234 lines
31 KiB
XML
234 lines
31 KiB
XML
<document id="E659E7EDEA9CCC4B2ADAB189BC6DC686" ID-DOI="10.5281/zenodo.182366" ID-GBIF-Dataset="1d12401f-e456-4790-86dd-85dc3b935a6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="182366" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459860504177" checkinUser="plazi" docAuthor="Liu, Xingyue, Hayashi, Fumio & Yang, Ding" docDate="2008" docId="03992C27AE45E714ADA0F929FE99FB9B" docLanguage="en" docName="zt01782p060.pdf" docOrigin="Zootaxa 1782" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Protohermes tortuosus Liu, Hayashi & Yang, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFA0545FAE43E71DAD37FFF5FC67FFB9" masterDocTitle="Species of the Protohermes sabahensis group (Megaloptera: Corydalidae) from eastern Malaysia" masterLastPageNumber="60" masterPageNumber="49" pageNumber="55" updateTime="1698225994607" updateUser="plazi">
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<mods:mods id="0206C5369A8C10437124DFD1A1E671FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6118017E5E684D6923310325423AD846">
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<mods:title id="056848C9A993680BDBDCA0054ABD8483">Species of the Protohermes sabahensis group (Megaloptera: Corydalidae) from eastern Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="6B330629340BB1E0C6FD9C76E3B1599B" type="personal">
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<mods:roleTerm id="70AF3CA4602134FD69798068BF8B25BB">Author</mods:roleTerm>
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<mods:namePart id="12D1B92EA44BEE14DF2EFEE15B2C9F1D">Liu, Xingyue</mods:namePart>
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<mods:roleTerm id="BB307188874874FE2272B6CCD9C5A3FD">Author</mods:roleTerm>
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<mods:namePart id="4044C6C01115CD2895C25FBE109CE99E">Hayashi, Fumio</mods:namePart>
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<mods:name id="D57C662E0DEF5A7924DA31CE580258D5" type="personal">
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<mods:roleTerm id="3649932A648BE266B58D6252BD806488">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="070147BAFE08BC23F2CE6CDB9276DB0D">Yang, Ding</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A281E82B5318D3F7AE91CD395CC36483">text</mods:typeOfResource>
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<mods:title id="03E2FF56A2A221D943420F69880AC212">Zootaxa</mods:title>
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<mods:part id="419CFBF0FA12176B0C5DF47267BEBB5E">
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<mods:date id="A1BB1F8A12822198325CBC8E7FF920B9">2008</mods:date>
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<mods:detail id="4F69342A92079C098574E2D2549ACB01" type="volume">
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<mods:number id="667847E0F56C8A73347E68BE1ED0EE7F">1782</mods:number>
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<mods:start id="A8C5044DA77B253FD236D597833FFA33">49</mods:start>
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<mods:classification id="F5212DC85EA775D4C0104C8BFDB9C3C8">journal article</mods:classification>
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<mods:identifier id="0E2F10A4AF735E430D5AAAD3A1920E9B" type="DOI">10.5281/zenodo.182366</mods:identifier>
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<mods:identifier id="C816B624D27E6F7362A326319A2DF403" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03992C27AE45E714ADA0F929FE99FB9B" ID-DOI="http://doi.org/10.5281/zenodo.6234901" ID-GBIF-Taxon="119368121" ID-Zenodo-Dep="6234901" LSID="urn:lsid:plazi:treatment:03992C27AE45E714ADA0F929FE99FB9B" httpUri="http://treatment.plazi.org/id/03992C27AE45E714ADA0F929FE99FB9B" lastPageId="9" lastPageNumber="58" pageId="6" pageNumber="55">
