213 lines
22 KiB
XML
213 lines
22 KiB
XML
<document id="06F9172859D165427770BBD02D055AC6" ID-DOI="10.11646/zootaxa.5448.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="11232285" ID-ZooBank="B82C87CC-501B-4844-ACAB-21BD1377E2A3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716280072248" checkinUser="plazi" docAuthor="Kehlmaier, Christian, Kuchling, Gerald, Shea, Glenn, Georges, Arthur, Zhang, Xiuwen, Fritz, Uwe & Thomson, Scott" docDate="2024" docId="0A7087BBFFAAFFBB41897BEEFE860741" docLanguage="en" docName="zootaxa.5448.4.6.pdf" docOrigin="Zootaxa 5448 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5448.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chelodina intergularis Fry 1915" docType="treatment" docVersion="2" lastPageNumber="542" masterDocId="F649FFC3FFA1FFB0411E795CFF950041" masterDocTitle="History and historical DNA: Identity of Chelodina intergularis Fry, 1915 and type localities of C. intergularis and C. rugosa Ogilby, 1890" masterLastPageNumber="555" masterPageNumber="531" pageNumber="542" updateTime="1716281573474" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="06ECDE1D2E331F1589F5B2BF43F29DF5">History and historical DNA: Identity of Chelodina intergularis Fry, 1915 and type localities of C. intergularis and C. rugosa Ogilby, 1890</mods:title>
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<mods:namePart id="FE69EE1D6D276052378E6A192116DEA7">Kehlmaier, Christian</mods:namePart>
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<mods:affiliation id="E05F41688F207E9A912C77192944765F">Museum of Zoology, Senckenberg Dresden, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="4B8069F120D955935046B58677F826DF">Kuchling, Gerald</mods:namePart>
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<mods:affiliation id="91879FF5280C97C4E1B9D32A45512FCB">School of Biological Sciences, The University of Western Australia, Crawley, WA 6009, Australia</mods:affiliation>
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</mods:name>
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<mods:name id="B854C59B64FFC575C436FDEAF305C487" type="personal">
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<mods:namePart id="1968FE7D6AFFB8632A5A561182D3B4F8">Shea, Glenn</mods:namePart>
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<mods:affiliation id="B2044637F4A15C3C032BD529A4A3EEAF">Sydney School of Veterinary Science, B 01, University of Sydney, NSW 2006, Australia & Australian Museum Research Institute, Australian Museum, 1 William Street, Sydney, NSW 2010, Australia</mods:affiliation>
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</mods:name>
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<mods:name id="DA9FA491453C0A2F4DD9DCD5AA0B6DE1" type="personal">
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<mods:namePart id="7E78F85F9B2240D17650D38466F10750">Georges, Arthur</mods:namePart>
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<mods:affiliation id="19A5A2C8D1C2102BF09DDD9A261D2062">Institute for Applied Ecology, University of Canberra, Canberra, ACT 2601, Australia</mods:affiliation>
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</mods:name>
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<mods:namePart id="091731C75A4ADC5960A74A1F87599C9E">Zhang, Xiuwen</mods:namePart>
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<mods:affiliation id="18A2A6BD7D9919E8FE9540E52DDDCE29">Institute for Applied Ecology, University of Canberra, Canberra, ACT 2601, Australia</mods:affiliation>
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</mods:name>
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<mods:namePart id="E0230F1C9FBD05E44E68CEC428828296">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="74283DC469614E8DD42DBD0A1E4DD458">Museum of Zoology, Senckenberg Dresden, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="7810BFC2020D94281BE04FF9674DD2B3">Thomson, Scott</mods:namePart>
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<mods:affiliation id="14E098002851C6EB5D48E30867FAF7AD">Centro de Estudos dos Quelônios da Amazônia, Manaus 69055 - 010, Brazil & Research Institute for the Environment and Livelihoods, Charles Darwin University, Darwin, NT 0909, Australia</mods:affiliation>
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<mods:typeOfResource id="A492E71BA00A5D5E86592BBBA9A7E053">text</mods:typeOfResource>
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<mods:title id="436D01D1D10396F7CC5CE8690500743D">Zootaxa</mods:title>
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<mods:part id="4048A9DB44629CB3FAD4A1C791FE2B5A">
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<mods:date id="AF8C8D1E129DE215F3307F8CA97BA344">2024</mods:date>
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<mods:number id="F9079EBA6890D23399535A1D880F9D7C">2024-05-14</mods:number>
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<mods:number id="E90F185946E1ABD4DD58D6B2B6189A64">5448</mods:number>
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<mods:url id="EB4D9277FF4F2DFDFC39293391714A20">http://dx.doi.org/10.11646/zootaxa.5448.4.6</mods:url>
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<mods:classification id="6CC418FFEA4C0FD508D450E2DFF3B717">journal article</mods:classification>
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<mods:identifier id="E8FD56D10C000DDA9FB2229D7B1F4D77" type="DOI">10.11646/zootaxa.5448.4.6</mods:identifier>
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<mods:identifier id="E0823B20ADB8E5939C49654B4E8464A8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="67EC930CAB96E5E1E1429828640D9249" type="Zenodo-Dep">11232285</mods:identifier>
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<mods:identifier id="61A9B76F2DFC3417D762F960F579FF12" type="ZooBank">B82C87CC-501B-4844-ACAB-21BD1377E2A3</mods:identifier>
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</mods:mods>
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<treatment id="0A7087BBFFAAFFBB41897BEEFE860741" LSID="urn:lsid:plazi:treatment:0A7087BBFFAAFFBB41897BEEFE860741" httpUri="http://treatment.plazi.org/id/0A7087BBFFAAFFBB41897BEEFE860741" lastPageNumber="542" pageId="11" pageNumber="542">
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<subSubSection id="CAC36526FFAAFFBB41897BEEFDB2028D" box="[151,551,690,717]" pageId="11" pageNumber="542" type="nomenclature">
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<paragraph id="826636ADFFAAFFBB41897BEEFDB2028D" blockId="11.[151,551,690,717]" box="[151,551,690,717]" pageId="11" pageNumber="542">
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<heading id="D92E81C1FFAAFFBB41897BEEFDB2028D" bold="true" box="[151,551,690,717]" fontSize="11" level="1" pageId="11" pageNumber="542" reason="1">
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<taxonomicName id="45D94D2EFFAAFFBB41897BEEFDB2028D" ID-CoL="87WDP" authority="Fry, 1915" authorityName="Fry" authorityYear="1915" box="[151,551,690,717]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="intergularis">
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<emphasis id="B0ADEABFFFAAFFBB41897BEEFDB2028D" bold="true" box="[151,551,690,717]" pageId="11" pageNumber="542">
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<emphasis id="B0ADEABFFFAAFFBB41897BEEFE3F028C" bold="true" box="[151,426,690,717]" italics="true" pageId="11" pageNumber="542">Chelodina intergularis</emphasis>
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<bibRefCitation id="E6484B5CFFAAFFBB40AF7BEFFDB2028D" author="Fry, D. B." box="[433,551,690,717]" pageId="11" pageNumber="542" pagination="88 - 90" refId="ref15120" refString="Fry, D. B. (1915) On a new Chelodina from Australia, with a key to the genus. Proceedings of the Royal Society of Queensland, 27 (1), 88 - 90, pl. 4." type="journal article" year="1915">Fry, 1915</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CAC36526FFAAFFBB41897BABFC9A034E" box="[151,783,759,783]" pageId="11" pageNumber="542" type="materials_examined">
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<paragraph id="826636ADFFAAFFBB41897BABFC9A034E" blockId="11.[151,783,759,783]" box="[151,783,759,783]" pageId="11" pageNumber="542">
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<materialsCitation id="32B13CF0FFAAFFBB41897BABFC9A034E" box="[151,783,759,783]" collectionCode="AM" pageId="11" pageNumber="542" specimenCode="R6255" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5D62880FFFAAFFBB41897BABFF6C034E" box="[151,249,759,783]" pageId="11" pageNumber="542" type="holotype">Holotype</typeStatus>
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: Australian Museum, Sydney (
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<collectionCode id="E4C8AE68FFAAFFBB432E7BABFDCF034E" box="[560,602,759,783]" country="Australia" name="Australian Museum" pageId="11" pageNumber="542" type="Museum">AM</collectionCode>
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<specimenCode id="D27F9ED6FFAAFFBB437F7BABFD32034E" box="[609,679,759,783]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="542">R6255</specimenCode>
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),
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<figureCitation id="1AE22A28FFAAFFBB43A77BABFC9A034E" box="[697,783,759,783]" captionStart="FIGURE 1" captionStartId="5.[151,250,1152,1176]" captionTargetBox="[175,1412,297,1106]" captionTargetId="figure-63@5.[151,1436,273,1128]" captionTargetPageId="5" captionText="FIGURE 1. Holotype of Chelodina intergularis, Australian Museum, Sydney (AM R6255). Photos: G. Shea." figureDoi="http://doi.org/10.5281/zenodo.11232287" httpUri="https://zenodo.org/record/11232287/files/figure.png" pageId="11" pageNumber="542">Figure 1</figureCitation>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="CAC36526FFAAFFBB41897A61FE860741" pageId="11" pageNumber="542" type="description">
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<paragraph id="826636ADFFAAFFBB41897A61FB1A0385" blockId="11.[151,1437,829,1792]" pageId="11" pageNumber="542">
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<emphasis id="B0ADEABFFFAAFFBB41897A61FEDB0316" bold="true" box="[151,334,829,855]" pageId="11" pageNumber="542">Measurements.</emphasis>
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Straight carapacial length (midline)
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<quantity id="45219B48FFAAFFBB43F37A61FCF70319" box="[749,866,829,856]" metricMagnitude="-1" metricUnit="m" metricValue="1.933" pageId="11" pageNumber="542" unit="mm" value="193.3">193.3 mm</quantity>
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; maximum carapacial width
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<quantity id="45219B48FFAAFFBB45B77A61FA8A0319" box="[1193,1311,829,856]" metricMagnitude="-1" metricUnit="m" metricValue="1.364" pageId="11" pageNumber="542" unit="mm" value="136.4">136.4 mm</quantity>
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; carapacial width at mid-bridge level
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<quantity id="45219B48FFAAFFBB40A47A3DFDBA033D" box="[442,559,865,892]" metricMagnitude="-1" metricUnit="m" metricValue="1.301" pageId="11" pageNumber="542" unit="mm" value="130.1">130.1 mm</quantity>
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; maximum plastral length (from level of paired caudal tips to extreme anterior margin)
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<quantity id="45219B48FFAAFFBB41E47AD9FEE603E1" box="[250,371,901,928]" metricMagnitude="-1" metricUnit="m" metricValue="1.575" pageId="11" pageNumber="542" unit="mm" value="157.5">157.5 mm</quantity>
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; midline plastral length
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<quantity id="45219B48FFAAFFBB438F7AD9FC9F03E1" box="[657,778,901,928]" metricMagnitude="-1" metricUnit="m" metricValue="1.411" pageId="11" pageNumber="542" unit="mm" value="141.1">141.1 mm</quantity>
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; plastral width at anterior extremity of bridge 72.0 mm; maximum posterior plastral width
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<quantity id="45219B48FFAAFFBB43057AF5FD170385" box="[539,642,937,964]" metricMagnitude="-2" metricUnit="m" metricValue="6.88" pageId="11" pageNumber="542" unit="mm" value="68.8">68.8 mm</quantity>
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; total shell depth at middle of bridge
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<quantity id="45219B48FFAAFFBB453A7AF5FB1E0385" box="[1060,1163,937,964]" metricMagnitude="-2" metricUnit="m" metricValue="7.06" pageId="11" pageNumber="542" unit="mm" value="70.6">70.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="826636ADFFAAFFBB41D97A91FAED0412" blockId="11.[151,1437,829,1792]" pageId="11" pageNumber="542">
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<emphasis id="B0ADEABFFFAAFFBB41D97A91FECC03A6" bold="true" box="[199,345,973,999]" pageId="11" pageNumber="542">Description.</emphasis>
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Specimen consists of a complete carapace with articulated plastron; all scutes have been removed and the specimen was highly polished during preparation. No ancillary skeletal components have been retained. The thoracic vertebrae have been lost, likely deliberately removed based on the damage to the rib heads. There is no evidence this specimen was cooked for consumption and was likely collected and prepared curatorially instead.
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</paragraph>
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<paragraph id="826636ADFFAAFFBB41D97D01FB010532" blockId="11.[151,1437,829,1792]" pageId="11" pageNumber="542">
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Dorsally there is an open space previously occupied by a 4
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<superScript id="75AC9BE5FFAAFFBB426B7D01FC17042A" attach="left" box="[885,898,1117,1131]" fontSize="6" pageId="11" pageNumber="542">th</superScript>
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neural bone (at the midline junction of the 3
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<superScript id="75AC9BE5FFAAFFBB44917D01FA08042A" attach="none" box="[1423,1437,1117,1131]" fontSize="6" pageId="11" pageNumber="542">rd</superScript>
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and 4
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<superScript id="75AC9BE5FFAAFFBB41C87DDDFF7604CE" attach="left" box="[214,227,1153,1167]" fontSize="6" pageId="11" pageNumber="542">th</superScript>
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pleurals) which is four-sided and diamond-shaped (following
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<bibRefCitation id="E6484B5CFFAAFFBB42B67DDDFBC704DA" author="Pritchard, P. C. H." box="[936,1106,1153,1179]" pageId="11" pageNumber="542" pagination="625 - 686" refId="ref17596" refString="Pritchard, P. C. H. (1988) A survey of neural bone variation among recent chelonian species, with functional interpretations. Acta Zoologica Cracoviensia, 31 (26), 625 - 686." type="journal article" year="1988">Pritchard 1988</bibRefCitation>
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;
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<bibRefCitation id="E6484B5CFFAAFFBB45407DDDFA1A04DA" author="Thomson, S. & Georges, A." box="[1118,1423,1153,1179]" pageId="11" pageNumber="542" pagination="82 - 86" refId="ref18752" refString="Thomson, S. & Georges, A. (1996) Neural bones in Australian chelid turtles. Chelonian Conservation and Biology, 2 (1), 82 - 86." type="journal article" year="1996">Thomson & Georges 1996</bibRefCitation>
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); the neural itself was lost when its supporting thoracic centrum was removed. This neural shape is indicative of a breakthrough neural that occurs in occasional specimens of any chelid species that would normally not have any visible neural bones. The carapace is ovoid in shape, typical of
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<taxonomicName id="45D94D2EFFAAFFBB42617DB1FBD90546" authorityName="Ogilby" authorityYear="1890" box="[895,1100,1261,1287]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" sensu="lato" species="rugosa">
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<emphasis id="B0ADEABFFFAAFFBB42617DB1FBD90546" box="[895,1100,1261,1287]" italics="true" pageId="11" pageNumber="542">Chelodina rugosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AB9E57C4FFAAFFBB45487DB2FB5D0546" box="[1110,1224,1261,1288]" pageId="11" pageNumber="542" sensu="lato">sensu lato</taxonomicNameLabel>
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(i.e., including
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<taxonomicName id="45D94D2EFFAAFFBB449D7DB2FED0056A" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="kurrichalpongo">
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<emphasis id="B0ADEABFFFAAFFBB449D7DB2FED0056A" italics="true" pageId="11" pageNumber="542">C. kurrichalpongo</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="45D94D2EFFAAFFBB40677C4EFD8C056A" authorityName="Werner" authorityYear="1901" box="[377,537,1297,1324]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="siebenrocki">
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<emphasis id="B0ADEABFFFAAFFBB40677C4EFD8C056A" box="[377,537,1297,1324]" italics="true" pageId="11" pageNumber="542">C. siebenrocki</emphasis>
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</taxonomicName>
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). There is a reticulated pattern on the surface of the shell bone. Pleurals 1 through 5 have lateral fontanelles at the rib/gomphosis indicating this is not a fully grown individual. Based on size, growth and generalized shape, it is in all likelihood a subadult female. However, this is not certain.
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</paragraph>
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<paragraph id="826636ADFFAAFFBB41D97C21FB3B059E" blockId="11.[151,1437,829,1792]" pageId="11" pageNumber="542">
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The plastron is also complete and possesses a reticulated surface patterning resembling that of the carapace. This is a feature of
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<taxonomicName id="45D94D2EFFAAFFBB406D7CFDFDDD05FA" box="[371,584,1441,1467]" class="Testudines" family="Chelidae" genus="Chelydera" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B0ADEABFFFAAFFBB406D7CFDFE7D05FA" box="[371,488,1441,1467]" italics="true" pageId="11" pageNumber="542">Chelydera</emphasis>
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species.
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</taxonomicName>
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Unusually, the intergular scute is extended forwards and reaches the rim of the plastron. At the same time there is a slight indentation of the anterior edge of the plastron.
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</paragraph>
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<paragraph id="826636ADFFAAFFBB41D97CB5FE860741" blockId="11.[151,1437,829,1792]" pageId="11" pageNumber="542">
|
|
Inside the shell, there is significant damage to many rib heads; all the thoracic centra are missing. There is also a small hole drilled into the anterior vertebral arches where the bone sample for our analysis was taken. The pelvic suture insertion is square to oval in general shape and is significantly extending onto the 7
|
|
<superScript id="75AC9BE5FFAAFFBB45687F6DFB16067E" attach="left" box="[1142,1155,1585,1599]" fontSize="6" pageId="11" pageNumber="542">th</superScript>
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pleural and pygal as well as completely crossing the 8
|
|
<superScript id="75AC9BE5FFAAFFBB40CD7F09FE750622" attach="left" box="[467,480,1621,1635]" fontSize="6" pageId="11" pageNumber="542">th</superScript>
|
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pleural. This is the typical condition in
|
|
<taxonomicName id="45D94D2EFFAAFFBB42827F0AFB9F062E" authorityName="Ogilby" authorityYear="1890" box="[924,1034,1622,1648]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="species" species="rugosa">
|
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<emphasis id="B0ADEABFFFAAFFBB42827F0AFB9F062E" box="[924,1034,1622,1648]" italics="true" pageId="11" pageNumber="542">C. rugosa</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="AB9E57C4FFAAFFBB450E7F0AFB17062E" box="[1040,1154,1621,1648]" pageId="11" pageNumber="542" sensu="lato">sensu lato</taxonomicNameLabel>
|
|
. The anterior bridge strut (for terminology, see
|
|
<bibRefCitation id="E6484B5CFFAAFFBB40957F25FDED06D2" author="Thomson, S. & White, A. & Georges, A." box="[395,632,1657,1684]" pageId="11" pageNumber="542" pagination="327 - 336" refId="ref18838" refString="Thomson, S., White, A. & Georges, A. (1997) Re-evaluation of Emydura lavarackorum: Identification of a living fossil. Memoirs of the Queensland Museum, 42 (1), 327 - 336." type="journal article" year="1997">
|
|
Thomson
|
|
<emphasis id="B0ADEABFFFAAFFBB40E37F26FDA506D2" box="[509,560,1657,1683]" italics="true" pageId="11" pageNumber="542">et al</emphasis>
|
|
. 1997
|
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</bibRefCitation>
|
|
) is enlarged and robust, angled back at approximately 45° and reaches the halfway point of the 1
|
|
<superScript id="75AC9BE5FFAAFFBB40DE7FC1FE5E06EA" attach="left" box="[448,459,1693,1707]" fontSize="6" pageId="11" pageNumber="542">st</superScript>
|
|
rib within the 1
|
|
<superScript id="75AC9BE5FFAAFFBB439A7FC1FD1A06EA" attach="left" box="[644,655,1693,1707]" fontSize="6" pageId="11" pageNumber="542">st</superScript>
|
|
pleural. This is the typical condition of
|
|
<taxonomicName id="45D94D2EFFAAFFBB45427FC1FABF06F6" box="[1116,1322,1693,1719]" pageId="11" pageNumber="542">
|
|
<emphasis id="B0ADEABFFFAAFFBB45427FC1FB4406F6" box="[1116,1233,1693,1719]" italics="true" pageId="11" pageNumber="542">Chelydera</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
(compare Thomson 2000;
|
|
<bibRefCitation id="E6484B5CFFAAFFBB40517F9DFDAC069A" author="Thomson, S. & Kennett, R. & Georges, A." box="[335,569,1729,1756]" pageId="11" pageNumber="542" pagination="675 - 685" refId="ref18785" refString="Thomson, S., Kennett, R. & Georges, A. (2000) A new species of long-necked turtle (Testudines: Chelidae) from the Arnhem Land Plateau, Northern Territory, Australia. Chelonian Conservation and Biology, 3 (4), 675 - 685." type="journal article" year="2000">
|
|
Thomson
|
|
<emphasis id="B0ADEABFFFAAFFBB40DE7F9EFE64069A" box="[448,497,1729,1755]" italics="true" pageId="11" pageNumber="542">et al</emphasis>
|
|
. 2000
|
|
</bibRefCitation>
|
|
) and excludes allocation of the
|
|
<typeStatus id="5D62880FFFAAFFBB42807F9DFB95069A" box="[926,1024,1729,1755]" pageId="11" pageNumber="542" type="holotype">holotype</typeStatus>
|
|
to the subgenera
|
|
<taxonomicName id="45D94D2EFFAAFFBB45D87F9DFAEB069A" authorityName="Wells & Wellington" authorityYear="1985" box="[1222,1406,1729,1755]" class="Reptilia" family="Chelidae" genus="Macrochelodina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="542" phylum="Chordata" rank="genus">
|
|
<emphasis id="B0ADEABFFFAAFFBB45D87F9DFAEB069A" box="[1222,1406,1729,1755]" italics="true" pageId="11" pageNumber="542">Macrochelodina</emphasis>
|
|
</taxonomicName>
|
|
or
|
|
<taxonomicName id="45D94D2EFFAAFFBB41897FB9FE9906BE" authorityName="Fitzinger" authorityYear="1826" box="[151,268,1765,1791]" class="Testudines" family="Chelidae" genus="Chelodina" kingdom="Animalia" order="Pleurodira" pageId="11" pageNumber="542" phylum="Chordata" rank="genus">
|
|
<emphasis id="B0ADEABFFFAAFFBB41897FB9FE9906BE" box="[151,268,1765,1791]" italics="true" pageId="11" pageNumber="542">Chelodina</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |