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<document id="FDC16C7535813B36242251085A9C69A5" ID-CLB-Dataset="8891" ID-DOI="10.11646/zootaxa.4868.3.2" ID-GBIF-Dataset="03a9be04-f633-4adc-8397-058c282f216b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417761" ID-ZooBank="256D29E7-5148-415C-AD06-5F1F6D986887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609808401120" checkinUser="plazi" docAuthor="Cao, Yanghui, Dmitriev, Dmitry A., Dietrich, Christopher H. &amp; Zhang, Yalin" docDate="2020" docId="03998E57FFD9FFE68B84FA4BFEC2BA9A" docLanguage="en" docName="zootaxa.4868.3.2.pdf" docOrigin="Zootaxa 4868 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Etmaria brevis Cao &amp; Dmitriev 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="339" masterDocId="FFA0F62FFFDFFFEE8B13FFAFFD37BB13" masterDocTitle="Review of the leafhopper genus Thaia Ghauri and related genera (Hemiptera: Cicadellidae: Typhlocybinae: Erythroneurini)" masterLastPageNumber="367" masterPageNumber="331" pageNumber="337" updateTime="1698852417663" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="943E65F086E4B239434E765064BF6116" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE8F9C7A19916927CFCBE0A3E5EA861B">
<mods:title id="43F010544EE8C1143D8697C1F7E53D24">Review of the leafhopper genus Thaia Ghauri and related genera (Hemiptera: Cicadellidae: Typhlocybinae: Erythroneurini)</mods:title>
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<mods:name id="D8F7DFD221E23A4E46DCF507E34F85D8" type="personal">
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<mods:namePart id="E7D29FD542B42D4C286CD6705BB981B1">Cao, Yanghui</mods:namePart>
<mods:nameIdentifier id="F9DDBF6C2AA7BC9C2E1140F47C70B3BB" type="ORCID">0000-0002-0515-0767</mods:nameIdentifier>
<mods:affiliation id="190CAAC9DB307CCADDB8C9F8AADBB65D">Key Laboratory of Plant Protection Resources and Pest Management, National Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, P. R. China &amp; Illinois Nature History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S Oak St., Champaign, IL 61820, USA &amp; caoyh @ illinois. edu; https: // orcid. org / 0000 - 0002 - 0515 - 0767</mods:affiliation>
<mods:nameIdentifier id="BDC90201A8D4BFEA7B66841B280A3928" type="email">caoyh@illinois.edu</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="67338952BED54FB572D00F16320EC705">Dmitriev, Dmitry A.</mods:namePart>
<mods:nameIdentifier id="FFB5CFEFFA402C3B1CBC85755C720598" type="ORCID">0000-0002-3293-4794</mods:nameIdentifier>
<mods:affiliation id="5D90A5DD7EB55806DD46C4BE0846997B">Illinois Nature History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S Oak St., Champaign, IL 61820, USA &amp; arboridia @ gmail. com; https: // orcid. org / 0000 - 0002 - 3293 - 4794</mods:affiliation>
<mods:nameIdentifier id="E5D566E970C59254CC8645E52BCD877C" type="email">arboridia@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="41D1378C220D4CA85694253E26DA310D" type="personal">
<mods:role id="707602682311B224A6F9764328655B6B">
<mods:roleTerm id="D14DC67344CF982065C67A55C9C1A085">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E0887166F18400DC20C9DF2BC1960CB">Dietrich, Christopher H.</mods:namePart>
<mods:nameIdentifier id="1EEC55B6FA1D02EDD08A69ED82424161" type="ORCID">0000-0003-4005-4305</mods:nameIdentifier>
<mods:affiliation id="65B89A7AEAAFF70778CCE89CB6107E5A">Illinois Nature History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S Oak St., Champaign, IL 61820, USA &amp; chdietri @ illinois. edu; https: // orcid. org / 0000 - 0003 - 4005 - 4305</mods:affiliation>
<mods:nameIdentifier id="81B1F8470A2560908615590E45A9DCFC" type="email">chdietri@illinois.edu</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="C5E8EFDB0ABC27096142CC232AC50911">Zhang, Yalin</mods:namePart>
<mods:affiliation id="9B5778FBA486B379407008A7BF385ED2">Key Laboratory of Plant Protection Resources and Pest Management, National Ministry of Education, Entomological Museum, Northwest A &amp; F University, Yangling, Shaanxi 712100, P. R. China</mods:affiliation>
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<mods:title id="59E8D808F582A3CB2DF51FD39509C52C">Zootaxa</mods:title>
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<mods:part id="7A3E01BDB109FEE85E734CFDF697102D">
<mods:date id="2CBD0F210CA160704541DDBBA123B482">2020</mods:date>
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<mods:number id="A4E87E1C546BF9E6ADD01CA97F35C1B4">2020-10-28</mods:number>
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<mods:number id="D33FCCC7A38FA4E8867A4A5300EC285B">4868</mods:number>
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<mods:number id="F64C6FC265F626860EBF838732FCB62D">3</mods:number>
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<mods:identifier id="A9ACAF43532B53019F54E0B1EF6FC486" type="CLB-Dataset">8891</mods:identifier>
<mods:identifier id="3A7CBA2ADD35D49F2FEEBE72E07D50C5" type="DOI">10.11646/zootaxa.4868.3.2</mods:identifier>
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<treatment id="03998E57FFD9FFE68B84FA4BFEC2BA9A" ID-DOI="http://doi.org/10.5281/zenodo.4417826" ID-GBIF-Taxon="176583710" ID-Zenodo-Dep="4417826" LSID="urn:lsid:plazi:treatment:03998E57FFD9FFE68B84FA4BFEC2BA9A" httpUri="http://treatment.plazi.org/id/03998E57FFD9FFE68B84FA4BFEC2BA9A" lastPageId="8" lastPageNumber="339" pageId="6" pageNumber="337">
<subSubSection id="C32A6CCAFFD9FFE88B84FA4BFFAABEED" box="[151,669,1507,1534]" pageId="6" pageNumber="337" type="nomenclature">
<paragraph id="8B8F3F41FFD9FFE88B84FA4BFFAABEED" blockId="6.[151,669,1507,1570]" box="[151,669,1507,1534]" pageId="6" pageNumber="337">
<heading id="D0C7882DFFD9FFE88B84FA4BFFAABEED" bold="true" box="[151,669,1507,1534]" fontSize="11" level="1" pageId="6" pageNumber="337" reason="1">
<emphasis id="B944E353FFD9FFE88B84FA4BFFAABEED" bold="true" box="[151,669,1507,1534]" pageId="6" pageNumber="337">
2.
<taxonomicName id="4C3044C2FFD9FFE88BA6FA4BFF0FBEED" ID-CoL="8KMYP" authority="Cao &amp; Dmitriev" authorityName="Cao &amp; Dmitriev" authorityYear="2020" box="[181,568,1507,1534]" class="Insecta" family="Cicadellidae" genus="Etmaria" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B944E353FFD9FFE88BA6FA4BFC50BEED" bold="true" box="[181,359,1507,1534]" italics="true" pageId="6" pageNumber="337">Etmaria brevis</emphasis>
Cao &amp; Dmitriev
</taxonomicName>
<taxonomicNameLabel id="A2775E28FFD9FFE88953FA4BFFAABEED" box="[576,669,1508,1534]" pageId="6" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFD9FFE88B84F9A7FF45BCC0" pageId="6" pageNumber="337" type="description">
<paragraph id="8B8F3F41FFD9FFE88B84F9A7FC13BD31" blockId="6.[151,669,1507,1570]" box="[151,292,1544,1570]" pageId="6" pageNumber="337">
<figureCitation id="130B23C4FFD9FFE88B84F9A7FDEABD31" box="[151,221,1544,1570]" captionStart="FIGURE 3" captionStartId="7.[151,250,1850,1874]" captionTargetBox="[161,1436,181,1815]" captionTargetId="figure-19@7.[151,1436,181,1827]" captionTargetPageId="7" captionText="FIGURE 3. Etmaria brevis Cao &amp; Dmitriev sp. nov. a. abdominal apodemes 2S; b. genital capsule; c. anal tube, pygofer lobe and pygofer ventral appendages, caudal view; d. anal tube appendage, lateral view; e. subgenital plate, style, connective and sternite IX; f. subgenital plate, dorsal view; g. style, dorsal view; h. connective; i. aedeagus, lateral view; j. aedeagus, caudal view." figureDoi="http://doi.org/10.5281/zenodo.4417767" httpUri="https://zenodo.org/record/4417767/files/figure.png" pageId="6" pageNumber="337">Figs 3</figureCitation>
,
<figureCitation id="130B23C4FFD9FFE88BF9F9A7FC33BD31" box="[234,260,1544,1570]" captionStart="FIGURE 18" captionStartId="31.[151,250,1866,1890]" captionTargetBox="[159,1428,192,1832]" captionTargetId="figure-19@31.[148,1439,181,1842]" captionTargetPageId="31" captionText="FIGURE 18. a–d. Nlunga parareeneni Cao &amp; Dietrich sp. nov.; e–h. Nlunga reeneni Dworakowska, 1974; i–l. Etmaria brevis Cao &amp; Dmitriev sp. nov.; m–p. Etmaria chaiyahumica Cao &amp; Dmitriev sp. nov.; q–t. Etmaria dentata Cao &amp; Dmitriev sp. nov.; u–x. Etmaria indonesica Cao &amp; Dietrich sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4417801" httpUri="https://zenodo.org/record/4417801/files/figure.png" pageId="6" pageNumber="337">18</figureCitation>
i–l
</paragraph>
<paragraph id="8B8F3F41FFD9FFE88B84F9FFFCECBDA1" blockId="6.[151,1437,1616,2003]" pageId="6" pageNumber="337">
<emphasis id="B944E353FFD9FFE88B84F9FFFC1EBD79" bold="true" box="[151,297,1616,1642]" pageId="6" pageNumber="337">Description.</emphasis>
Ground color light tawny, dorsum infuscated (specimen preserved in ethanol before examined), eyes grey, frontoclypeal area and anteclypeus tawny. Crown fore margin parallel to hind margin, coronal suture distinct; anteclypeus relatively broad.
</paragraph>
<paragraph id="8B8F3F41FFD9FFE88BD4F913FE4EBDC5" blockId="6.[151,1437,1616,2003]" box="[199,889,1724,1751]" pageId="6" pageNumber="337">Anal tube appendage broad, extended from apex of anal tube.</paragraph>
<paragraph id="8B8F3F41FFD9FFE88BD4F94FF9E8BCBD" blockId="6.[151,1437,1616,2003]" pageId="6" pageNumber="337">Pygofer with oblique boundary between basal and distal lobes; ventral appendage tubular, curved inwards, not surpassing hind margin of lobe, without rigid setae at base. Subgenital plate with about 3 macrosetae. Style with apical part slim and long, curved outwards in dorsal view. Connective with manubrium long and relatively broad, widened apically, margin of central lobe straight. Aedeagal shaft long, curved dorso-cephalad, atrium long but narrow in lateral view, broadened in caudal view, with serrated lamellae on dorso-lateral edges near base; dorsal apodeme rudimentary; paired preatrial processes short and straight, shifted to base of shaft, near each other.</paragraph>
<paragraph id="8B8F3F41FFD9FFE88BD4F817FF45BCC0" blockId="6.[151,1437,1616,2003]" box="[199,626,1976,2003]" pageId="6" pageNumber="337">
<emphasis id="B944E353FFD9FFE88BD4F817FC44BCC1" bold="true" box="[199,371,1976,2002]" pageId="6" pageNumber="337">Measurement.</emphasis>
Male length 3.10 mm.
</paragraph>
</subSubSection>
<caption id="DF4F6FC9FFD8FFE98B84F895FDF9BCAC" ID-DOI="http://doi.org/10.5281/zenodo.4417767" ID-Zenodo-Dep="4417767" httpUri="https://zenodo.org/record/4417767/files/figure.png" pageId="7" pageNumber="338" startId="7.[151,250,1850,1874]" targetBox="[161,1436,181,1815]" targetPageId="7">
<paragraph id="8B8F3F41FFD8FFE98B84F895FDF9BCAC" blockId="7.[151,1437,1850,1983]" pageId="7" pageNumber="338">
<emphasis id="B944E353FFD8FFE98B84F895FC3ABC40" bold="true" box="[151,269,1850,1875]" pageId="7" pageNumber="338">FIGURE 3</emphasis>
.
<taxonomicName id="4C3044C2FFD8FFE98A09F894FF6ABC40" authority="Cao &amp; Dmitriev" authorityName="Cao &amp; Dmitriev" authorityYear="2020" box="[282,605,1851,1875]" class="Insecta" family="Cicadellidae" genus="Etmaria" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="B944E353FFD8FFE98A09F894FC86BC40" box="[282,433,1851,1875]" italics="true" pageId="7" pageNumber="338">Etmaria brevis</emphasis>
Cao &amp; Dmitriev
</taxonomicName>
<emphasis id="B944E353FFD8FFE98977F894FF82BC40" bold="true" box="[612,693,1851,1875]" pageId="7" pageNumber="338">
<taxonomicNameLabel id="A2775E28FFD8FFE98977F894FF82BC40" box="[612,693,1851,1875]" pageId="7" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
a. abdominal apodemes 2S; b. genital capsule; c. anal tube, pygofer lobe and pygofer ventral appendages, caudal view; d. anal tube appendage, lateral view; e. subgenital plate, style, connective and sternite IX; f. subgenital plate, dorsal view; g. style, dorsal view; h. connective; i. aedeagus, lateral view; j. aedeagus, caudal view.
</paragraph>
</caption>
<subSubSection id="C32A6CCAFFD7FFE68BD4FF39F9C6BBC6" pageId="8" pageNumber="339" type="materials_examined">
<paragraph id="8B8F3F41FFD7FFE68BD4FF39F9C6BBC6" blockId="8.[151,1437,150,393]" pageId="8" pageNumber="339">
<emphasis id="B944E353FFD7FFE68BD4FF39FC82BBA3" bold="true" box="[199,437,150,176]" pageId="8" pageNumber="339">Material examined.</emphasis>
<materialsCitation id="3B58351CFFD7FFE68AACFF38F9C6BBC6" ID-GBIF-Occurrence="3015520324" collectingDate="2008-10-19" collectingDateMax="2008-10-26" collectingDateMin="2008-10-19" collectionCode="QSBG" collectorName="Yai &amp; Amnad." country="Thailand" elevation="200" latitude="12.21315" location="Prachuab Khiri Khan" longLatPrecision="1" longitude="99.9777" municipality="Checkpoint" pageId="8" pageNumber="339" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="548B81E3FFD7FFE68AACFF38FF1BBBA2" box="[447,556,151,177]" pageId="8" pageNumber="339" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D36F4C8FFD7FFE68929FF38FF66BBA2" box="[570,593,151,177]" pageId="8" pageNumber="339" type="male">♂</specimenCount>
,
<collectingCountry id="F3277FD1FFD7FFE68972FF38FFC2BBA2" box="[609,757,151,177]" name="Thailand" pageId="8" pageNumber="339">THAILAND</collectingCountry>
,
<location id="8EEF699AFFD7FFE68817FF38FECBBBA2" LSID="urn:lsid:plazi:treatment:03998E57FFD9FFE68B84FA4BFEC2BA9A:8EEF699AFFD7FFE68817FF38FECBBBA2" box="[772,1020,151,177]" country="Thailand" latitude="12.21315" longLatPrecision="1" longitude="99.9777" municipality="Checkpoint" name="Prachuab Khiri Khan" pageId="8" pageNumber="339">Prachuab Khiri Khan</location>
,
<collectorName id="26C55A97FFD7FFE68F1FFF38F82EBBA2" box="[1036,1305,151,177]" pageId="8" pageNumber="339">Khao Sam Roi Yot NP</collectorName>
,
<quantity id="4CC892A4FFD7FFE68E34FF38F85FBBA2" box="[1319,1384,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="339" unit="m" value="200.0">
<elevation id="001DD872FFD7FFE68E34FF38F85FBBA2" box="[1319,1384,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="339" unit="m" value="200.0">200m</elevation>
</quantity>
/s of
<collectingMunicipality id="6BEBA53BFFD7FFE68B84FF14FC17BBC6" box="[151,288,187,213]" pageId="8" pageNumber="339">Checkpoint</collectingMunicipality>
1,
<geoCoordinate id="EE045986FFD7FFE68A27FF14FCF3BBC6" box="[308,452,187,213]" degrees="12" direction="north" minutes="12.789" orientation="latitude" pageId="8" pageNumber="339" precision="1" value="12.21315">12°12.789'N</geoCoordinate>
,
<geoCoordinate id="EE045986FFD7FFE68AC3FF15FF56BBC6" box="[464,609,186,213]" degrees="99" direction="east" minutes="58.662" orientation="longitude" pageId="8" pageNumber="339" precision="1" value="99.9777">99°58.662‘E</geoCoordinate>
,
<date id="FF8E1981FFD7FFE6897FFF14FE3EBBC6" box="[620,777,187,213]" pageId="8" pageNumber="339" value="2008-10-19" valueMax="2008-10-26" valueMin="2008-10-19">
<collectingDate id="EFCAE069FFD7FFE6897FFF14FE3EBBC6" box="[620,777,187,213]" pageId="8" pageNumber="339" value="2008-10-19" valueMax="2008-10-26" valueMin="2008-10-19">19–26 x 2008</collectingDate>
</date>
,
<collectingMethod id="52714756FFD7FFE68807FF14FE95BBC6" box="[788,930,187,213]" pageId="8" pageNumber="339">Malaise trap</collectingMethod>
, coll.
<collectorName id="26C55A97FFD7FFE688F7FF14F93DBBC6" box="[996,1034,187,213]" pageId="8" pageNumber="339">Yai</collectorName>
&amp;
<collectorName id="26C55A97FFD7FFE68F3FFF14F9B1BBC6" box="[1068,1158,187,213]" pageId="8" pageNumber="339">Amnad.</collectorName>
[
<collectionCode id="ED21A784FFD7FFE68F87FF15F9D0BBC6" box="[1172,1255,186,213]" pageId="8" pageNumber="339">QSBG</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFD7FFE68BD4FF70FF6BBA0E" pageId="8" pageNumber="339" type="etymology">
<paragraph id="8B8F3F41FFD7FFE68BD4FF70FF6BBA0E" blockId="8.[151,1437,150,393]" pageId="8" pageNumber="339">
<emphasis id="B944E353FFD7FFE68BD4FF70FC79BBEA" bold="true" box="[199,334,223,249]" pageId="8" pageNumber="339">Etymology.</emphasis>
The new specific epithet is derived from the Latin adjective “
<taxonomicName id="4C3044C2FFD7FFE68F09FF70F96ABBEA" authorityName="Cao &amp; Dmitriev" authorityYear="2020" box="[1050,1117,223,249]" class="Insecta" family="Cicadellidae" genus="Etmaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="B944E353FFD7FFE68F09FF70F96ABBEA" box="[1050,1117,223,249]" italics="true" pageId="8" pageNumber="339">brevis</emphasis>
</taxonomicName>
” which means short, small, referring to the short preatrial processes.
</paragraph>
</subSubSection>
<subSubSection id="C32A6CCAFFD7FFE68BD4FE89FEC2BA9A" pageId="8" pageNumber="339" type="diagnosis">
<paragraph id="8B8F3F41FFD7FFE68BD4FE89FEC2BA9A" blockId="8.[151,1437,150,393]" pageId="8" pageNumber="339">
<emphasis id="B944E353FFD7FFE68BD4FE89FC75BA53" bold="true" box="[199,322,294,320]" pageId="8" pageNumber="339">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="4C3044C2FFD7FFE68980FE88FED4BA51" authority="(Dworakowska)" authorityName="Cao &amp; Dmitriev &amp; Dietrich &amp; Zhang" authorityYear="2020" baseAuthorityName="Dworakowska" baseAuthorityYear="1984" box="[659,995,295,322]" class="Insecta" family="Cicadellidae" genus="Etmaria" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="decembris">
<emphasis id="B944E353FFD7FFE68980FE88FE12BA52" box="[659,805,295,321]" italics="true" pageId="8" pageNumber="339">E. decembris</emphasis>
(Dworakowska)
</taxonomicName>
, but the anal tube appendage is broader and curved, the pygofer ventral appendage is curved, the style has a much longer apex, and the aedeagal shaft is curved dorso-cephalad with the dorso-lateral edges serrated and lamelliform.
</paragraph>
</subSubSection>
</treatment>
</document>
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