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<subSubSection id="C32ACEBAAE45E71BADA0F929FD0FF8A0" pageId="6" pageNumber="55" type="nomenclature">
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<paragraph id="8B8F9D31AE45E71BADA0F929FE69F94C" blockId="6.[151,526,1755,1817]" box="[151,526,1755,1782]" pageId="6" pageNumber="55">
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<heading id="D0C72A5DAE45E71BADA0F929FE69F94C" bold="true" box="[151,526,1755,1782]" fontSize="11" level="1" pageId="6" pageNumber="55" reason="1">
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<emphasis id="B9444123AE45E71BADA0F929FE69F94C" bold="true" box="[151,526,1755,1782]" pageId="6" pageNumber="55">
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<taxonomicName id="4C30E6B2AE45E71BADA0F929FDCEF94F" ID-CoL="4N5JW" box="[151,425,1756,1782]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="tortuosus" status="sp. nov.">
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<emphasis id="B9444123AE45E71BADA0F929FDCEF94F" bold="true" box="[151,425,1756,1782]" italics="true" pageId="6" pageNumber="55">Protohermes tortuosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277FC58AE45E71BAC87F92EFE69F94C" box="[432,526,1755,1781]" pageId="6" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F9D31AE45E71BADA0F90AFD0FF8A0" blockId="6.[151,526,1755,1817]" box="[151,360,1791,1817]" pageId="6" pageNumber="55">
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(
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<figureCitation id="130B81B4AE45E71BADA8F90AFD6EF8A0" box="[159,265,1791,1817]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,269,1850,1874]" captionTargetBox="[230,1356,194,1831]" captionTargetId="figure@5.[230,1356,194,1836]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Adults of Protohermes spp. 4. P. spectabilis sp. nov. (paratype), female; 5. P. t o r t u o s u s sp. nov. (holotype), male; 6. P. tortuosus sp. nov. (paratype), female. Scale lines: 5.0 mm." httpUri="https://zenodo.org/record/182368/files/figure.png" pageId="6" pageNumber="55">Figs 5–6</figureCitation>
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,
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<figureCitation id="130B81B4AE45E71BAC22F90AFD07F8A0" box="[277,352,1791,1817]" captionStart-0="FIGURES 23 – 31" captionStart-1="FIGURE 32" captionStartId-0="8.[151,269,1556,1580]" captionStartId-1="9.[151,255,1735,1759]" captionTargetBox-0="[178,1399,315,1462]" captionTargetBox-1="[452,1134,1114,1734]" captionTargetId-0="figure@8.[151,1436,277,1533]" captionTargetId-1="figure@9.[452,1136,1114,1761]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." captionText-1="FIGURE 32. Distribution map of the P. sabahensis species – group. P. sabahensis sp. nov. (); P. spectabilis sp. nov. (); P. tortuosus sp. nov. ()." httpUri-0="https://zenodo.org/record/182371/files/figure.png" httpUri-1="https://zenodo.org/record/182372/files/figure.png" pageId="6" pageNumber="55">23–32</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32ACEBAAE45E714ADA0F8BBF9E7FEC3" lastPageId="9" lastPageNumber="58" pageId="6" pageNumber="55" type="diagnosis">
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<paragraph id="8B8F9D31AE45E71BADA0F8BBF811F858" blockId="6.[151,1436,1870,2017]" pageId="6" pageNumber="55">
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<emphasis id="B9444123AE45E71BADA0F8BBFD70F8D1" bold="true" box="[151,279,1870,1896]" pageId="6" pageNumber="55">Diagnosis.</emphasis>
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This new species is defined by a combination of a yellow pronotum with two pairs of broad black markings, by the forewing having several median longitudinal veins alternating in black and yellow coloration, by the male ninth sternum being deeply incised posteriorly, and by the distal half of male tenth tergum being strongly broadened medially then abruptly narrowed to a dorsally pointing tip.
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</paragraph>
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<paragraph id="8B8F9D31AE44E71AADF1FF6DFF5CFEEB" blockId="7.[151,1436,152,338]" pageId="7" pageNumber="56">
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Male. Body length
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<quantity id="4CC830D4AE44E71AAC9FFF6DFD9CFF0B" box="[424,507,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="56" unit="mm" value="16.0">16 mm</quantity>
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(n=1); forewing length
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<quantity id="4CC830D4AE44E71AAE23FF6DFFF4FF0B" box="[788,915,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="7" pageNumber="56" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25–28 mm</quantity>
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(n=2), hindwing length
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<quantity id="4CC830D4AE44E71AA987FF6DF954FF0B" box="[1200,1331,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="7" pageNumber="56" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22–25 mm</quantity>
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. Head (
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<figureCitation id="130B81B4AE44E71AAC23FF35FDFBFF63" box="[276,412,192,218]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="7" pageNumber="56">Figs 23–24</figureCitation>
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) yellow; vertex with pair of small suboblong black markings near posterolateral margins, but black markings occasionally absent; postocular spine absent. Compound eyes brown; ocelli yellow, medially margined black; posterior ocelli close to median ocellus. Antenna black with scape and pedicel yellow. Mouthparts yellow; mandibles with distal half black.
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</paragraph>
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<caption id="DF4FCDB9AE44E71AADA0FA19FE30F9D2" httpUri="https://zenodo.org/record/182370/files/figure.png" pageId="7" pageNumber="56" targetBox="[170,1400,413,1453]" targetPageId="7">
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<paragraph id="8B8F9D31AE44E71AADA0FA19FE30F9D2" blockId="7.[151,1437,1516,1643]" pageId="7" pageNumber="56">
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<emphasis id="B9444123AE44E71AADA0FA19FD3AF9BD" bold="true" box="[151,349,1516,1540]" pageId="7" pageNumber="56">FIGURES 15–22.</emphasis>
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<taxonomicName id="4C30E6B2AE44E71AAC54FA18FE03F9BD" box="[355,612,1517,1540]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="spectabilis" status="sp. nov.">
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<emphasis id="B9444123AE44E71AAC54FA18FE03F9BD" box="[355,612,1517,1540]" italics="true" pageId="7" pageNumber="56">Protohermes spectabilis</emphasis>
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</taxonomicName>
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<emphasis id="B9444123AE44E71AAF5CFA19FEA7F9BD" bold="true" box="[619,704,1516,1540]" pageId="7" pageNumber="56">
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<taxonomicNameLabel id="A277FC58AE44E71AAF5CFA19FEA7F9BD" box="[619,704,1516,1540]" pageId="7" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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15. Male head and prothorax, dorsal view; 16. Male genitalia, dorsal view; 17. Male genitalia, ventral view; 18. Male genitalia, lateral view; 19. Male ninth gonostylus, ventral view; 20. Male tenth sternum, ventral view; 21. Female genitalia, lateral view; 22. Female eighth sternum, ventral view. Scale lines: 1.0 mm. Pdm: dorsomedial process.
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</paragraph>
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</caption>
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<paragraph id="8B8F9D31AE44E715ADF2F96AF803FF63" blockId="7.[151,1436,1695,2004]" lastBlockId="8.[151,1436,152,222]" lastPageId="8" lastPageNumber="57" pageId="7" pageNumber="56">
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Thorax yellow; pronotum (
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<figureCitation id="130B81B4AE44E71AAF34F96AFE3FF900" box="[515,600,1695,1721]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="7" pageNumber="56">Fig. 23</figureCitation>
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) with two pairs of broad black markings near lateral margins, posterior pair of markings slightly larger than anterior pair. Thoracic pilosity yellowish, much longer on meso– and metathorax than that on prothorax. Legs yellow, with dense, yellowish short setae; tarsal claws pale reddish brown. Wings (
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<figureCitation id="130B81B4AE44E71AAC7CF8E2FDF7F888" box="[331,400,1815,1841]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,269,1850,1874]" captionTargetBox="[230,1356,194,1831]" captionTargetId="figure@5.[230,1356,194,1836]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Adults of Protohermes spp. 4. P. spectabilis sp. nov. (paratype), female; 5. P. t o r t u o s u s sp. nov. (holotype), male; 6. P. tortuosus sp. nov. (paratype), female. Scale lines: 5.0 mm." httpUri="https://zenodo.org/record/182368/files/figure.png" pageId="7" pageNumber="56">Fig. 5</figureCitation>
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) hyaline, with several yellowish markings. Forewing slightly brownish; proximally with a subtriangular marking, medially with several irregular markings usually fused with each other, and with a round marking at distal 1/3; membrane between proximal and median markings darkened as a narrow transverse band, posteriorly extending to Cu1. Hindwing almost hyaline with marginal area slightly brownish. Veins brown but pale yellow in yellowish markings and proximal half of hindwings, longitudinal veins on proximal half of forewing alternating in black and yellow. Rs 8 to 9–branched, last branch bifurcate; 7–9 crossveins between R1 and Rs; M1+2 6–branched, M3+4 2–branched; 1A 3–branched.
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</paragraph>
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<caption id="DF4FCDB9AE4BE715ADA0F9E1FF65F92A" httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="8" pageNumber="57" targetBox="[178,1399,315,1462]" targetPageId="8">
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<paragraph id="8B8F9D31AE4BE715ADA0F9E1FF65F92A" blockId="8.[151,1436,1556,1683]" pageId="8" pageNumber="57">
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<emphasis id="B9444123AE4BE715ADA0F9E1FD39F995" bold="true" box="[151,350,1556,1580]" pageId="8" pageNumber="57">FIGURES 23–31.</emphasis>
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<taxonomicName id="4C30E6B2AE4BE715AC53F9E0FE30F995" box="[356,599,1557,1580]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="tortuosus" status="sp. nov.">
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<emphasis id="B9444123AE4BE715AC53F9E0FE30F995" box="[356,599,1557,1580]" italics="true" pageId="8" pageNumber="57">Protohermes tortuosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277FC58AE4BE715AF69F9E1FED3F995" box="[606,692,1556,1580]" pageId="8" pageNumber="57" rank="species">
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<emphasis id="B9444123AE4BE715AF69F9E1FED3F995" bold="true" box="[606,692,1556,1580]" pageId="8" pageNumber="57">sp. nov.</emphasis>
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</taxonomicNameLabel>
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23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm.
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</paragraph>
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</caption>
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<paragraph id="8B8F9D31AE4BE715ADF2F932FEDCF868" blockId="8.[151,1437,1735,2001]" pageId="8" pageNumber="57">
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Abdomen yellow. Ninth tergum (
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<figureCitation id="130B81B4AE4BE715AF7FF932FEFBF958" box="[584,668,1735,1761]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="8" pageNumber="57">Fig. 25</figureCitation>
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) subtrapezoidal, with deeply concave anterior and shallowly convex posterior margins. Ninth sternum (
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<figureCitation id="130B81B4AE4BE715AF6CF91AFED6F8B0" box="[603,689,1775,1801]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="8" pageNumber="57">Fig. 26</figureCitation>
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) with posterior margin deeply incised, forming a pair of slender attenuate processes. Ninth gonostylus (
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<figureCitation id="130B81B4AE4BE715AF5FF8E2FEA7F888" box="[616,704,1815,1841]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="8" pageNumber="57">Fig. 28</figureCitation>
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) slender, unguiform, nearly as long as ninth tergum, with tip slightly curved. Tenth tergum (
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<figureCitation id="130B81B4AE4BE715AF31F8CAFEF3F8E0" box="[518,660,1855,1881]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="8" pageNumber="57">Figs. 25–27</figureCitation>
|
||
) flattened, about twice as long as ninth tergum, twisted medially, bearing long hair–like setae medially; in lateral view strongly broadened medially with tip constricted and curve dorsad to form a distinct point. Tenth sternum (
|
||
<figureCitation id="130B81B4AE4BE715AE3AF87AFF04F810" box="[781,867,1935,1961]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="8" pageNumber="57">Fig. 29</figureCitation>
|
||
) arched; dorsomedial process nearly absent; lateral lobes long digitiform, straightly produced.
|
||
</paragraph>
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FF6DFD28FF63" blockId="9.[151,1437,152,1058]" pageId="9" pageNumber="58">
|
||
Female. Body length
|
||
<quantity id="4CC830D4AE4AE714ACF5FF6DFE72FF0B" box="[450,533,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="58" unit="mm" value="19.0">19 mm</quantity>
|
||
(n=1); forewing length
|
||
<quantity id="4CC830D4AE4AE714AE19FF6DFFCAFF0B" box="[814,941,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="9" pageNumber="58" unit="mm" value="33.0" valueMax="35.0" valueMin="31.0">31–35 mm</quantity>
|
||
(n=6), hindwing length
|
||
<quantity id="4CC830D4AE4AE714A9FBFF6DF92AFF0B" box="[1228,1357,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="9" pageNumber="58" unit="mm" value="31.0" valueMax="33.0" valueMin="29.0">29–33 mm</quantity>
|
||
. Color similar to male.
|
||
</paragraph>
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FF1DF9E7FEC3" blockId="9.[151,1437,152,1058]" pageId="9" pageNumber="58">
|
||
Eighth sternum (
|
||
<figureCitation id="130B81B4AE4AE714ACBBFF1DFE7DFEBB" box="[396,538,232,258]" captionStart="FIGURES 23 – 31" captionStartId="8.[151,269,1556,1580]" captionTargetBox="[178,1399,315,1462]" captionTargetId="figure@8.[151,1436,277,1533]" captionTargetPageId="8" captionText="FIGURES 23 – 31. Protohermes tortuosus sp. nov. 23. Male head and prothorax, dorsal view (holotype); 24. Male head, dorsal view (paratype); 25. Male genitalia, dorsal view; 26. Male genitalia, ventral view; 27. Male genitalia, lateral view; 28. Male ninth gonostylus, ventral view; 29. Male tenth sternum, ventral view; 30. Female genitalia, lateral view; 31. Female eighth sternum, ventral view. Scale lines: 1.0 mm." httpUri="https://zenodo.org/record/182371/files/figure.png" pageId="9" pageNumber="58">Figs. 30–31</figureCitation>
|
||
) subtrapezoidal in lateral view, with posterior portion slightly rugose medially; in ventral view, posterior margin with small, medial incision. Ninth gonocoxite with distal portion subquadrate; posteroventral corner strongly curved anteriorly with a small gonostylus at tip. Tenth tergum short with posterior margin medially incised, leaving a short subtriangular dorsal and a semicircular ventral lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ACEBAAE4AE714ADF2FE72FFF5FCB3" pageId="9" pageNumber="58" type="materials_examined">
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FE72FD2CFDA3" blockId="9.[151,1437,152,1058]" pageId="9" pageNumber="58">
|
||
<emphasis id="B9444123AE4AE714ADF2FE72FDEBFE18" bold="true" box="[197,396,391,417]" pageId="9" pageNumber="58">
|
||
<typeStatus id="548B2393AE4AE714ADF2FE72FD65FE18" box="[197,258,391,417]" pageId="9" pageNumber="58">Type</typeStatus>
|
||
materials.
|
||
</emphasis>
|
||
<typeStatus id="548B2393AE4AE714ACAEFE7DFE61FE1B" box="[409,518,392,418]" pageId="9" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
ɗ, Eastern
|
||
<collectingCountry id="F327DDA1AE4AE714AFA8FE7DFF68FE1B" box="[671,783,392,418]" name="Malaysia" pageId="9" pageNumber="58">Malaysia</collectingCountry>
|
||
:
|
||
<collectingCountry id="F327DDA1AE4AE714AE28FE7DFFE0FE1B" box="[799,903,392,418]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
, Bau dist., Pangkalan Tebang,
|
||
<quantity id="4CC830D4AE4AE714A83CFE7DF9FFFE1B" box="[1291,1432,392,418]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="9" pageNumber="58" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300–450 m</quantity>
|
||
, 1°16N, 110°4E, 5/
|
||
<date id="FF8EBBF1AE4AE714AC44FE45FD8DFE73" box="[371,490,432,458]" pageId="9" pageNumber="58" value="1958-09-08">8.IX.1958</date>
|
||
, T. C. Maa (
|
||
<collectionCode id="ED2105F4AE4AE714AFB2FE45FEBAFE73" LSID="urn:lsid:biocol.org:col:34697" box="[645,733,432,458]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="58">BPBM</collectionCode>
|
||
);
|
||
<typeStatus id="548B2393AE4AE714AFC4FE45FF0FFE73" box="[755,872,432,458]" pageId="9" pageNumber="58" type="paratype">Paratypes</typeStatus>
|
||
, Eastern
|
||
<collectingCountry id="F327DDA1AE4AE714AEEFFE45F82FFE73" box="[984,1096,432,458]" name="Malaysia" pageId="9" pageNumber="58">Malaysia</collectingCountry>
|
||
: 1 ɗ,
|
||
<date id="FF8EBBF1AE4AE714A9A6FE45F96FFE73" box="[1169,1288,432,458]" pageId="9" pageNumber="58" value="1990-11-09">9.XI.1990</date>
|
||
(meso– and meta thorax damaged) (HC); 1 Ψ, Sabah, Marak Parak, 6°19N, 116°43E, 25/
|
||
<date id="FF8EBBF1AE4AE714A913FE2DF8FAFE4B" box="[1060,1181,472,498]" pageId="9" pageNumber="58" value="1985-05-26">26.V.1985</date>
|
||
, Schmidt, Schmidt & Starr (
|
||
<collectionCode id="ED2105F4AE4AE714ADE8FDF5FD27FDA3" box="[223,320,512,538]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="58">NMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FDD2FFF5FCB3" blockId="9.[151,1437,152,1058]" pageId="9" pageNumber="58">
|
||
<emphasis id="B9444123AE4AE714ADF2FDD2FE42FDF8" bold="true" box="[197,549,551,577]" pageId="9" pageNumber="58">Other specimens examined.</emphasis>
|
||
Eastern
|
||
<collectingCountry id="F327DDA1AE4AE714AFA4FDDDFF64FDFB" box="[659,771,552,578]" name="Malaysia" pageId="9" pageNumber="58">Malaysia</collectingCountry>
|
||
: 1 Ψ,
|
||
<collectingCountry id="F327DDA1AE4AE714AE66FDDDFFDEFDFB" box="[849,953,552,578]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
, Kuching, Mt. Serapi, 1°33N, 110°25E,
|
||
<date id="FF8EBBF1AE4AE714ADA0FDA5FD63FDD3" box="[151,260,592,618]" pageId="9" pageNumber="58" value="1990-10-04">4.X.1990</date>
|
||
(meso– and meta thorax and abdomen damaged) (HC); 1 Ψ,
|
||
<collectingCountry id="F327DDA1AE4AE714A92AFDA5F8E2FDD3" box="[1053,1157,592,618]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
, Kuching, Mt. Serapi,
|
||
<date id="FF8EBBF1AE4AE714ADA0FD8DFD74FD2B" box="[151,275,632,658]" pageId="9" pageNumber="58" value="1990-10-15">15.X.1990</date>
|
||
(meso– and meta thorax and abdomen damaged) (HC); 1 Ψ,
|
||
<collectingCountry id="F327DDA1AE4AE714A917FD8DF8EFFD2B" box="[1056,1160,632,658]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
, Kuching, Mt. Serapi,
|
||
<date id="FF8EBBF1AE4AE714ADA0FD55FD74FD03" box="[151,275,672,698]" pageId="9" pageNumber="58" value="1990-10-22">22.X.1990</date>
|
||
(meso– and meta thorax and abdomen damaged) (HC); 1 Ψ,
|
||
<collectingCountry id="F327DDA1AE4AE714A917FD55F8EFFD03" box="[1056,1160,672,698]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
, Kuching, Mt. Serapi,
|
||
<date id="FF8EBBF1AE4AE714ADA0FD3DFD74FD5B" box="[151,275,712,738]" pageId="9" pageNumber="58" value="1990-10-23">23.X.1990</date>
|
||
(meso– and meta thorax and abdomen damaged) (HC); 1 Ψ,
|
||
<collectingCountry id="F327DDA1AE4AE714A917FD3DF8EFFD5B" box="[1056,1160,712,738]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
, Kuching, Mt. Serapi,
|
||
<date id="FF8EBBF1AE4AE714ADA0FD05FD6AFCB3" box="[151,269,752,778]" pageId="9" pageNumber="58" value="1990-11-05">5.XI.1990</date>
|
||
(meso– and metathorax and abdomen damaged) (HC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ACEBAAE4AE714ADF2FCE2FF72FC8B" box="[197,789,791,818]" pageId="9" pageNumber="58" type="distribution">
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FCE2FF72FC8B" blockId="9.[151,1437,152,1058]" box="[197,789,791,818]" pageId="9" pageNumber="58">
|
||
<emphasis id="B9444123AE4AE714ADF2FCE2FD01FC88" bold="true" box="[197,358,791,817]" pageId="9" pageNumber="58">Distribution.</emphasis>
|
||
Eastern
|
||
<collectingCountry id="F327DDA1AE4AE714ACFBFCEDFE50FC8B" box="[460,567,792,818]" name="Malaysia" pageId="9" pageNumber="58">Malaysia</collectingCountry>
|
||
(Sabah,
|
||
<collectingCountry id="F327DDA1AE4AE714AFA8FCEDFF6EFC8B" box="[671,777,792,818]" name="Malaysia" pageId="9" pageNumber="58">Sarawak</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ACEBAAE4AE714ADF2FCCAF91DFCE3" box="[197,1402,831,858]" pageId="9" pageNumber="58" type="etymology">
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FCCAF91DFCE3" blockId="9.[151,1437,152,1058]" box="[197,1402,831,858]" pageId="9" pageNumber="58">
|
||
<emphasis id="B9444123AE4AE714ADF2FCCAFD35FCE0" bold="true" box="[197,338,831,857]" pageId="9" pageNumber="58">Etymology.</emphasis>
|
||
The specific epithet ‘
|
||
<taxonomicName id="4C30E6B2AE4AE714AF65FCB5FED8FCE0" box="[594,703,832,857]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="tortuosus">
|
||
<emphasis id="B9444123AE4AE714AF65FCB5FED8FCE0" box="[594,703,832,857]" italics="true" pageId="9" pageNumber="58">tortuosus</emphasis>
|
||
</taxonomicName>
|
||
’ refers to the twisted male tenth tergum of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ACEBAAE4AE714ADF2FC92FE99FB9B" pageId="9" pageNumber="58" type="discussion">
|
||
<paragraph id="8B8F9D31AE4AE714ADF2FC92FE99FB9B" blockId="9.[151,1437,152,1058]" pageId="9" pageNumber="58">
|
||
<emphasis id="B9444123AE4AE714ADF2FC92FD59FC38" bold="true" box="[197,318,871,897]" pageId="9" pageNumber="58">Remarks.</emphasis>
|
||
Although a series of specimens from HC were seriously damaged, the specific identity can be determined by the vertexal and prothoracic markings as well as the wing patterns. Fortunately, the genitalia from the HC male was intact and the same as that of the
|
||
<typeStatus id="548B2393AE4AE714AE08FC4DFFCEFC6B" box="[831,937,952,978]" pageId="9" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
. It was designated as a
|
||
<typeStatus id="548B2393AE4AE714A9F5FC4DF94CFC6B" box="[1218,1323,952,978]" pageId="9" pageNumber="58" type="paratype">paratype</typeStatus>
|
||
. The vertexal markings are absent in the
|
||
<typeStatus id="548B2393AE4AE714AF21FC15FEE7FC43" box="[534,640,992,1018]" pageId="9" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
, which might constitute individual variation. For the comparison to the related species, see the remarks of
|
||
<taxonomicName id="4C30E6B2AE4AE714AF6DFBFDFE91FB98" box="[602,758,1032,1057]" class="Insecta" family="Corydalidae" genus="Protohermes" kingdom="Animalia" order="Megaloptera" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B9444123AE4AE714AF6DFBFDFE91FB98" box="[602,758,1032,1057]" italics="true" pageId="9" pageNumber="58">P. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |