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<document id="9B9E64EC5F67E247B9D3E54CE8209003" ID-CLB-Dataset="3153" ID-DOI="10.5169/seals-787071" ID-GBIF-Dataset="a3876c73-615d-4cd3-88d7-779d2175638f" ID-ISSN="2624-9170" ID-Zenodo-Dep="5749772" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637933697018" checkinUser="carolina" docAuthor="Casale, Achille" docDate="2011" docId="03D34424C84CBE36C4925757FC25CDFD" docLanguage="en" docName="ContribNatHist.1.1-16.pdf.imf" docOrigin="Contributions to Natural History 16" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Duvalius (Biharotrechus) huberi Casale 2011, sp. nov." docType="treatment" docVersion="6" lastPageNumber="8" masterDocId="FFEA3C5CC84DBE31C4235232FFCACE40" masterDocTitle="Two new subterranean, microphthalmous trechine beetles from the Mediterranean area, and a synonymic note (Coleoptera: Carabidae, Trechini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="2" updateTime="1699470421717" updateUser="ExternalLinkService">
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<mods:title id="E8B9736504E8F6E087FDE7F626FE0D15">Two new subterranean, microphthalmous trechine beetles from the Mediterranean area, and a synonymic note (Coleoptera: Carabidae, Trechini)</mods:title>
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<mods:name id="8F0C8EE5DA30FEABF5CDCBFFDE73B60B" type="personal">
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</mods:role>
<mods:namePart id="5D794F0554A24514EA7AB8FAFCDCA7B3">Casale, Achille</mods:namePart>
<mods:affiliation id="2E3708BB5A3C8FBE06A4CBF95ED8A903">Università di Sassari Dipartimento di Zoologia e Genetica Evoluzionistica Via Muroni 25 I 07100 Sassari (Italy)</mods:affiliation>
<mods:nameIdentifier id="2699F54806C9C8EAA877E09DF21B57D3" type="email">casale@uniss.it &amp; a_casale@libero.it</mods:nameIdentifier>
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<mods:title id="381CA47C579FDD803FBDE82D3210C544">Contributions to Natural History</mods:title>
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<mods:part id="1DBFF5E7951C1BDEA7F74D050A674638">
<mods:date id="6ECFDF7B36471C994D7EB85E917EEA63">2011</mods:date>
<mods:detail id="788889F7D4C40A1E013F106D7B168C7E" type="pubDate">
<mods:number id="5A3914D96B1195FB81EB56D21D975919">2011-03-17</mods:number>
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<mods:identifier id="46878A61A4E5E74EA31CA791566E12B4" type="CLB-Dataset">3153</mods:identifier>
<mods:identifier id="D471BFA828E6011B4FD16208AD3D6A7C" type="DOI">10.5169/seals-787071</mods:identifier>
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<mods:identifier id="EAE1B2F4A917750909A44AE57AB2A1EE" type="Zenodo-Dep">5749772</mods:identifier>
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<treatment id="03D34424C84CBE36C4925757FC25CDFD" ID-DOI="http://doi.org/10.5281/zenodo.5749776" ID-GBIF-Taxon="190875632" ID-Zenodo-Dep="5749776" LSID="urn:lsid:plazi:treatment:03D34424C84CBE36C4925757FC25CDFD" httpUri="http://treatment.plazi.org/id/03D34424C84CBE36C4925757FC25CDFD" lastPageId="7" lastPageNumber="8" pageId="1" pageNumber="2">
<subSubSection id="C360A6B9C84CBE30C4925757FDB1CB3E" box="[177,635,1381,1406]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BC5F532C84CBE30C4925757FDB1CB3E" blockId="1.[177,635,1381,1406]" box="[177,635,1381,1406]" pageId="1" pageNumber="2">
<heading id="D08D425EC84CBE30C4925757FDB1CB3E" box="[177,635,1381,1406]" centered="true" fontSize="36" level="2" pageId="1" pageNumber="2" reason="4">
<emphasis id="B90E2920C84CBE30C4925757FDB1CB3E" bold="true" box="[177,635,1381,1406]" pageId="1" pageNumber="2">
<taxonomicName id="4C7A8EB1C84CBE30C4925757FDD4CB3E" ID-CoL="384Z2" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[177,542,1381,1406]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C84CBE30C4925757FEDECB3E" bold="true" box="[177,276,1381,1406]" italics="true" pageId="1" pageNumber="2">Duvalius</emphasis>
(
<emphasis id="B90E2920C84CBE30C5075757FE0FCB3E" bold="true" box="[292,453,1381,1406]" italics="true" pageId="1" pageNumber="2">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C84CBE30C5F75757FDD4CB3E" bold="true" box="[468,542,1381,1406]" italics="true" pageId="1" pageNumber="2">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC84CBE30C6075757FDB1CB3E" box="[548,635,1381,1406]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C84CBE30C4935782FD34CB88" box="[176,766,1456,1480]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BC5F532C84CBE30C4935782FD34CB88" blockId="1.[176,766,1456,1480]" box="[176,766,1456,1480]" pageId="1" pageNumber="2">
Type locality:
<collectingCountry id="F36DB5A2C84CBE30C56F5782FE50CB88" box="[332,410,1456,1480]" name="Turkey" pageId="1" pageNumber="2">Turkey</collectingCountry>
, Southern
<collectingCountry id="F36DB5A2C84CBE30C6305782FDBCCB88" box="[531,630,1456,1480]" name="Turkey" pageId="1" pageNumber="2">Anatolia</collectingCountry>
: Geyik DağI.
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C84CBE30C49357C8FE9CC852" box="[176,342,1530,1554]" pageId="1" pageNumber="2" type="multiple">
<paragraph id="8BC5F532C84CBE30C49357C8FE9CC852" blockId="1.[176,1067,1530,1629]" box="[176,342,1530,1554]" pageId="1" pageNumber="2">Type material:</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C84CBE30C4915412FD74C81D" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BC5F532C84CBE30C4915412FD74C81D" blockId="1.[176,1067,1530,1629]" pageId="1" pageNumber="2">
<materialsCitation id="3B12FF6FC84CBE30C4915412FD74C81D" ID-GBIF-Occurrence="3413090301" collectingDate="22. 5. 2007" collectionCode="NMBE" collectorName="C. Huber" country="Turkey" elevation="0" latitude="36.9" location="Geyik DagI" longLatPrecision="925" longitude="32.189583" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Antalya" typeStatus="holotype">
<typeStatus id="54C14B90C84CBE30C4915412FED0C878" box="[178,282,1568,1592]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7C3EBBC84CBE30C502542FFEFFC878" box="[289,309,1565,1592]" pageId="1" pageNumber="2" type="male"></specimenCount>
with the following data: &quot;TR
<collectingRegion id="49BE3BD0C84CBE30C6BE5412FD39C878" box="[669,755,1568,1592]" country="Turkey" name="Antalya" pageId="1" pageNumber="2">Antalya</collectingRegion>
<location id="8EA5A3E9C84CBE30C6D85412FCBFC878" LSID="urn:lsid:plazi:treatment:03D34424C84CBE36C4925757FC25CDFD:8EA5A3E9C84CBE30C6D85412FCBFC878" box="[763,885,1568,1592]" latitude="36.9" longLatPrecision="925" longitude="32.189583" name="Geyik DagI" pageId="1" pageNumber="2" stateProvince="Antalya">Geyik DağI</location>
<elevation id="00571201C84CBE30C75E5412FC2BC878" box="[893,993,1568,1592]" pageId="1" pageNumber="2">2600 mt</elevation>
,
<geoCoordinate id="EE4E93F5C84CBE30C7CF5412FF19C81D" degrees="36" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="36.9">N 36° 54'</geoCoordinate>
<geoCoordinate id="EE4E93F5C84CBE30C4FA5477FEF1C81D" box="[217,315,1605,1629]" pageId="1" pageNumber="2">E 32° 11'</geoCoordinate>
<collectingDate id="EF802A1AC84CBE30C5625477FE7EC81D" box="[321,436,1605,1629]" pageId="1" pageNumber="2">
<date id="FFC4D3F2C84CBE30C5625477FEB9C81D" box="[321,371,1605,1629]" pageId="1" pageNumber="2" value="2007-05-22">22.5</date>
.2007
</collectingDate>
leg.
<collectorName id="268F90E4C84CBE30C5CA5477FD9BC81D" box="[489,593,1605,1629]" pageId="1" pageNumber="2">C. Huber</collectorName>
&quot; (
<collectionCode id="ED6B6DF7C84CBE30C6475477FD7EC81D" box="[612,692,1605,1629]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="1" pageNumber="2">NMBE</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF05A5BAC84FBE33C4F4547BFC00C81B" ID-DOI="http://doi.org/10.5281/zenodo.5749774" ID-Zenodo-Dep="5749774" box="[215,970,1609,1627]" httpUri="https://zenodo.org/record/5749774/files/figure.png" pageId="2" pageNumber="3" startId="2.[215,244,1609,1627]" targetBox="[237,1077,148,1505]" targetPageId="2">
<paragraph id="8BC5F532C84FBE33C4F4547BFC00C81B" blockId="2.[215,970,1609,1627]" box="[215,970,1609,1627]" pageId="2" pageNumber="3">
Fig. 1:
<taxonomicName id="4C7A8EB1C84FBE33C532547BFDEDC81B" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[273,551,1609,1627]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C84FBE33C532547BFE97C81B" bold="true" box="[273,349,1609,1627]" italics="true" pageId="2" pageNumber="3">Duvalius</emphasis>
(
<emphasis id="B90E2920C84FBE33C54A547BFE2EC81B" bold="true" box="[361,484,1609,1627]" italics="true" pageId="2" pageNumber="3">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C84FBE33C5CC547BFDEDC81B" bold="true" box="[495,551,1609,1627]" italics="true" pageId="2" pageNumber="3">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC84FBE33C60F547BFDB8C81B" box="[556,626,1609,1627]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype, habitus, dorsal aspect.
</paragraph>
</caption>
<subSubSection id="C360A6B9C84EBE32C49152A7FE5BCEEE" box="[178,401,149,174]" pageId="3" pageNumber="4" type="multiple">
<paragraph id="8BC5F532C84EBE32C49152A7FE5BCEEE" blockId="3.[178,401,149,174]" box="[178,401,149,174]" pageId="3" pageNumber="4">
<emphasis id="B90E2920C84EBE32C49152A7FE5BCEEE" bold="true" box="[178,401,149,174]" pageId="3" pageNumber="4">Diagnostic features</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C84EBE32C49252D2FBEECFBD" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BC5F532C84EBE32C49252D2FDD6CFCD" blockId="3.[175,1067,224,621]" pageId="3" pageNumber="4">
A medium sized (TL:
<quantity id="4C8258D7C84EBE32C58D52D2FDD7CEB8" box="[430,541,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="4" unit="mm" value="5.5">5.50 mm</quantity>
), glabrous, depigmented, microphthalmous trechine species (
<figureCitation id="1341E9B7C84EBE32C55B5337FE7DCF5D" box="[376,439,261,285]" captionStart="Fig" captionStartId="2.[215,244,1609,1627]" captionTargetBox="[237,1077,148,1505]" captionTargetId="figure-11@2.[232,1084,148,1496]" captionTargetPageId="2" captionText="Fig. 1: Duvalius (Biharotrechus) huberi sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749774" httpUri="https://zenodo.org/record/5749774/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
), with the character states of the genus
<taxonomicName id="4C7A8EB1C84EBE32C7A25337FC2ECF5D" authorityName="Delarouzee" authorityYear="1859" box="[897,996,261,285]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84EBE32C7A25337FC2ECF5D" box="[897,996,261,285]" italics="true" pageId="3" pageNumber="4">Duvalius</emphasis>
</taxonomicName>
DELAROUzéE, 1859 of Subgenus
<taxonomicName id="4C7A8EB1C84EBE32C5D25318FCF8CF02" authority="BOKOR, 1922" authorityName="BOKOR" authorityYear="1922" box="[497,818,298,322]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Biharotrechus">
<emphasis id="B90E2920C84EBE32C5D25318FD58CF02" box="[497,658,298,322]" italics="true" pageId="3" pageNumber="4">Biharotrechus</emphasis>
BOKOR, 1922
</taxonomicName>
(
<taxonomicName id="4C7A8EB1C84EBE32C7665318FF23CF28" authority="JEANNEL, 1928" authorityName="JEANNEL" authorityYear="1928" class="Insecta" family="Carabidae" genus="Duvaliotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84EBE32C7665318FC74CF02" box="[837,958,298,322]" italics="true" pageId="3" pageNumber="4">Duvaliotes</emphasis>
<bibRefCitation id="EFEB88C3C84EBE32C7E45318FF23CF28" author="Jeannel, R." pageId="3" pageNumber="4" pagination="1 - 808" refId="ref4875" refString="Jeannel, R. (1928): Monographie des Trechinae 3. Morphologie comparee et distribution geographique d'un groupe de Coleopteres - L'Abeille, Paris 35: 1 - 808." type="journal article" year="1928">JEANNEL, 1928</bibRefCitation>
</taxonomicName>
, in the widest sense of
<bibRefCitation id="EFEB88C3C84EBE32C6255362FD6CCF28" author="Jeannel, R." box="[518,678,336,360]" pageId="3" pageNumber="4" pagination="1 - 808" refId="ref4875" refString="Jeannel, R. (1928): Monographie des Trechinae 3. Morphologie comparee et distribution geographique d'un groupe de Coleopteres - L'Abeille, Paris 35: 1 - 808." type="journal article" year="1928">Jeannel, 1928</bibRefCitation>
),
<bibRefCitation id="EFEB88C3C84EBE32C6E35362FC21CF28" author="Casale, A. &amp; Laneyrie, R." box="[704,1003,336,360]" pageId="3" pageNumber="4" refId="ref4679" refString="Casale, A. &amp; Laneyrie, R. (1982): Trechodinae et Trechinae du monde. Tableau des sousfamilles, tribus, series phyletiques, genres, et catalogue general des especes - Memoires de Biospeologie 9: 226 pp." type="book" year="1982">Casale &amp; Laneyrie (1982)</bibRefCitation>
, and
<bibRefCitation id="EFEB88C3C84EBE32C4925347FDDDCFCD" author="Casale, A. &amp; Vigna Taglianti, A." box="[177,535,373,397]" pageId="3" pageNumber="4" pagination="309 - 329" refId="ref4723" refString="Casale, A. &amp; Vigna Taglianti, A. (1984): Due nuovi Carabidi cavernicoli dell'Anatolia meridionale (Coleoptera, Carabidae) - Fragmenta entomologica (Roma) 17: 309 - 329." type="journal article" year="1984">Casale &amp; Vigna Taglianti (1984)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC5F532C84EBE32C4F453A8FBEECFBD" blockId="3.[175,1067,224,621]" pageId="3" pageNumber="4">
Colour dark reddish. Pronotum cordiform, widened in front, with lateral sides constricted to the base. Elytra elongate, depressed, with humeral angles rounded but evident. Chaetotaxy as in
<figureCitation id="1341E9B7C84EBE32C69453D7FD39CFBD" box="[695,755,485,509]" captionStart="Fig" captionStartId="2.[215,244,1609,1627]" captionTargetBox="[237,1077,148,1505]" captionTargetId="figure-11@2.[232,1084,148,1496]" captionTargetPageId="2" captionText="Fig. 1: Duvalius (Biharotrechus) huberi sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749774" httpUri="https://zenodo.org/record/5749774/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
. Male genitalia as in
<figureCitation id="1341E9B7C84EBE32C7C353D7FBD7CFBD" box="[992,1053,485,509]" captionStart="Fig" captionStartId="4.[215,243,154,172]" captionTargetBox="[545,1051,149,914]" captionTargetId="figure-213@4.[541,1053,144,916]" captionTargetPageId="4" captionText="Fig. 2: Duvalius (Biharo- trechus) huberi sp. nov., male holotype, aedeagus in left lateral aspect (a), copulatory piece in dor- sal aspect (b)." figureDoi="http://doi.org/10.5281/zenodo.5749778" httpUri="https://zenodo.org/record/5749778/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C84EBE32C4F45038FEE6CC2D" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BC5F532C84EBE32C4F45038FEE6CC2D" blockId="3.[175,1067,224,621]" pageId="3" pageNumber="4">
Close to
<taxonomicName id="4C7A8EB1C84EBE32C51A5038FC2BCC62" authority="CASALE &amp; VIGNA TAGLIANTI, 1984" authorityName="CASALE &amp; VIGNA TAGLIANTI" authorityYear="1984" box="[313,993,522,546]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bortesii" subGenus="Biharotrechus">
<emphasis id="B90E2920C84EBE32C51A5038FE83CC62" box="[313,329,522,546]" italics="true" pageId="3" pageNumber="4">D</emphasis>
. (
<emphasis id="B90E2920C84EBE32C5435038FDCACC62" box="[352,512,522,546]" italics="true" pageId="3" pageNumber="4">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C84EBE32C6335038FDA2CC62" box="[528,616,522,546]" italics="true" pageId="3" pageNumber="4">bortesii</emphasis>
<bibRefCitation id="EFEB88C3C84EBE32C64C5038FC2BCC62" author="Casale, A. &amp; Vigna Taglianti, A." box="[623,993,522,546]" pageId="3" pageNumber="4" pagination="309 - 329" refId="ref4723" refString="Casale, A. &amp; Vigna Taglianti, A. (1984): Due nuovi Carabidi cavernicoli dell'Anatolia meridionale (Coleoptera, Carabidae) - Fragmenta entomologica (Roma) 17: 309 - 329." type="journal article" year="1984">CASALE &amp; VIGNA TAGLIANTI, 1984</bibRefCitation>
</taxonomicName>
, from which is distinguished by several different morphological features (see: Relationships).
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C84EBE35C49150F7FDFFC81D" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="8BC5F532C84EBE32C49150F7FEFFCC9E" blockId="3.[178,309,709,734]" box="[178,309,709,734]" pageId="3" pageNumber="4">
<emphasis id="B90E2920C84EBE32C49150F7FEFFCC9E" bold="true" box="[178,309,709,734]" pageId="3" pageNumber="4">Description</emphasis>
</paragraph>
<paragraph id="8BC5F532C84EBE32C4925122FC40CD0D" blockId="3.[177,1067,784,1629]" pageId="3" pageNumber="4">
General features as in
<figureCitation id="1341E9B7C84EBE32C5955122FE3ECD68" box="[438,500,784,808]" captionStart="Fig" captionStartId="2.[215,244,1609,1627]" captionTargetBox="[237,1077,148,1505]" captionTargetId="figure-11@2.[232,1084,148,1496]" captionTargetPageId="2" captionText="Fig. 1: Duvalius (Biharotrechus) huberi sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749774" httpUri="https://zenodo.org/record/5749774/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
. Medium sized: TL:
<quantity id="4C8258D7C84EBE32C6FD5122FC88CD68" box="[734,834,784,808]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="4" unit="mm" value="5.5">5.50 mm</quantity>
; L: 6.0 mm. Colour dark reddish, with paler yellow reddish palpi, antennae and legs.
</paragraph>
<paragraph id="8BC5F532C84EBE32C4F45168FBEACDD8" blockId="3.[177,1067,784,1629]" pageId="3" pageNumber="4">Dorsal surface moderately shiny; transversal cuticular microlines evident on frons and pronotum, scarcely visible as polygonal meshes on elytral intervals.</paragraph>
<paragraph id="8BC5F532C84EBE32C4F45197FE31CAA8" blockId="3.[177,1067,784,1629]" pageId="3" pageNumber="4">Head elongate, with frontal furrows markedly impressed, angulate in the anterior third, with evident transveral wrinkles; frons convex; supraorbital setiferous punctures inserted on ranges slightly convergent backwards; eyes small, as long as 1/4 of genae, flat, oblique and whitish, with slightly distinguishable ommatidia, pigmented only at margins; genae regularly curved, convex, with deep transversal wrinkles, narrowed to the neck constriction; labrum slightly emarginate at the anterior side; mentum tooth prominent, emarginate at apex; antennae elongate, exceeding backwards the level of the fourth humeral setiferous puncture.</paragraph>
<paragraph id="8BC5F532C84EBE32C4F456C7FF38CB88" blockId="3.[177,1067,784,1629]" pageId="3" pageNumber="4">Pronotum cordiform transverse (PL/PW: 0.80), widest at the anterior third; disc moderately convex; sides markedly arcuate in front, shortly and slightly sinuate towards hind angles, which are evident but not prominent outside; marginal furrows deep; anterior margin gently arcuate, front angles effaced; base straight, slightly emarginate in the centre, fully beaded; basal foveae deep.</paragraph>
<paragraph id="8BC5F532C84EBE35C4F457E7FED5CADD" blockId="3.[177,1067,784,1629]" lastBlockId="4.[215,1105,970,1629]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Elytra elongate (EL/EW: 1.58), slightly widened at the apical third, much wider than prothorax; disc depressed; humeri rounded but evident, with prehumeral margins oblique; lateral furrows relatively wide and deep; striae all visible: striae 14 much deeper, 56 reduced to series of punctures, stria 7 almost effaced; apical striola very deep and curved, connected by some points with apex of stria 5; intervals flat; apical carina developed. Chaetotaxy as in
<figureCitation id="1341E9B7C849BE35C5345627FE9CCA6D" box="[279,342,1045,1069]" captionStart="Fig" captionStartId="2.[215,244,1609,1627]" captionTargetBox="[237,1077,148,1505]" captionTargetId="figure-11@2.[232,1084,148,1496]" captionTargetPageId="2" captionText="Fig. 1: Duvalius (Biharotrechus) huberi sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749774" httpUri="https://zenodo.org/record/5749774/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
: three setiferous discal punctures (two discal, one pre-apical) on stria 3; umbilicate pores 5 and 6 very close to each other; apical and anguloapical punctures present, forming with the pre-apical puncture an apical triangle.
</paragraph>
<caption id="DF05A5BAC849BE35C4F452A8FE9ECF71" ID-DOI="http://doi.org/10.5281/zenodo.5749778" ID-Zenodo-Dep="5749778" httpUri="https://zenodo.org/record/5749778/files/figure.png" pageId="4" pageNumber="5" startId="4.[215,243,154,172]" targetBox="[545,1051,149,914]" targetPageId="4">
<paragraph id="8BC5F532C849BE35C4F452A8FE9ECF71" blockId="4.[215,433,154,305]" pageId="4" pageNumber="5">
Fig. 2:
<taxonomicName id="4C7A8EB1C849BE35C53252A8FED4CE86" authority="(Biharotrechus)" authorityName="Delarouzee" authorityYear="1859" baseAuthorityName="Biharotrechus" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C849BE35C53252A8FE96CEEC" bold="true" box="[273,348,154,172]" italics="true" pageId="4" pageNumber="5">Duvalius</emphasis>
(
<emphasis id="B90E2920C849BE35C54452A8FED4CE86" bold="true" pageId="4" pageNumber="5">
<emphasis id="B90E2920C849BE35C54452A8FEDDCE86" bold="true" italics="true" pageId="4" pageNumber="5">Biharotrechus</emphasis>
)
</emphasis>
</taxonomicName>
<emphasis id="B90E2920C849BE35C5005286FF3BCF71" bold="true" pageId="4" pageNumber="5">
<taxonomicName id="4C7A8EB1C849BE35C5005286FE91CE86" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[291,347,180,198]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="B90E2920C849BE35C5005286FE91CE86" bold="true" box="[291,347,180,198]" italics="true" pageId="4" pageNumber="5">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC849BE35C57C5286FE6ECE86" box="[351,420,180,198]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, male
<typeStatus id="54C14B90C849BE35C52652FDFE9FCEA1" box="[261,341,207,225]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, aedeagus in left lateral aspect (a), copulatory piece in dorsal
</emphasis>
aspect (b).
</paragraph>
</caption>
<paragraph id="8BC5F532C849BE35C4DE5698FCBECB4D" blockId="4.[215,1105,970,1629]" pageId="4" pageNumber="5">Legs long, slender; protibiae slightly dilated and sparsely pubescent at apex, each with a long groove on the external side; protarsi in the male each with two basal tarsomeres dilated and denticulate inwards.</paragraph>
<paragraph id="8BC5F532C849BE35C4DE5728FCE0C878" blockId="4.[215,1105,970,1629]" pageId="4" pageNumber="5">
Male genitalia as in
<figureCitation id="1341E9B7C849BE35C5C65728FDE9CB72" box="[485,547,1306,1330]" captionStart="Fig" captionStartId="4.[215,243,154,172]" captionTargetBox="[545,1051,149,914]" captionTargetId="figure-213@4.[541,1053,144,916]" captionTargetPageId="4" captionText="Fig. 2: Duvalius (Biharo- trechus) huberi sp. nov., male holotype, aedeagus in left lateral aspect (a), copulatory piece in dor- sal aspect (b)." figureDoi="http://doi.org/10.5281/zenodo.5749778" httpUri="https://zenodo.org/record/5749778/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
. Median lobe of aedeagus relatively short (
<quantity id="4C8258D7C849BE35C03B5728FECFCB18" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="4" pageNumber="5" unit="mm" value="1.08">1.08 mm</quantity>
) and thickened, in lateral aspect regularly bent basally and slightly sinuate in the apical third, regularly narrowed at apex; basal bulb large, inflated, with wide basal orifice; sagittal carina absent. Endophallus armed with an elongate, &quot;isotopic&quot; copulatory piece, obliquely truncate at apex in lateral aspect, regularly rounded in dorsal aspect, bifid at base, and a series of small, sclerotized spines that fringe the basal tract of lateral sides of the copulatory piece. Parameres robust, each with five apical setae.
</paragraph>
<paragraph id="8BC5F532C849BE35C4DE5477FDFFC81D" blockId="4.[215,1105,970,1629]" box="[253,565,1605,1629]" pageId="4" pageNumber="5">Female genitalia: unknown.</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C848BE34C49152A7FEAFCEEE" box="[178,357,149,174]" pageId="5" pageNumber="6" type="multiple">
<paragraph id="8BC5F532C848BE34C49152A7FEAFCEEE" blockId="5.[178,357,149,174]" box="[178,357,149,174]" pageId="5" pageNumber="6">
<emphasis id="B90E2920C848BE34C49152A7FEAFCEEE" bold="true" box="[178,357,149,174]" pageId="5" pageNumber="6">Specific epithet</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C848BE34C49152D2FCACCF28" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BC5F532C848BE34C49152D2FCACCF28" blockId="5.[178,1061,224,360]" pageId="5" pageNumber="6">
I wish to dedicate this interesting new species to
<collectorName id="268F90E4C848BE34C6C152D2FC66CEB8" box="[738,940,224,248]" pageId="5" pageNumber="6">Dr. Charles Huber</collectorName>
, curator of the Entomological Department at the
<collectingRegion id="49BE3BD0C848BE34C64A5337FD66CF5D" box="[617,684,261,285]" country="Switzerland" name="Bern" pageId="5" pageNumber="6">Berne</collectingRegion>
Museum (
<collectingCountry id="F36DB5A2C848BE34C7045337FC7ECF5D" box="[807,948,261,285]" name="Switzerland" pageId="5" pageNumber="6">Switzerland</collectingCountry>
) and well known specialist of carabids of the genus
<taxonomicName id="4C7A8EB1C848BE34C6A65318FD1ACF02" box="[645,720,298,322]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C848BE34C6A65318FD1ACF02" box="[645,720,298,322]" italics="true" pageId="5" pageNumber="6">Nebria</emphasis>
</taxonomicName>
, who sampled with
<collectorName id="268F90E4C848BE34C78C5318FECBCF28" pageId="5" pageNumber="6">Dr. Werner Marggi</collectorName>
the
<typeStatus id="54C14B90C848BE34C5115362FE5DCF28" box="[306,407,336,360]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
specimen, and offered it to me for study.
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C848BE34C491538DFDAACF98" box="[178,608,447,472]" pageId="5" pageNumber="6" type="multiple">
<paragraph id="8BC5F532C848BE34C491538DFDAACF98" blockId="5.[178,608,447,472]" box="[178,608,447,472]" pageId="5" pageNumber="6">
<emphasis id="B90E2920C848BE34C491538DFDAACF98" bold="true" box="[178,608,447,472]" pageId="5" pageNumber="6">Geographical distribution and habitat</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C848BE34C4935038FCB4CCF8" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BC5F532C848BE34C4935038FCB4CCF8" blockId="5.[176,1067,522,696]" pageId="5" pageNumber="6">
The only male individual of the new species here described was sampled on the Northern slope of the Geyik DağI Mt. (Southern
<collectingCountry id="F36DB5A2C848BE34C7045002FC47CC08" box="[807,909,560,584]" name="Turkey" pageId="5" pageNumber="6">Anatolia</collectingCountry>
, Toros chain,
<collectingRegion id="49BE3BD0C848BE34C4925067FEC6CC2D" box="[177,268,597,621]" country="Turkey" name="Antalya" pageId="5" pageNumber="6">Antalya</collectingRegion>
) by Charles Huber and Werner Marggi at
<quantity id="4C8258D7C848BE34C6D45067FC9ACC2D" box="[759,848,597,621]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="6" unit="m" value="2600.0">2600 m</quantity>
of altitude, under stones near the snow (
<figureCitation id="1341E9B7C848BE34C5945048FE32CCD2" box="[439,504,634,658]" captionStart="Fig" captionStartId="6.[215,244,1535,1553]" captionTargetBox="[215,1099,147,1486]" captionTargetPageId="6" captionText="Fig. 3: Duvalius (Biharotrechus) huberi sp. nov., type locality: Geyik Daği Mt. (Southern Anatolia, Toros chain, Antalya) (a); detail of the habitat (b) (photo C. Huber)." figureDoi="http://doi.org/10.5281/zenodo.5749782" httpUri="https://zenodo.org/record/5749782/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
). As many
<taxonomicName id="4C7A8EB1C848BE34C6585048FD14CCD2" box="[635,734,634,658]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90E2920C848BE34C6585048FD14CCD2" box="[635,734,634,658]" italics="true" pageId="5" pageNumber="6">Duvalius</emphasis>
</taxonomicName>
species of the Euro-Mediterranean area, it is an upper hypogean species in the alpine zone.
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C848BE34C491513DFE85CD68" box="[178,335,783,808]" pageId="5" pageNumber="6" type="multiple">
<paragraph id="8BC5F532C848BE34C491513DFE85CD68" blockId="5.[178,335,783,808]" box="[178,335,783,808]" pageId="5" pageNumber="6">
<emphasis id="B90E2920C848BE34C491513DFE85CD68" bold="true" box="[178,335,783,808]" pageId="5" pageNumber="6">Relationships</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C360A6B9C848BE36C4925168FC25CDFD" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BC5F532C848BE34C4925168FDE4CDD8" blockId="5.[175,1067,858,1629]" pageId="5" pageNumber="6">
<taxonomicName id="4C7A8EB1C848BE34C4925168FE1ECD32" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[177,468,858,882]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C848BE34C4925168FF0BCD32" box="[177,193,858,882]" italics="true" pageId="5" pageNumber="6">D</emphasis>
. (
<emphasis id="B90E2920C848BE34C4FA5168FEB3CD32" box="[217,377,858,882]" italics="true" pageId="5" pageNumber="6">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C848BE34C5A95168FE1ECD32" box="[394,468,858,882]" italics="true" pageId="5" pageNumber="6">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC848BE34C5F85168FDF9CD32" box="[475,563,858,882]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
is particularly interesting for both taxonomical and biogeographical aspects.
</paragraph>
<paragraph id="8BC5F532C848BE34C4F45197FEAFC852" blockId="5.[175,1067,858,1629]" pageId="5" pageNumber="6">
First, from the taxonomical point of view, it is apparently the first adelphotaxon known so far of
<taxonomicName id="4C7A8EB1C848BE34C59B51F8FF23CA48" authority="CASALE &amp; VIGNA TAGLIANTI, 1984" authorityName="CASALE &amp; VIGNA TAGLIANTI" authorityYear="1984" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bortesii" subGenus="Biharotrechus">
<emphasis id="B90E2920C848BE34C59B51F8FE02CDA2" box="[440,456,970,994]" italics="true" pageId="5" pageNumber="6">D</emphasis>
. (
<emphasis id="B90E2920C848BE34C5C251F8FD4BCDA2" box="[481,641,970,994]" italics="true" pageId="5" pageNumber="6">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C848BE34C6B051F8FD21CDA2" box="[659,747,970,994]" italics="true" pageId="5" pageNumber="6">bortesii</emphasis>
<bibRefCitation id="EFEB88C3C848BE34C6D051F8FF23CA48" author="Casale, A. &amp; Vigna Taglianti, A." pageId="5" pageNumber="6" pagination="309 - 329" refId="ref4723" refString="Casale, A. &amp; Vigna Taglianti, A. (1984): Due nuovi Carabidi cavernicoli dell'Anatolia meridionale (Coleoptera, Carabidae) - Fragmenta entomologica (Roma) 17: 309 - 329." type="journal article" year="1984">CASALE &amp; VIGNA TAGLIANTI, 1984</bibRefCitation>
</taxonomicName>
geographically vicariant in a massif (Bey DağlarI Mt.,
<collectingRegion id="49BE3BD0C848BE34C74E51C2FBD5CA48" box="[877,1055,1008,1032]" country="Turkey" name="Mugla" pageId="5" pageNumber="6">Muğla province</collectingRegion>
) at 80 kms SW of Geyik DağI to which it seems related by many morphological features, and the peculiar shape of both median lobe of aedeagus and copulatory piece. From the latter,
<taxonomicName id="4C7A8EB1C848BE34C61A5652FC94CA38" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[569,862,1120,1144]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C848BE34C61A5652FD83CA38" box="[569,585,1120,1144]" italics="true" pageId="5" pageNumber="6">D</emphasis>
. (
<emphasis id="B90E2920C848BE34C6415652FCC8CA38" box="[610,770,1120,1144]" italics="true" pageId="5" pageNumber="6">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C848BE34C7375652FC94CA38" box="[788,862,1120,1144]" italics="true" pageId="5" pageNumber="6">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC848BE34C7455652FC0ACA38" box="[870,960,1120,1144]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished by the more elongate head, the wider pronotum with basal angles smaller and not prominent; the shorter, more depressed elytra, with shallower and less deeply punctuate striae; and the different shape of aedeagus: in
<taxonomicName id="4C7A8EB1C848BE34C03056E2FEE1CB4D" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C848BE34C03056E2FBE9CAA8" box="[1043,1059,1232,1256]" italics="true" pageId="5" pageNumber="6">D</emphasis>
. (
<emphasis id="B90E2920C848BE34C49956C7FF1BCB4D" box="[186,209,1269,1293]" italics="true" pageId="5" pageNumber="6">B.</emphasis>
)
<emphasis id="B90E2920C848BE34C4C256C7FEE1CB4D" box="[225,299,1269,1293]" italics="true" pageId="5" pageNumber="6">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC848BE34C51056C7FE46CB4D" box="[307,396,1269,1293]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
the median lobe is shorter and more thickened, and endophallus has reduced sclerotized spines and scales, and much more elongate copulatory piece. It should also be recalled that individuals of
<taxonomicName id="4C7A8EB1C848BE34C75E5772FBEFCB18" authorityName="CASALE &amp; VIGNA TAGLIANTI" authorityYear="1984" box="[893,1061,1344,1368]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bortesii" subGenus="Biharotrechus">
<emphasis id="B90E2920C848BE34C75E5772FC47CB18" box="[893,909,1344,1368]" italics="true" pageId="5" pageNumber="6">D</emphasis>
. (
<emphasis id="B90E2920C848BE34C7865772FC76CB18" box="[933,956,1344,1368]" italics="true" pageId="5" pageNumber="6">B.</emphasis>
)
<emphasis id="B90E2920C848BE34C7EE5772FBEFCB18" box="[973,1061,1344,1368]" italics="true" pageId="5" pageNumber="6">bortesii</emphasis>
</taxonomicName>
were sampled in a pit at
<quantity id="4C8258D7C848BE34C5E45757FDEACB3D" box="[455,544,1381,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="m" value="1200.0">1200 m</quantity>
, whereas
<taxonomicName id="4C7A8EB1C848BE34C6B35757FD32CB3D" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[656,760,1381,1405]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="B90E2920C848BE34C6B35757FD32CB3D" box="[656,760,1381,1405]" italics="true" pageId="5" pageNumber="6">D. huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC848BE34C6DD5757FC9DCB3D" box="[766,855,1381,1405]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
in the alpine zone at
<quantity id="4C8258D7C848BE34C4F357B8FEEDCBE2" box="[208,295,1418,1442]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="6" unit="m" value="2600.0">2600 m</quantity>
near the snow. But this datum is scarcely informative: in fact, species with this degree of low specialization to the subterranean environment can be found both in caves at lower altitude and in the shallow hypogean zone at high altitude.
</paragraph>
<paragraph id="8BC5F532C848BE36C4F45412FDDBCFCD" blockId="5.[175,1067,858,1629]" lastBlockId="7.[177,1067,149,957]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
Validity and limits of the Subgenus
<taxonomicName id="4C7A8EB1C848BE34C6575412FC53C878" authority="JEANNEL, 1928" authorityName="JEANNEL" authorityYear="1928" box="[628,921,1568,1592]" class="Insecta" family="Carabidae" genus="Duvaliotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C848BE34C6575412FD27C878" box="[628,749,1568,1592]" italics="true" pageId="5" pageNumber="6">Duvaliotes</emphasis>
<bibRefCitation id="EFEB88C3C848BE34C6D55412FC53C878" author="Jeannel, R." box="[758,921,1568,1592]" pageId="5" pageNumber="6" pagination="1 - 808" refId="ref4875" refString="Jeannel, R. (1928): Monographie des Trechinae 3. Morphologie comparee et distribution geographique d'un groupe de Coleopteres - L'Abeille, Paris 35: 1 - 808." type="journal article" year="1928">JEANNEL, 1928</bibRefCitation>
</taxonomicName>
, and its distinction from the Subgenus
<taxonomicName id="4C7A8EB1C848BE34C5DC5477FCE1C81D" authority="JEANNEL, 1928" authorityName="JEANNEL" authorityYear="1928" box="[511,811,1605,1629]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C848BE34C5DC5477FDB0C81D" box="[511,634,1605,1629]" italics="true" pageId="5" pageNumber="6">Duvalidius</emphasis>
<bibRefCitation id="EFEB88C3C848BE34C6A05477FCE1C81D" author="Jeannel, R." box="[643,811,1605,1629]" pageId="5" pageNumber="6" pagination="1 - 808" refId="ref4875" refString="Jeannel, R. (1928): Monographie des Trechinae 3. Morphologie comparee et distribution geographique d'un groupe de Coleopteres - L'Abeille, Paris 35: 1 - 808." type="journal article" year="1928">JEANNEL, 1928</bibRefCitation>
</taxonomicName>
, currently treated as synonyms of
<taxonomicName id="4C7A8EB1C84ABE36C56F52A7FD40CEED" authority="BOKOR, 1922" authorityName="BOKOR" authorityYear="1922" box="[332,650,149,173]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Biharotrechus">
<emphasis id="B90E2920C84ABE36C56F52A7FE26CEED" box="[332,492,149,173]" italics="true" pageId="7" pageNumber="8">Biharotrechus</emphasis>
BOKOR, 1922
</taxonomicName>
, and
<taxonomicName id="4C7A8EB1C84ABE36C6EE52A7FBE9CEED" authority="BOKOR, 1922" authorityName="BOKOR" authorityYear="1922" box="[717,1059,149,173]" class="Insecta" family="Carabidae" genus="Hungarotrechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84ABE36C6EE52A7FC4FCEED" box="[717,901,149,173]" italics="true" pageId="7" pageNumber="8">Hungarotrechus</emphasis>
BOKOR, 1922
</taxonomicName>
, respectively (see
<bibRefCitation id="EFEB88C3C84ABE36C5A75288FDA7CE92" author="Moravec, P. &amp; Ueno, S. - I. &amp; Belousov, I. A." box="[388,621,186,210]" pageId="7" pageNumber="8" pagination="288 - 346" refId="ref5087" refString="Moravec, P., Ueno, S. - I. &amp; Belousov, I. A. (2003): Carabidae: Trechinae: Trechini. - In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Vol. 1., pp. 288 - 346, Apollo Books, Stenstrup." type="book chapter" year="2003">Moravec &amp; al. 2003</bibRefCitation>
, and
<bibRefCitation id="EFEB88C3C84ABE36C6915288FC8ECE92" author="Lorenz, W." box="[690,836,186,210]" pageId="7" pageNumber="8" refId="ref4932" refString="Lorenz, W. (2005): Systematic list of extant ground beetles of the world (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae. incl. Paussinae, Cicindelinae, Rhysodinae). Tutzing, iv + 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
) have been widely debated by
<bibRefCitation id="EFEB88C3C84ABE36C51852D2FD60CEB8" author="Casale, A. &amp; Vigna Taglianti, A." box="[315,682,224,248]" pageId="7" pageNumber="8" pagination="309 - 329" refId="ref4723" refString="Casale, A. &amp; Vigna Taglianti, A. (1984): Due nuovi Carabidi cavernicoli dell'Anatolia meridionale (Coleoptera, Carabidae) - Fragmenta entomologica (Roma) 17: 309 - 329." type="journal article" year="1984">Casale &amp; Vigna Taglianti (1984)</bibRefCitation>
. What is evident, is the fact that
<taxonomicName id="4C7A8EB1C84ABE36C4925337FE80CF5D" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[177,330,261,285]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C84ABE36C4925337FF0BCF5D" box="[177,193,261,285]" italics="true" pageId="7" pageNumber="8">D</emphasis>
. (
<emphasis id="B90E2920C84ABE36C4FA5337FF3ACF5D" box="[217,240,261,285]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
)
<emphasis id="B90E2920C84ABE36C5235337FE80CF5D" box="[256,330,261,285]" italics="true" pageId="7" pageNumber="8">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC84ABE36C5735337FE63CF5D" box="[336,425,261,285]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C7A8EB1C84ABE36C5C35337FD4DCF5D" authorityName="CASALE &amp; VIGNA TAGLIANTI" authorityYear="1984" box="[480,647,261,285]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bortesii" subGenus="Biharotrechus">
<emphasis id="B90E2920C84ABE36C5C35337FE3ACF5D" box="[480,496,261,285]" italics="true" pageId="7" pageNumber="8">D</emphasis>
. (
<emphasis id="B90E2920C84ABE36C6245337FDD5CF5D" box="[519,543,261,285]" italics="true" pageId="7" pageNumber="8">B.</emphasis>
)
<emphasis id="B90E2920C84ABE36C60C5337FD4DCF5D" box="[559,647,261,285]" italics="true" pageId="7" pageNumber="8">bortesii</emphasis>
</taxonomicName>
(the latter, described as
<taxonomicName id="4C7A8EB1C84ABE36C7805337FBD6CF5D" authorityName="JEANNEL" authorityYear="1928" box="[931,1052,261,285]" class="Insecta" family="Carabidae" genus="Duvaliotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84ABE36C7805337FBD6CF5D" box="[931,1052,261,285]" italics="true" pageId="7" pageNumber="8">Duvaliotes</emphasis>
</taxonomicName>
) form a homogeneous species group, and are so far the only representatives in Southern
<collectingCountry id="F36DB5A2C84ABE36C51E5362FE50CF28" box="[317,410,336,360]" name="Turkey" pageId="7" pageNumber="8">Anatolia</collectingCountry>
of a lineage widely spread in the Carpathians, the Balkan peninsula and the Dinaric Alps.
</paragraph>
<caption id="DF05A5BAC84BBE37C4F457CDFCD4C86B" ID-DOI="http://doi.org/10.5281/zenodo.5749782" ID-Zenodo-Dep="5749782" httpUri="https://zenodo.org/record/5749782/files/figure.png" pageId="6" pageNumber="7" startId="6.[215,244,1535,1553]" targetBox="[215,1099,147,1486]" targetPageId="6">
<paragraph id="8BC5F532C84BBE37C4F457CDFCD4C86B" blockId="6.[215,1100,1534,1579]" pageId="6" pageNumber="7">
<emphasis id="B90E2920C84BBE37C4F457CDFCBAC851" bold="true" box="[215,880,1534,1553]" pageId="6" pageNumber="7">
Fig. 3:
<taxonomicName id="4C7A8EB1C84BBE37C53457CDFDFFC851" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[279,565,1535,1553]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov." subGenus="Biharotrechus">
<emphasis id="B90E2920C84BBE37C53457CDFEAEC851" bold="true" box="[279,356,1535,1553]" italics="true" pageId="6" pageNumber="7">Duvalius</emphasis>
(
<emphasis id="B90E2920C84BBE37C55157CDFE24C851" bold="true" box="[370,494,1535,1553]" italics="true" pageId="6" pageNumber="7">Biharotrechus</emphasis>
)
<emphasis id="B90E2920C84BBE37C5DF57CDFDFFC851" bold="true" box="[508,565,1535,1553]" italics="true" pageId="6" pageNumber="7">huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC84BBE37C61F57CDFD4FC851" box="[572,645,1535,1553]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, type locality: Geyik Daği
</emphasis>
Mt. (Southern Anatolia, Toros chain, Antalya) (a); detail of the habitat (b) (photo C. Huber).
</paragraph>
</caption>
<paragraph id="8BC5F532C84ABE36C4F653A8FC25CDFD" blockId="7.[177,1067,149,957]" pageId="7" pageNumber="8">
Therefore, as shown by
<bibRefCitation id="EFEB88C3C84ABE36C5D153A8FCA0CFF2" author="Casale, A. &amp; Vigna Taglianti, A." box="[498,874,410,434]" pageId="7" pageNumber="8" pagination="277 - 406" refId="ref4760" refString="Casale, A. &amp; Vigna Taglianti, A. (1999): Caraboid beetles (excl. Cicindelidae) of Anatolia, and their biogeographical significance (Coleoptera, Caraboidea) - Biogeographia 20: 277 - 406." type="journal article" year="1999">Casale &amp; Vigna Taglianti (1999)</bibRefCitation>
, in
<collectingCountry id="F36DB5A2C84ABE36C7B453A8FC3CCFF2" box="[919,1014,410,434]" name="Turkey" pageId="7" pageNumber="8">Anatolia</collectingCountry>
the genus
<taxonomicName id="4C7A8EB1C84ABE36C52253F2FEACCF98" authorityName="Delarouzee" authorityYear="1859" box="[257,358,448,472]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84ABE36C52253F2FEACCF98" box="[257,358,448,472]" italics="true" pageId="7" pageNumber="8">Duvalius</emphasis>
</taxonomicName>
includes so far representatives of some different lineages: in the central and eastern Pontic chain, and in eastern
<collectingCountry id="F36DB5A2C84ABE36C71353D7FC5BCFBD" box="[816,913,485,509]" name="Turkey" pageId="7" pageNumber="8">Anatolia</collectingCountry>
, some forest dweller or troglophilic species, all probably belonging to the Caucasian
<emphasis id="B90E2920C84ABE36C7CC5038FF28CC08" italics="true" pageId="7" pageNumber="8">antoniae</emphasis>
species group. Markedly isolated seem to be
<taxonomicName id="4C7A8EB1C84ABE36C6C25002FBE9CC08" authority="PERRAULT, 1971" authorityName="PERRAULT" authorityYear="1971" box="[737,1059,560,584]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bicikensis">
<emphasis id="B90E2920C84ABE36C6C25002FCA7CC08" box="[737,877,560,584]" italics="true" pageId="7" pageNumber="8">D. bicikensis</emphasis>
PERRAULT, 1971
</taxonomicName>
, from the
<collectingRegion id="49BE3BD0C84ABE36C53A5067FEA5CC2D" box="[281,367,597,621]" country="Turkey" name="Giresun" pageId="7" pageNumber="8">Giresun</collectingRegion>
DağlarI, apparently related to the Eastern-European
<taxonomicName id="4C7A8EB1C84ABE36C7E35067FECFCCD2" authorityName="BOKOR" authorityYear="1922" class="Insecta" family="Carabidae" genus="Hungarotrechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84ABE36C7E35067FECFCCD2" italics="true" pageId="7" pageNumber="8">Hungarotrechus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C7A8EB1C84ABE36C5375048FE40CCD2" authorityName="JEANNEL" authorityYear="1928" box="[276,394,634,658]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B90E2920C84ABE36C5375048FE40CCD2" box="[276,394,634,658]" italics="true" pageId="7" pageNumber="8">Duvalidius</emphasis>
</taxonomicName>
of authors) complex, and
<taxonomicName id="4C7A8EB1C84ABE36C68E5048FBE8CCD2" authority="VIGNA TAGLIANTI, 1999" authorityName="VIGNA TAGLIANTI" authorityYear="1999" box="[685,1058,634,658]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bruschii">
<emphasis id="B90E2920C84ABE36C68E5048FCE9CCD2" box="[685,803,634,658]" italics="true" pageId="7" pageNumber="8">D. bruschii</emphasis>
VIGNA TAGLIANTI, 1999
</taxonomicName>
, probably related to the southern Balkan
<emphasis id="B90E2920C84ABE36C6AC5092FD25CCF8" box="[655,751,672,696]" italics="true" pageId="7" pageNumber="8">krueperi</emphasis>
-group. By contrast, in the Western Toros, the genus is represented by three peculiar, subterranean but oculate species, with very different relationships:
<taxonomicName id="4C7A8EB1C84ABE36C6C950D8FCAACD42" authorityName="CASALE &amp; VIGNA TAGLIANTI" authorityYear="1984" box="[746,864,746,770]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bortesii">
<emphasis id="B90E2920C84ABE36C6C950D8FCAACD42" box="[746,864,746,770]" italics="true" pageId="7" pageNumber="8">D. bortesii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A8EB1C84ABE36C7BA50D8FBCBCD42" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[921,1025,746,770]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="B90E2920C84ABE36C7BA50D8FBCBCD42" box="[921,1025,746,770]" italics="true" pageId="7" pageNumber="8">D. huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D945BC84ABE36C02A50D8FF2FCD68" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, treated in the present contribution, are apparently related to the Balkan
<taxonomicName id="4C7A8EB1C84ABE36C4925107FE9ACD0D" authorityName="BOKOR" authorityYear="1922" box="[177,336,821,845]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Biharotrechus">
<emphasis id="B90E2920C84ABE36C4925107FE9ACD0D" box="[177,336,821,845]" italics="true" pageId="7" pageNumber="8">Biharotrechus</emphasis>
</taxonomicName>
lineage; and
<taxonomicName id="4C7A8EB1C84ABE36C5C45107FCC0CD0D" authority="JEANNEL, 1934" authorityName="JEANNEL" authorityYear="1934" box="[487,778,821,845]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="huetheri">
<emphasis id="B90E2920C84ABE36C5C45107FDA9CD0D" box="[487,611,821,845]" italics="true" pageId="7" pageNumber="8">D. huetheri</emphasis>
JEANNEL, 1934
</taxonomicName>
, present in several caves of
<collectingRegion id="49BE3BD0C84ABE36C4F25168FEECCD32" box="[209,294,858,882]" country="Turkey" name="Antalya" pageId="7" pageNumber="8">Antalya</collectingRegion>
and
<collectingRegion id="49BE3BD0C84ABE36C5415168FE6DCD32" box="[354,423,858,882]" country="Turkey" name="Mugla" pageId="7" pageNumber="8">Muğla</collectingRegion>
provinces, redescribed with different names, is related to the &quot;Southern
<collectingRegion id="49BE3BD0C84ABE36C55F51B2FE1FCDD8" box="[380,469,896,920]" country="Greece" name="Voreio Aigaio" pageId="7" pageNumber="8">Aegean</collectingRegion>
&quot; lineage of
<taxonomicName id="4C7A8EB1C84ABE36C67D51B2FC1BCDD8" authority="ROTTENBERG, 1874" authorityName="ROTTENBERG" authorityYear="1874" box="[606,977,896,920]" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="diaphanus">
<emphasis id="B90E2920C84ABE36C67D51B2FD3CCDD8" box="[606,758,896,920]" italics="true" pageId="7" pageNumber="8">D. diaphanus</emphasis>
ROTTENBERG, 1874
</taxonomicName>
, and
<taxonomicName id="4C7A8EB1C84ABE36C03051B2FEB5CDFD" authority="DEUVE, 2001" authorityName="DEUVE" authorityYear="2001" class="Insecta" family="Carabidae" genus="Duvalidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mirei">
<emphasis id="B90E2920C84ABE36C03051B2FF21CDFD" italics="true" pageId="7" pageNumber="8">D. mirei</emphasis>
DEUVE, 2001
</taxonomicName>
, both endemic to Mt. Taygetos (
<collectingCountry id="F36DB5A2C84ABE36C6C35197FCFCCDFD" box="[736,822,933,957]" name="Greece" pageId="7" pageNumber="8">Greece</collectingCountry>
,
<collectingRegion id="49BE3BD0C84ABE36C71C5197FC2CCDFD" box="[831,998,933,957]" country="Greece" name="Peloponnisos" pageId="7" pageNumber="8">Peloponnesus</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>