5CC0F196193F4F4CDADFD59BDA2F09D8">
<mods:roleTerm id="583F1D9054D5D3C04F947833A4DEF310">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F4B84C1D9065488B5E03BA117E542A0">Zhang, Xiuwen</mods:namePart>
<mods:affiliation id="988191C6E5D0C17CC42F87D24C0EA404">Institute for Applied Ecology, University of Canberra, Canberra, ACT 2601, Australia</mods:affiliation>
</mods:name>
<mods:name id="E99F85AFC705E24D781A9CA00B1EE8E9" type="personal">
<mods:role id="72AF31CFBCCFF18245A596EB30B649D3">
<mods:roleTerm id="5129FEC771748F1666D2CF0BC7742F5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DAB23CBF29E639A6BC892469DBC16864">Fritz, Uwe</mods:namePart>
<mods:affiliation id="7231386B353C849F0E567AFCD7DD1678">Museum of Zoology, Senckenberg Dresden, 01109 Dresden, Germany</mods:affiliation>
</mods:name>
<mods:name id="78F686D2B58806F2F491A923B76CA289" type="personal">
<mods:role id="910D00011076E323C1A94E3B66535DE5">
<mods:roleTerm id="3AE7309B13DDCABE951082CFA1AA55CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BF5CE10196EA3366A467557E07C1E3A">Thomson, Scott</mods:namePart>
<mods:affiliation id="2B217E622264FA23F1DFE54FF0AE474D">Centro de Estudos dos Quelônios da Amazônia, Manaus 69055 - 010, Brazil &amp; Research Institute for the Environment and Livelihoods, Charles Darwin University, Darwin, NT 0909, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="56BB4F8A615CCE5F6602C38712AFE8C5">text</mods:typeOfResource>
<mods:relatedItem id="F470878F696C5223366923023AF3A493" type="host">
<mods:titleInfo id="C1D781A309F4F38D967226B259D77D0A">
<mods:title id="C7C805D195A896502EF17DEB7A592702">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5D29EF1E91AB19543665572F153E039B">
<mods:date id="BD113DBB4EEAE4D913C60D8C1F84171D">2024</mods:date>
<mods:detail id="882ED443C3DD41FEE511E76B9125B4E2" type="pubDate">
<mods:number id="D6B707A4CB897C966D08B5682ECA70C0">2024-05-14</mods:number>
</mods:detail>
<mods:detail id="A97A6FBA5A20264770B27C6E44B6DD06" type="volume">
<mods:number id="BAEFA9AC7D3B260C84FC36A61162C190">5448</mods:number>
</mods:detail>
<mods:detail id="980BE5096823EB32589F142594D7F42F" type="issue">
<mods:number id="6F95220DDA09F3CAE2E227CF4291D75C">4</mods:number>
</mods:detail>
<mods:extent id="0E31E4632EA1B1CD0AA6A4BC719E05FE" unit="page">
<mods:start id="6556350FF199FA7BF6CEA2FAA5157098">531</mods:start>
<mods:end id="7BFF513DCE0A19EE25C04D9683481460">555</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="CA69AA24AF4442E6B0EC6D7A1BB339FF">
<mods:url id="AD9F80CD98E6AB1B6D48507FE89CB523">http://dx.doi.org/10.11646/zootaxa.5448.4.6</mods:url>
</mods:location>
<mods:classification id="BC1288D030D36EBB83F6DB29C7CA49F1">journal article</mods:classification>
<mods:identifier id="D516C3FE732E5AF4A72687E5C9C97221" type="DOI">10.11646/zootaxa.5448.4.6</mods:identifier>
<mods:identifier id="8987C169AF707A57DC4D03100EBC2098" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1A3F2282911884D326972537FC291400" type="Zenodo-Dep">11232285</mods:identifier>
<mods:identifier id="42C05D37FFB712FC57E23E0A88F1466C" type="ZooBank">B82C87CC-501B-4844-ACAB-21BD1377E2A3</mods:identifier>
</mods:mods>
<treatment id="0A7087BBFFAAFFBE41897E0DFD46015C" LSID="urn:lsid:plazi:treatment:0A7087BBFFAAFFBE41897E0DFD46015C" httpUri="http://treatment.plazi.org/id/0A7087BBFFAAFFBE41897E0DFD46015C" lastPageId="14" lastPageNumber="544" pageId="11" pageNumber="542">
<subSubSection id="CAC36526FFAAFFBB41897E0DFD86072A" box="[151,531,1873,1900]" pageId="11" pageNumber="542" type="nomenclature">
<paragraph id="826636ADFFAAFFBB41897E0DFD86072A" blockId="11.[151,531,1873,1900]" box="[151,531,1873,1900]" pageId="11" pageNumber="542">
<heading id="D92E81C1FFAAFFBB41897E0DFD86072A" bold="true" box="[151,531,1873,1900]" fontSize="11" level="1" pageId="11" pageNumber="542" reason="1">
<taxonomicName id="45D94D2EFFAAFFBB41897E0DFD86072A" ID-CoL="87WDQ" authority="Ogilby, 1890" authorityName="Ogilby" authorityYear="1890" box="[151,531,1873,1900]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B0ADEABFFFAAFFBB41897E0DFD86072A" bold="true" box="[151,531,1873,1900]" pageId="11" pageNumber="542">
<emphasis id="B0ADEABFFFAAFFBB41897E0DFEFA072D" bold="true" box="[151,367,1873,1900]" italics="true" pageId="11" pageNumber="542">Chelodina rugosa</emphasis>
<bibRefCitation id="E6484B5CFFAAFFBB40697E0DFD86072A" author="Ogilby, J. D." box="[375,531,1873,1899]" pageId="11" pageNumber="542" pagination="56 - 59" refId="ref17547" refString="Ogilby, J. D. (1890) Description of a new Australian tortoise. Records of the Australian Museum, 1 (3), 56 - 59, pl. VIII. https: // doi. org / 10.3853 / j. 0067 - 1975.1.1890.1226" type="journal article" year="1890">Ogilby, 1890</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CAC36526FFAAFFBB41897ECAFC9A07EF" box="[151,783,1942,1966]" pageId="11" pageNumber="542" type="materials_examined">
<paragraph id="826636ADFFAAFFBB41897ECAFC9A07EF" blockId="11.[151,783,1942,1966]" box="[151,783,1942,1966]" pageId="11" pageNumber="542">
<materialsCitation id="32B13CF0FFAAFFBB41897ECAFC9A07EF" box="[151,783,1942,1966]" collectionCode="AM" pageId="11" pageNumber="542" specimenCode="R6256" specimenCount="1" typeStatus="holotype">
<typeStatus id="5D62880FFFAAFFBB41897ECAFF6C07EF" box="[151,249,1942,1966]" pageId="11" pageNumber="542" type="holotype">Holotype</typeStatus>
: Australian Museum, Sydney (
<collectionCode id="E4C8AE68FFAAFFBB432E7ECAFDCF07EF" box="[560,602,1942,1966]" country="Australia" name="Australian Museum" pageId="11" pageNumber="542" type="Museum">AM</collectionCode>
<specimenCode id="D27F9ED6FFAAFFBB437F7ECAFD3D07EF" box="[609,680,1942,1966]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="542">R6256</specimenCode>
),
<figureCitation id="1AE22A28FFAAFFBB43A77ECAFC9A07EF" box="[697,783,1942,1966]" captionStart="FIGURE 3" captionStartId="12.[151,250,1232,1256]" captionTargetBox="[170,1417,292,1190]" captionTargetId="figure-72@12.[151,1436,273,1209]" captionTargetPageId="12" captionText="FIGURE 3. Holotype of Chelodina rugosa, Australian Museum, Sydney (AM R6256). Photos: G. Shea." figureDoi="http://doi.org/10.5281/zenodo.11232291" httpUri="https://zenodo.org/record/11232291/files/figure.png" pageId="11" pageNumber="542">Figure 3</figureCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CAC36526FFAAFFBE41897E80FD46015C" lastPageId="14" lastPageNumber="545" pageId="11" pageNumber="542" type="description">
<paragraph id="826636ADFFAAFFBC41897E80FB1A00B8" blockId="11.[151,1436,2012,2039]" lastBlockId="12.[151,1436,151,249]" lastPageId="12" lastPageNumber="543" pageId="11" pageNumber="542">
<emphasis id="B0ADEABFFFAAFFBB41897E80FEDB07B7" bold="true" box="[151,334,2012,2038]" pageId="11" pageNumber="542">Measurements.</emphasis>
Straight carapacial length (midline)
<quantity id="45219B48FFAAFFBB43F37E80FCF607B6" box="[749,867,2012,2039]" metricMagnitude="-1" metricUnit="m" metricValue="2.558" pageId="11" pageNumber="542" unit="mm" value="255.8">255.8 mm</quantity>
; maximum carapacial width
<quantity id="45219B48FFAAFFBB45B77E80FA8A07B6" box="[1193,1311,2012,2039]" metricMagnitude="-1" metricUnit="m" metricValue="1.7169999999999999" pageId="11" pageNumber="542" unit="mm" value="171.7">171.7 mm</quantity>
; carapacial width at mid-bridge level 160.0 mm; maximum plastral length (from level of paired caudal tips to extreme anterior margin)
<quantity id="45219B48FFADFFBC41E479E7FEE70094" box="[250,370,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="2.114" pageId="12" pageNumber="543" unit="mm" value="211.4">211.4 mm</quantity>
; midline plastral length
<quantity id="45219B48FFADFFBC438E79E7FC9C0094" box="[656,777,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.9169999999999998" pageId="12" pageNumber="543" unit="mm" value="191.7">191.7 mm</quantity>
; plastral width at anterior extremity of bridge
<quantity id="45219B48FFADFFBC443379E6FA0D0094" box="[1325,1432,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="9.57" pageId="12" pageNumber="543" unit="mm" value="95.7">95.7 mm</quantity>
; maximum posterior plastral width
<quantity id="45219B48FFADFFBC43057983FD1700B8" box="[539,642,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="9.29" pageId="12" pageNumber="543" unit="mm" value="92.9">92.9 mm</quantity>
; total shell depth at middle of bridge
<quantity id="45219B48FFADFFBC453A7983FB1E00B8" box="[1060,1163,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="9.39" pageId="12" pageNumber="543" unit="mm" value="93.9">93.9 mm</quantity>
.
</paragraph>
<caption id="D6A66625FFADFFBC41897D8CFB2804A9" ID-DOI="http://doi.org/10.5281/zenodo.11232291" ID-Zenodo-Dep="11232291" box="[151,1213,1232,1257]" httpUri="https://zenodo.org/record/11232291/files/figure.png" pageId="12" pageNumber="543" startId="12.[151,250,1232,1256]" targetBox="[170,1417,292,1190]" targetPageId="12" targetType="figure">
<paragraph id="826636ADFFADFFBC41897D8CFB2804A9" blockId="12.[151,1213,1232,1257]" box="[151,1213,1232,1257]" pageId="12" pageNumber="543">
<emphasis id="B0ADEABFFFADFFBC41897D8CFE8604A9" bold="true" box="[151,275,1232,1256]" pageId="12" pageNumber="543">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="45D94D2EFFADFFBC40877D8CFDDA04A9" authorityName="Ogilby" authorityYear="1890" box="[409,591,1232,1256]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B0ADEABFFFADFFBC40877D8CFDDA04A9" box="[409,591,1232,1256]" italics="true" pageId="12" pageNumber="543">Chelodina rugosa</emphasis>
</taxonomicName>
, Australian Museum, Sydney (AM R6256). Photos: G. Shea.
</paragraph>
</caption>
<caption id="D6A66625FFADFFBC41897F25FE1D0760" ID-DOI="http://doi.org/10.5281/zenodo.11232293" ID-Zenodo-Dep="11232293" httpUri="https://zenodo.org/record/11232293/files/figure.png" pageId="12" pageNumber="543" startId="12.[151,250,1657,1681]" targetBox="[177,1423,1293,1618]" targetPageId="12" targetType="figure">
<paragraph id="826636ADFFADFFBC41897F25FE1D0760" blockId="12.[151,1437,1657,1825]" pageId="12" pageNumber="543">
<emphasis id="B0ADEABFFFADFFBC41897F25FE8706D0" bold="true" box="[151,274,1657,1681]" pageId="12" pageNumber="543">FIGURE 4.</emphasis>
Comparison of (A) the holotype of
<taxonomicName id="45D94D2EFFADFFBC43697F25FCBE06D0" authorityName="Ogilby" authorityYear="1890" box="[631,811,1657,1681]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B0ADEABFFFADFFBC43697F25FCBE06D0" box="[631,811,1657,1681]" italics="true" pageId="12" pageNumber="543">Chelodina rugosa</emphasis>
</taxonomicName>
(AM R6256), (B) the holotype of
<taxonomicName id="45D94D2EFFADFFBC459C7F26FA8006D0" authorityName="Fry" authorityYear="1915" box="[1154,1301,1657,1681]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="intergularis">
<emphasis id="B0ADEABFFFADFFBC459C7F26FA8006D0" box="[1154,1301,1657,1681]" italics="true" pageId="12" pageNumber="543">C. intergularis</emphasis>
</taxonomicName>
(AM R6255) and (C) an individual of
<taxonomicName id="45D94D2EFFADFFBC408E7FC2FDAE06F4" box="[400,571,1693,1717]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="12" pageNumber="543" phylum="Chordata" rank="species" species="burrungandjii">
<emphasis id="B0ADEABFFFADFFBC408E7FC2FDAE06F4" box="[400,571,1693,1717]" italics="true" pageId="12" pageNumber="543">C. burrungandjii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB9E57C4FFADFFBC435C7FC2FD2A06F4" box="[578,703,1693,1717]" pageId="12" pageNumber="543" sensu="stricto">sensu stricto</taxonomicNameLabel>
(UC 2101; redrawn from
<bibRefCitation id="E6484B5CFFADFFBC42D97FC1FB0806F4" author="Thomson, S. &amp; Kennett, R. &amp; Georges, A." box="[967,1181,1693,1717]" pageId="12" pageNumber="543" pagination="675 - 685" refId="ref18785" refString="Thomson, S., Kennett, R. &amp; Georges, A. (2000) A new species of long-necked turtle (Testudines: Chelidae) from the Arnhem Land Plateau, Northern Territory, Australia. Chelonian Conservation and Biology, 3 (4), 675 - 685." type="journal article" year="2000">
Thomson
<emphasis id="B0ADEABFFFADFFBC45337FC2FBCF06F4" box="[1069,1114,1693,1717]" italics="true" pageId="12" pageNumber="543">et al</emphasis>
. 2000
</bibRefCitation>
). Not to scale. Epidermal scutes (blue): G = gular; H = humeral; I = intergular; P = pectoral. Bony plates (black): Epi = epiplastron; Ent = entoplastron; Hyo = hyoplastron. In AM R6256 the bony sutures are visible at closer inspection through the translucent epidermal scutes. Drawings: S. Thomson.
</paragraph>
</caption>
<caption id="D6A66625FFACFFBD41897A61FEFB0383" ID-DOI="http://doi.org/10.5281/zenodo.11232295" ID-Zenodo-Dep="11232295" httpUri="https://zenodo.org/record/11232295/files/figure.png" pageId="13" pageNumber="544" startId="13.[151,250,829,853]" targetBox="[234,1345,186,795]" targetPageId="13" targetType="figure">
<paragraph id="826636ADFFACFFBD41897A61FEFB0383" blockId="13.[151,1437,829,962]" pageId="13" pageNumber="544">
<emphasis id="B0ADEABFFFACFFBD41897A61FE870317" bold="true" box="[151,274,829,854]" pageId="13" pageNumber="544">FIGURE 5.</emphasis>
Plastral aspects of
<taxonomicName id="45D94D2EFFACFFBD40CB7A62FD370317" authorityName="Cann" authorityYear="1997" box="[469,674,830,854]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="13" pageNumber="544" phylum="Chordata" rank="species" species="kuchlingi">
<emphasis id="B0ADEABFFFACFFBD40CB7A62FD370317" box="[469,674,830,854]" italics="true" pageId="13" pageNumber="544">Chelodina kuchlingi</emphasis>
</taxonomicName>
; specimens from the Western Australian Museum, Perth. From left to right: WAM R29411 (holotype), WAM R28116, WAM R28117 (some scute outlines in this specimen highlighted in blue). To scale, midline plastral length of holotype 185 mm. Note that the intergular reaches the anterior plastral rim only in the holotype. Photos: G. Kuchling.
</paragraph>
</caption>
<paragraph id="826636ADFFACFFBD41D97D48FF7704A3" blockId="13.[151,1437,1044,2042]" pageId="13" pageNumber="544">
<emphasis id="B0ADEABFFFACFFBD41D97D48FECC046F" bold="true" box="[199,345,1044,1070]" pageId="13" pageNumber="544">Description.</emphasis>
Specimen consists of a complete carapace and articulated plastron, the anterior thoracic centra are in situ as well as the posterior pelvis. Preserved separately are the 7
<superScript id="75AC9BE5FFACFFBD42617D64FC190407" attach="left" box="[895,908,1080,1094]" fontSize="6" pageId="13" pageNumber="544">th</superScript>
and 8
<superScript id="75AC9BE5FFACFFBD42D07D64FC4E0407" attach="left" box="[974,987,1080,1094]" fontSize="6" pageId="13" pageNumber="544">th</superScript>
cervical vertebrae. The scutes are in situ with this specimen. This specimen was clearly either collected alive or found recently deceased and then prepared curatorially for the museum. The entire carapace has a fine reticulate pattern to the scutes which is more rugose and antero-posteriorly aligned in the 2
<superScript id="75AC9BE5FFACFFBD430A7DF8FDB104F3" attach="left" box="[532,548,1188,1202]" fontSize="6" pageId="13" pageNumber="544">nd</superScript>
to 4
<superScript id="75AC9BE5FFACFFBD43487DF8FDF604F3" attach="left" box="[598,611,1188,1202]" fontSize="6" pageId="13" pageNumber="544">th</superScript>
vertebral scutes. This pattern is typical of
<taxonomicName id="45D94D2EFFACFFBD45207DF8FB3804FF" authorityName="Ogilby" authorityYear="1890" box="[1086,1197,1188,1214]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="13" pageNumber="544" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B0ADEABFFFACFFBD45207DF8FB3804FF" box="[1086,1197,1188,1214]" italics="true" pageId="13" pageNumber="544">C. rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB9E57C4FFACFFBD45AA7DF8FAB604FF" box="[1204,1315,1188,1214]" pageId="13" pageNumber="544" sensu="lato">sensu lato</taxonomicNameLabel>
as defined above.
</paragraph>
<paragraph id="826636ADFFACFFBD41D97DB0FB2305D7" blockId="13.[151,1437,1044,2042]" pageId="13" pageNumber="544">
The 4
<superScript id="75AC9BE5FFACFFBD40197DB0FE8104BB" attach="left" box="[263,276,1260,1274]" fontSize="6" pageId="13" pageNumber="544">th</superScript>
vertebral scute is reduced in size compared to other specimens of
<taxonomicName id="45D94D2EFFACFFBD42F17DB0FBC90547" authorityName="Ogilby" authorityYear="1890" box="[1007,1116,1260,1286]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="13" pageNumber="544" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B0ADEABFFFACFFBD42F17DB0FBC90547" box="[1007,1116,1260,1286]" italics="true" pageId="13" pageNumber="544">C. rugosa</emphasis>
</taxonomicName>
, with an extra scute between this and the 5
<superScript id="75AC9BE5FFACFFBD40377C4CFEA3055F" attach="left" box="[297,310,1296,1310]" fontSize="6" pageId="13" pageNumber="544">th</superScript>
vertebral which is also reduced. Between the 1
<superScript id="75AC9BE5FFACFFBD42237C4CFCDD055F" attach="left" box="[829,840,1296,1310]" fontSize="6" pageId="13" pageNumber="544">st</superScript>
costal on the left side and the 1
<superScript id="75AC9BE5FFACFFBD45BE7C4CFB3E055F" attach="left" box="[1184,1195,1296,1310]" fontSize="6" pageId="13" pageNumber="544">st</superScript>
vertebral scutes there is an extra anomalous scute which has encroached into the normal size and shape of both scutes that border it. The 2
<superScript id="75AC9BE5FFACFFBD41BB7C04FF200527" attach="left" box="[165,181,1368,1382]" fontSize="6" pageId="13" pageNumber="544">nd</superScript>
marginal is approximately 30% longer than the 3
<superScript id="75AC9BE5FFACFFBD43FD7C04FD640527" attach="none" box="[739,753,1368,1382]" fontSize="6" pageId="13" pageNumber="544">rd</superScript>
, otherwise marginals appear normal in growth (note damage left side below) and number 11 on each side. A pygal scute and cervical scute are also present.
</paragraph>
<paragraph id="826636ADFFACFFBD41D97CFCFD4B0643" blockId="13.[151,1437,1044,2042]" pageId="13" pageNumber="544">At the left side rear quarter of the carapace there is significant damage above the hind leg. This would appear to be the result of a bite from a predator. This damage has fully healed with overgrown scute material and hence occurred many years before the death of the animal.</paragraph>
<paragraph id="826636ADFFACFFBD41D97F50FEAD0763" blockId="13.[151,1437,1044,2042]" pageId="13" pageNumber="544">
The plastron is complete, includes the attached posterior pelvis, and also has a fine reticulate pattern throughout. The museum accession number (R 6256) is written in permanent ink on the left humeral scute. There is some old larger-font-sized handwritten text on the plastron, twice “Cape
<collectingRegion id="401DF84FFFACFFBD424D7F08FC18062F" box="[851,909,1620,1646]" country="United Kingdom" name="York" pageId="13" pageNumber="544">York</collectingRegion>
” in different orientations, one longitudinal, one transverse. The transverse inscription also has “Thorpe,” additional text is present but illegible. On the left rear of the plastron, within the anal scute, is a break where some bone has been lost, interestingly it is in a similar position to the injury of the carapace. There is evidence of some healing, and hence the injury occurred before the animal died. However, the plastral fracture is far less healed over than the carapace injury, indicating that the plastral damage occurred later.
</paragraph>
<paragraph id="826636ADFFACFFBD41D97E70FBB207F3" blockId="13.[151,1437,1044,2042]" pageId="13" pageNumber="544">
The scute pattern of the plastron is typical of
<taxonomicName id="45D94D2EFFACFFBD43A27E70FCBC0707" authorityName="Ogilby" authorityYear="1890" box="[700,809,1836,1862]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="13" pageNumber="544" phylum="Chordata" rank="species" species="rugosa">
<emphasis id="B0ADEABFFFACFFBD43A27E70FCBC0707" box="[700,809,1836,1862]" italics="true" pageId="13" pageNumber="544">C. rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AB9E57C4FFACFFBD42317E70FC340707" box="[815,929,1836,1862]" pageId="13" pageNumber="544" sensu="lato">sensu lato</taxonomicNameLabel>
. Importantly, this includes the shape, size and extent of the intergular scute. This scute is large, hexagonal though elongated. It partially divides the humerals and separates the pectorals. It is enclosed anteriorly by the gular scutes, which is the typical
<taxonomicName id="45D94D2EFFACFFBD45777E28FB4B07CF" authorityName="Fitzinger" authorityYear="1826" box="[1129,1246,1908,1934]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="13" pageNumber="544" phylum="Chordata" rank="genus">
<emphasis id="B0ADEABFFFACFFBD45777E28FB4B07CF" box="[1129,1246,1908,1934]" italics="true" pageId="13" pageNumber="544">Chelodina</emphasis>
</taxonomicName>
condition. There is a small indentation between the epiplastra/gulars at the anterior of the plastron.
</paragraph>
<paragraph id="826636ADFFACFFBE41D97EE0FD46015C" blockId="13.[151,1437,1044,2042]" lastBlockId="14.[151,1436,151,285]" lastPageId="14" lastPageNumber="545" pageId="13" pageNumber="544">
Internally, the first four thoracic centra are present.The 4
<superScript id="75AC9BE5FFACFFBD422C7EE0FCAA078B" attach="left" box="[818,831,1980,1994]" fontSize="6" pageId="13" pageNumber="544">th</superScript>
thoracic centrum is articulated and damaged anteriorly, the other thoracic centra seem to have become disarticulated over time, or during preparation, but were not cut out as the rib heads are largely undamaged. This differs from the
<taxonomicName id="45D94D2EFFAFFFBE424D79CBFC6D00F0" authorityName="Fry" authorityYear="1915" box="[851,1016,151,177]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="14" pageNumber="545" phylum="Chordata" rank="species" species="intergularis">
<emphasis id="B0ADEABFFFAFFFBE424D79CBFC6D00F0" box="[851,1016,151,177]" italics="true" pageId="14" pageNumber="545">C. intergularis</emphasis>
</taxonomicName>
<typeStatus id="5D62880FFFAFFFBE451E79CBFBA500F0" box="[1024,1072,151,177]" pageId="14" pageNumber="545">type</typeStatus>
where the centra were removed during preparation as indicated by the extensive rib damage. The pelvis is postero-laterally rotated (sensu
<bibRefCitation id="E6484B5CFFAFFFBE442C79E7FE8500B8" author="Thomson, S. A. &amp; Friol, N. R. &amp; White, A. &amp; Wedd, D. &amp; Georges, A." pageId="14" pageNumber="545" pagination="237 - 256" refId="ref18881" refString="Thomson, S. A., Friol, N. R., White, A., Wedd, D. &amp; Georges, A. (2023) The Australian gulf snapping turtle Elseya lavarackorum (Testudines: Chelidae) revisited - Is the late Pleistocene fossil species extant? Vertebrate Zoology, 73, 237 - 256. https: // doi. org / 10.3897 / vz. 73. e 99495" type="journal article" year="2023">
Thomson
<emphasis id="B0ADEABFFFAFFFBE41897983FF5D00B8" box="[151,200,223,249]" italics="true" pageId="14" pageNumber="545">et al</emphasis>
. 2023
</bibRefCitation>
) and attached to the 1
<superScript id="75AC9BE5FFAFFFBE431A7983FD9A00AC" attach="left" box="[516,527,223,237]" fontSize="6" pageId="14" pageNumber="545">st</superScript>
and 2
<superScript id="75AC9BE5FFAFFFBE434D7982FDF600AD" attach="left" box="[595,611,222,236]" fontSize="6" pageId="14" pageNumber="545">nd</superScript>
sacral ribs (ribs 1011) which are in situ. The pelvis covers the 8
<superScript id="75AC9BE5FFAFFFBE44257982FADD00AD" attach="left" box="[1339,1352,222,236]" fontSize="6" pageId="14" pageNumber="545">th</superScript>
pleural and extends onto both the 7
<superScript id="75AC9BE5FFAFFFBE40D2785EFE4C0151" attach="left" box="[460,473,258,272]" fontSize="6" pageId="14" pageNumber="545">th</superScript>
pleural and the pygal.
</paragraph>
</subSubSection>
</treatment>
</document>