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<document id="1478A05893AD0C3760D837B759162230" ID-DOI="10.11646/zootaxa.5529.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14022748" ID-ZooBank="716EDF67-C933-484D-911B-4585B11187A6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1730453336952" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2024" docId="0394B500FFD7FFB8FF322B6DD0CDFB70" docLanguage="en" docName="zootaxa.5529.3.3.pdf" docOrigin="Zootaxa 5529 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5529.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colossobolus olympiacus Wesener 2024, new combination" docType="treatment" docVersion="1" lastPageNumber="466" masterDocId="FFADCD78FFD4FFBDFFA52C0BD158FF99" masterDocTitle="Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="486" masterPageNumber="461" pageNumber="464" updateTime="1730714569688" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="997389381DD4E9658F05DB6C2369770B">Five new species and numerous locality records of Spirobolida millipedes from Madagascar (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:date id="2C0A7112C5679729E855BF8B002CE4CD">2024</mods:date>
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<subSubSection id="C327579DFFD7FFBEFF322B6DD23CF818" box="[151,868,1894,1921]" pageId="3" pageNumber="464" type="nomenclature">
<paragraph id="8B820416FFD7FFBEFF322B6DD23CF818" blockId="3.[151,868,1894,1957]" box="[151,868,1894,1921]" pageId="3" pageNumber="464">
<heading id="D0CAB37AFFD7FFBEFF322B6DD23CF818" bold="true" box="[151,868,1894,1921]" fontSize="11" level="1" pageId="3" pageNumber="464" reason="1">
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<taxonomicName id="4C3D7F95FFD7FFBEFF322B6DD3DEF819" authority="(Karsch, 1881)" authorityName="Wesener" authorityYear="2024" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[151,646,1894,1921]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="olympiacus" status="comb. nov.">
<emphasis id="B949D804FFD7FFBEFF322B6DD09FF818" bold="true" box="[151,455,1894,1921]" italics="true" pageId="3" pageNumber="464">Colossobolus olympiacus</emphasis>
(
<bibRefCitation id="EFAC79E7FFD7FFBEFE722B6DD326F819" author="Karsch, F." box="[471,638,1894,1920]" pageId="3" pageNumber="464" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
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<paragraph id="8B820416FFD7FFBEFF322B80D057F83C" blockId="3.[151,868,1894,1957]" box="[151,271,1931,1957]" pageId="3" pageNumber="464">
<figureCitation id="13061893FFD7FFBEFF322B80D057F83C" box="[151,271,1931,1957]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="3" pageNumber="464">Fig. 1A–H</figureCitation>
</paragraph>
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<paragraph id="8B820416FFD7FFBEFF322BC4D263F87E" blockId="3.[151,827,1999,2023]" box="[151,827,1999,2023]" pageId="3" pageNumber="464">
<treatmentCitationGroup id="AB2D2338FFD7FFBEFF322BC4D263F87E" box="[151,827,1999,2023]" pageId="3" pageNumber="464">
<treatmentCitation id="0A9C2207FFD7FFBEFF322BC4D36AF87E" author="Karsch, F." box="[151,562,1999,2023]" page="64" pageId="3" pageNumber="464" year="1881">
<taxonomicName id="4C3D7F95FFD7FFBEFF322BC4D36AF87E" ID-CoL="4Z7NL" authority="Karsch, 1881: 64" authorityName="Karsch" authorityPageNumber="64" authorityYear="1881" box="[151,562,1999,2023]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="olympiacus">
<emphasis id="B949D804FFD7FFBEFF322BC4D025F87E" box="[151,381,1999,2023]" italics="true" pageId="3" pageNumber="464">Spirobolus olympiacus</emphasis>
<bibRefCitation id="EFAC79E7FFD7FFBEFE212BC4D36AF87E" author="Karsch, F." box="[388,562,1999,2023]" pageId="3" pageNumber="464" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881: 64</bibRefCitation>
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(first description, female)
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</paragraph>
<paragraph id="8B820416FFD0FFB9FF322C9CD2F8FF36" blockId="4.[151,928,151,267]" box="[151,928,151,175]" pageId="4" pageNumber="465">
<taxonomicName id="4C3D7F95FFD0FFB9FF322C9CD351FF36" ID-CoL="4Z7NL" authorityName="Karsch" authorityYear="1881" box="[151,521,151,175]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="olympiacus" subGenus="Trigoniulus">
<emphasis id="B949D804FFD0FFB9FF322C9CD351FF36" box="[151,521,151,175]" italics="true" pageId="4" pageNumber="465">Spirobolus (Trigoniulus) olympiacus</emphasis>
</taxonomicName>
deSaussure &amp; Zehntner 1902: 129 (list)
</paragraph>
<paragraph id="8B820416FFD0FFB9FF322CBED3D9FF54" blockId="4.[151,928,151,267]" box="[151,641,181,205]" pageId="4" pageNumber="465">
<treatmentCitationGroup id="AB2D2338FFD0FFB9FF322CBED3D9FF54" box="[151,641,181,205]" pageId="4" pageNumber="465">
<treatmentCitation id="0A9C2207FFD0FFB9FF322CBED313FF54" author="Enghoff, H." box="[151,587,181,205]" page="624" pageId="4" pageNumber="465" year="2003">
<taxonomicName id="4C3D7F95FFD0FFB9FF322CBED313FF54" authority="Enghoff 2003: 624" authorityName="Enghoff" authorityPageNumber="624" authorityYear="2003" box="[151,587,181,205]" class="Diplopoda" family="Pachybolidae" genus="Trigoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="olympiacus">
<emphasis id="B949D804FFD0FFB9FF322CBED0DDFF54" box="[151,389,181,205]" italics="true" pageId="4" pageNumber="465">Trigoniulus olympiacus</emphasis>
<bibRefCitation id="EFAC79E7FFD0FFB9FE292CBED313FF54" author="Enghoff, H." box="[396,587,181,205]" pageId="4" pageNumber="465" pagination="617 - 627" refId="ref13366" refString="Enghoff, H. (2003) Diplopoda, Millipedes. In: Goodman, S. M. &amp; Benstead, J. P. (Eds.), The Natural History of Madagascar. University of Chicago Press, Chicago, Illinois, pp. 617 - 627." type="book chapter" year="2003">Enghoff 2003: 624</bibRefCitation>
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(list)
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</paragraph>
<paragraph id="8B820416FFD0FFB9FF322CDFD210FF75" blockId="4.[151,928,151,267]" box="[151,840,212,236]" pageId="4" pageNumber="465">
<treatmentCitationGroup id="AB2D2338FFD0FFB9FF322CDFD210FF75" box="[151,840,212,236]" pageId="4" pageNumber="465">
<taxonomicName id="4C3D7F95FFD0FFB9FF322CDFD0D1FF75" authorityName="Karsch" authorityYear="1881" box="[151,393,212,236]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="olympiaceus">
<emphasis id="B949D804FFD0FFB9FF322CDFD0D1FF75" box="[151,393,212,236]" italics="true" pageId="4" pageNumber="465">Spirobolus olympiaceus</emphasis>
</taxonomicName>
(sic!)
<treatmentCitation id="0A9C2207FFD0FFB9FE6E2CDFD38AFF75" author="Wesener T. &amp; Enghoff H. &amp; Sierwald P." box="[459,722,212,236]" page="120" pageId="4" pageNumber="465" year="2009">
<bibRefCitation id="EFAC79E7FFD0FFB9FE6E2CDFD38AFF75" author="Wesener T. &amp; Enghoff H. &amp; Sierwald P." box="[459,722,212,236]" pageId="4" pageNumber="465" pagination="1 - 128" refId="ref15399" refString="Wesener T., Enghoff H., Sierwald P. (2009 a) Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda). ZooKeys, 19, 1 - 128. https: // doi. org / 10.3897 / zookeys. 19.221" type="journal article" year="2009">
Wesener
<emphasis id="B949D804FFD0FFB9FD8D2CDED303FF75" box="[552,603,212,236]" italics="true" pageId="4" pageNumber="465">et al.</emphasis>
2009a: 120
</bibRefCitation>
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(comment)
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</paragraph>
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<treatmentCitationGroup id="AB2D2338FFD0FFB9FF322CF8D3AFFE92" box="[151,759,243,267]" pageId="4" pageNumber="465">
<treatmentCitation id="0A9C2207FFD0FFB9FF322CF8D39AFE92" author="Wesener, T. &amp; Enghoff, H." box="[151,706,243,267]" page="929" pageId="4" pageNumber="465" year="2022">
<taxonomicName id="4C3D7F95FFD0FFB9FF322CF8D39AFE92" authority="Wesener &amp; Enghoff 2022: 929" authorityName="Wesener &amp; Enghoff" authorityPageNumber="929" authorityYear="2022" box="[151,706,243,267]" class="Diplopoda" family="Pachybolidae" genus="Trigoniulus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="olympiacus">
<emphasis id="B949D804FFD0FFB9FF322CF8D0DDFE92" box="[151,389,243,267]" italics="true" pageId="4" pageNumber="465">Trigoniulus olympiacus</emphasis>
<bibRefCitation id="EFAC79E7FFD0FFB9FE292CF8D39AFE92" author="Wesener, T. &amp; Enghoff, H." box="[396,706,243,267]" pageId="4" pageNumber="465" pagination="918 - 933" refId="ref15196" refString="Wesener, T. &amp; Enghoff, H. (2022) Diplopoda, Millipedes, Menavetraka, Ankodavitra, Ankodiavitry, Marotanana, Sakolavitsy. In: Goodmann, S. M. (Ed.), The New Natural History of Madagascar, 1 (9), pp. 918 - 933. https: // doi. org / 10.2307 / j. ctv 2 ks 6 tbb. 118" type="book chapter" year="2022">Wesener &amp; Enghoff 2022: 929</bibRefCitation>
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(list)
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</paragraph>
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<subSubSection id="C327579DFFD0FFB9FF322D32D410F88D" pageId="4" pageNumber="465" type="materials_examined">
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<emphasis id="B949D804FFD0FFB9FF322D32D025FECA" bold="true" box="[151,381,313,339]" pageId="4" pageNumber="465">Material examined</emphasis>
.
<materialsCitation id="3B550E4BFFD0FFB9FE2A2D32D5CAFE02" collectingDate="2003-03-13" collectingDateMax="2003-03-16" collectingDateMin="2003-03-13" collectionCode="BLF" collectorName="D. Andriamalala &amp; Andriamalala, C &amp; Griswold, H &amp; D. Silva" country="Madagascar" county="Province" elevation="100" latitude="-13.415555" location="Parc National de Lokobe" longLatPrecision="21" longitude="48.3075" municipality="Nosy Be" pageId="4" pageNumber="465" specimenCode="BLF 7999" specimenCount="1" stateProvince="Antsiranana">
1 M,
<emphasis id="B949D804FFD0FFB9FE772D32D21DFECB" bold="true" box="[466,837,312,339]" pageId="4" pageNumber="465">
<specimenCode id="DB9BAC6DFFD0FFB9FE772D32D312FECA" box="[466,586,313,339]" collectionCode="BLF" pageId="4" pageNumber="465">BLF 7999</specimenCode>
(CASENT 9005459)
</emphasis>
.
<collectingCountry id="F32A4486FFD0FFB9FCF22D32D2B8FECA" box="[855,992,313,339]" name="Madagascar" pageId="4" pageNumber="465">Madagascar</collectingCountry>
,
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<collectingRegion id="49F9CAF4FFD0FFB9FBFE2D32D5BDFECA" box="[1115,1253,313,339]" country="Madagascar" name="Antsiranana" pageId="4" pageNumber="465">Antsiranana</collectingRegion>
,
<collectingMunicipality id="6BE69E6CFFD0FFB9FB512D32D404FECA" box="[1268,1372,313,339]" pageId="4" pageNumber="465">Nosy Be</collectingMunicipality>
,
<location id="8EE252CDFFD0FFB9FACF2D32D0DAFEEE" LSID="urn:lsid:plazi:treatment:0394B500FFD7FFB8FF322B6DD0CDFB70:8EE252CDFFD0FFB9FACF2D32D0DAFEEE" country="Madagascar" county="Province" latitude="-13.415555" longLatPrecision="21" longitude="48.3075" municipality="Nosy Be" name="Parc National de Lokobe" pageId="4" pageNumber="465" stateProvince="Antsiranana">Parc National de Lokobe</location>
,
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125° ESE Hellville
</locationDeviation>
, rainforest,
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,
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,
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, coll.
<collectorName id="26C861C0FFD0FFB9FA242D56D06AFE02" pageId="4" pageNumber="465">D. Andriamalala</collectorName>
, C.
<collectorName id="26C861C0FFD0FFB9FEC22D8AD0A1FE02" box="[359,505,385,411]" pageId="4" pageNumber="465">Griswold, H</collectorName>
. Ratsirarson &amp;
<collectorName id="26C861C0FFD0FFB9FD102D8AD240FE02" box="[693,792,385,411]" pageId="4" pageNumber="465">D. Silva</collectorName>
, hand collecting,
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</date>
</materialsCitation>
.
<materialsCitation id="3B550E4BFFD0FFB9FB042D8AD357FE7A" collectingDate="2013-02-21" collectionCode="BLF" collectorName="B. L. Fisher" country="Madagascar" elevation="520" latitude="-13.585" location="Mont Galoko" longLatPrecision="21" longitude="48.718056" municipality="Galoko" pageId="4" pageNumber="465" specimenCode="BLF30972" specimenCount="1" stateProvince="Antsiranana">
1 M, 2 F,
<emphasis id="B949D804FFD0FFB9FA852D8AD4C4FE02" bold="true" box="[1312,1436,385,411]" pageId="4" pageNumber="465">
<specimenCode id="DB9BAC6DFFD0FFB9FA852D8AD4C4FE02" box="[1312,1436,385,411]" collectionCode="BLF" pageId="4" pageNumber="465">BLF30972</specimenCode>
</emphasis>
(
<emphasis id="B949D804FFD0FFB9FF042DAFD0DFFE27" bold="true" box="[161,391,420,446]" pageId="4" pageNumber="465">CASENT 9068308)</emphasis>
,
<collectingRegion id="49F9CAF4FFD0FFB9FE302DAED347FE26" box="[405,543,421,447]" country="Madagascar" name="Antsiranana" pageId="4" pageNumber="465">Antsiranana</collectingRegion>
,
<collectingMunicipality id="6BE69E6CFFD0FFB9FD882DAED327FE26" box="[557,639,421,447]" pageId="4" pageNumber="465">Galoko</collectingMunicipality>
chain,
<location id="8EE252CDFFD0FFB9FD732DAED229FE26" LSID="urn:lsid:plazi:treatment:0394B500FFD7FFB8FF322B6DD0CDFB70:8EE252CDFFD0FFB9FD732DAED229FE26" box="[726,881,421,447]" country="Madagascar" latitude="-13.585" longLatPrecision="21" longitude="48.718056" municipality="Galoko" name="Mont Galoko" pageId="4" pageNumber="465" stateProvince="Antsiranana">Mont Galoko</location>
, rainforest,
<quantity id="4CC5A9F3FFD0FFB9FC5E2DAED51FFE26" box="[1019,1095,421,448]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="465" unit="m" value="520.0">
<elevation id="0010E325FFD0FFB9FC5E2DAED51FFE26" box="[1019,1095,421,448]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="465" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<geoCoordinate id="EE0962D1FFD0FFB9FBF02DAED5BEFE26" box="[1109,1254,421,448]" degrees="13" direction="south" minutes="35" orientation="latitude" pageId="4" pageNumber="465" precision="15" seconds="06" value="-13.585">13°35’06’’ S</geoCoordinate>
,
<geoCoordinate id="EE0962D1FFD0FFB9FB532DAED4CFFE26" box="[1270,1431,421,448]" degrees="048" direction="east" minutes="43" orientation="longitude" pageId="4" pageNumber="465" precision="15" seconds="05" value="48.718056">048°43’05’’ E</geoCoordinate>
, coll.
<collectorName id="26C861C0FFD0FFB9FF6A2DC2D016FE7A" box="[207,334,457,483]" pageId="4" pageNumber="465">B.L. Fisher</collectorName>
<emphasis id="B949D804FFD0FFB9FEF02DC1D0DEFE7A" box="[341,390,457,483]" italics="true" pageId="4" pageNumber="465">et al</emphasis>
.,
<date id="FF8322D6FFD0FFB9FE3E2DC2D357FE7A" box="[411,527,457,483]" pageId="4" pageNumber="465" value="2013-02-21">
<collectingDate id="EFC7DB3EFFD0FFB9FE3E2DC2D357FE7A" box="[411,527,457,483]" pageId="4" pageNumber="465" value="2013-02-21">21.ii.2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="DF42549EFFD0FFB9FF3228EBD3E4F981" ID-DOI="http://doi.org/10.5281/zenodo.14022750" ID-Zenodo-Dep="14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465" startId="4.[151,250,1248,1272]" targetBox="[417,1171,507,1224]" targetPageId="4" targetType="figure">
<paragraph id="8B820416FFD0FFB9FF3228EBD3E4F981" blockId="4.[151,1437,1248,1560]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FF3228EBD04DFB61" bold="true" box="[151,277,1248,1272]" pageId="4" pageNumber="465">FIGURE 1.</emphasis>
<taxonomicName id="4C3D7F95FFD0FFB9FEB828EBD39EFB61" authority="(Karsch, 1881)" authorityName="Wesener" authorityYear="2024" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[285,710,1248,1272]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="olympiacus" status="comb. nov.">
<emphasis id="B949D804FFD0FFB9FEB828EBD379FB61" box="[285,545,1248,1272]" italics="true" pageId="4" pageNumber="465">Colossobolus olympiacus</emphasis>
(
<bibRefCitation id="EFAC79E7FFD0FFB9FD9428EBD3E6FB61" author="Karsch, F." box="[561,702,1248,1272]" pageId="4" pageNumber="465" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B949D804FFD0FFB9FD7728EBD268FB61" bold="true" box="[722,816,1248,1272]" pageId="4" pageNumber="465">
<taxonomicNameLabel id="A27A657FFFD0FFB9FD7728EBD268FB61" box="[722,816,1248,1272]" pageId="4" pageNumber="465" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, topotype (
<emphasis id="B949D804FFD0FFB9FC0D28EBD537FB61" bold="true" box="[936,1135,1248,1272]" pageId="4" pageNumber="465">CASENT 9005459</emphasis>
).
<emphasis id="B949D804FFD0FFB9FB2328EBD5C0FB61" bold="true" box="[1158,1176,1248,1272]" pageId="4" pageNumber="465">A</emphasis>
. Multi-layer photograph, male anterior body, lateral view;
<emphasis id="B949D804FFD0FFB9FE5D290FD351FA85" bold="true" box="[504,521,1284,1308]" pageId="4" pageNumber="465">B</emphasis>
. Multi-layer photograph, male leg pairs 1–7, ventral view;
<emphasis id="B949D804FFD0FFB9FB26290FD5CDFA85" bold="true" box="[1155,1173,1284,1308]" pageId="4" pageNumber="465">C</emphasis>
. Multi-layer photograph, male posterior body end with telson;
<emphasis id="B949D804FFD0FFB9FDBD2923D369FAD9" bold="true" box="[536,561,1320,1344]" pageId="4" pageNumber="465">D.</emphasis>
Right anterior gonopod, anterior view;
<emphasis id="B949D804FFD0FFB9FC6E2923D284FAD9" bold="true" box="[971,988,1320,1344]" pageId="4" pageNumber="465">E</emphasis>
. Right anterior gonopod, posterior view;
<emphasis id="B949D804FFD0FFB9FA2C2923D4C4FAD9" bold="true" box="[1417,1436,1320,1344]" pageId="4" pageNumber="465">F.</emphasis>
Left posterior gonopod, anterior view;
<emphasis id="B949D804FFD0FFB9FD832947D319FAFD" bold="true" box="[550,577,1356,1380]" pageId="4" pageNumber="465">G.</emphasis>
Left posterior gonopod, posterior view;
<emphasis id="B949D804FFD0FFB9FC462947D2A5FAFD" bold="true" box="[995,1021,1356,1380]" pageId="4" pageNumber="465">H.</emphasis>
Male right midbody leg, posterior view.
<emphasis id="B949D804FFD0FFB9FF32297BD068FA11" bold="true" box="[151,304,1392,1416]" pageId="4" pageNumber="465">Abbreviations</emphasis>
: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number.
<emphasis id="B949D804FFD0FFB9FE542A0BD307F981" bold="true" box="[497,607,1536,1560]" pageId="4" pageNumber="465">Scale bars</emphasis>
= 1 mm.
</paragraph>
</caption>
<paragraph id="8B820416FFD0FFB9FF622A61D410F88D" blockId="4.[151,1437,1642,2029]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FF622A61D00EF91D" bold="true" box="[199,342,1642,1668]" pageId="4" pageNumber="465">Rediagnosis</emphasis>
.
<taxonomicName id="4C3D7F95FFD0FFB9FEC02A61D30DF91D" box="[357,597,1642,1668]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FEC02A61D0A4F91D" box="[357,508,1642,1668]" italics="true" pageId="4" pageNumber="465">Colossobolus</emphasis>
species
</taxonomicName>
with a finger-shaped coxite branch on the posterior gonopod (
<figureCitation id="13061893FFD0FFB9FABD2A61D4D7F91D" box="[1304,1423,1642,1668]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1F, G</figureCitation>
), like in
<taxonomicName id="4C3D7F95FFD0FFB9FF4C2A84D360F931" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[233,568,1678,1705]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="semicyclus">
<emphasis id="B949D804FFD0FFB9FF4C2A84D0DDF931" box="[233,389,1678,1705]" italics="true" pageId="4" pageNumber="465">C. semicyclus</emphasis>
Wesener, 2009
</taxonomicName>
,
<taxonomicName id="4C3D7F95FFD0FFB9FDE02A84D2EDF931" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[581,949,1678,1705]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="oblongopedus">
<emphasis id="B949D804FFD0FFB9FDE02A84D25DF931" box="[581,773,1678,1705]" italics="true" pageId="4" pageNumber="465">C. oblongopedus</emphasis>
Wesener, 2009
</taxonomicName>
and
<taxonomicName id="4C3D7F95FFD0FFB9FC4A2A84D46BF931" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1007,1331,1678,1705]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B949D804FFD0FFB9FC4A2A84D527F931" box="[1007,1151,1679,1705]" italics="true" pageId="4" pageNumber="465">C. giganteus</emphasis>
Wesener, 2009
</taxonomicName>
. Anterior gonopods lacking a retrorse process on the mesal margin of the telopodite (
<figureCitation id="13061893FFD0FFB9FC562AB9D537F955" box="[1011,1135,1714,1740]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1D, E</figureCitation>
), like in
<taxonomicName id="4C3D7F95FFD0FFB9FB732AB8D4CDF955" authorityName="Wesener" authorityYear="2009" box="[1238,1429,1714,1741]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="oblongopedus">
<emphasis id="B949D804FFD0FFB9FB732AB8D4CDF955" box="[1238,1429,1714,1741]" italics="true" pageId="4" pageNumber="465">C. oblongopedus</emphasis>
</taxonomicName>
. Posterior gonopods most similar to those of
<taxonomicName id="4C3D7F95FFD0FFB9FDD12ADCD277F969" authorityName="Wesener" authorityYear="2009" box="[628,815,1750,1777]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="oblongopedus">
<emphasis id="B949D804FFD0FFB9FDD12ADCD277F969" box="[628,815,1750,1777]" italics="true" pageId="4" pageNumber="465">C. oblongopedus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D7F95FFD0FFB9FCC42ADCD2B4F969" authorityName="Wesener" authorityYear="2009" box="[865,1004,1751,1777]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B949D804FFD0FFB9FCC42ADCD2B4F969" box="[865,1004,1751,1777]" italics="true" pageId="4" pageNumber="465">C. giganteus</emphasis>
</taxonomicName>
, but differing in the narrowly elongated tip of the lateral branch, as well as the apical part of the mesal branch. Male legs with tarsal pads (
<figureCitation id="13061893FFD0FFB9FB472AF1D463F88D" box="[1250,1339,1786,1812]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD0FFB8FF622B15D306FC3C" lastPageId="5" lastPageNumber="466" pageId="4" pageNumber="465" type="description">
<paragraph id="8B820416FFD0FFB9FF622B15D23BF8A1" blockId="4.[151,1437,1642,2029]" box="[199,867,1822,1849]" pageId="4" pageNumber="465">
<emphasis id="B949D804FFD0FFB9FF622B15D035F8A1" bold="true" box="[199,365,1822,1848]" pageId="4" pageNumber="465">Redescription</emphasis>
(based on topotypic material from Nosy Be)
</paragraph>
<paragraph id="8B820416FFD0FFB9FF622B48D5B3F8C5" blockId="4.[151,1437,1642,2029]" box="[199,1259,1858,1885]" pageId="4" pageNumber="465">
Measurements: 56+0 body rings plus telson.
<quantity id="4CC5A9F3FFD0FFB9FD1A2B49D244F8C4" box="[703,796,1858,1885]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="4" pageNumber="465" unit="mm" value="178.0">178 mm</quantity>
long,
<quantity id="4CC5A9F3FFD0FFB9FCC72B49D28CF8C4" box="[866,980,1858,1885]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" pageId="4" pageNumber="465" unit="mm" value="12.15">12.15 mm</quantity>
wide (midbody, widest).
</paragraph>
<paragraph id="8B820416FFD0FFB9FF622B6DD29AF83D" blockId="4.[151,1437,1642,2029]" pageId="4" pageNumber="465">
Colour: mesozonite orange-grey, metazonite black, telson blood red. Ozopores black, head, antenna, mouthparts red. Collum at margins black, surface red (
<figureCitation id="13061893FFD0FFB9FDD02B81D396F83D" box="[629,718,1930,1956]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1A</figureCitation>
). Legs red to greyish.
</paragraph>
<paragraph id="8B820416FFD0FFB8FF622BA5D3DCFF4C" blockId="4.[151,1437,1642,2029]" lastBlockId="5.[151,1437,151,1257]" lastPageId="5" lastPageNumber="466" pageId="4" pageNumber="465">
Head with 35/36 black ommatidia (
<figureCitation id="13061893FFD0FFB9FDF72BA5D3F3F851" box="[594,683,1966,1992]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1A</figureCitation>
). Vertical suture not going through whole head, head with the usual 2+2 setiferous foveolae (
<figureCitation id="13061893FFD0FFB9FE122BD9D349F875" box="[439,529,2002,2028]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="4" pageNumber="465">Fig. 1B</figureCitation>
). Gnathochilarium typical for the order, lamellae linguales each with 1+1 seta behind one another, stipes each with three setae (
<figureCitation id="13061893FFD1FFB8FD762C9CD275FF28" box="[723,813,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1B</figureCitation>
). Mandible basal lobe not projecting. Antennae short, protruding back to third body ring (
<figureCitation id="13061893FFD1FFB8FD812CB0D323FF4C" box="[548,635,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1A</figureCitation>
)
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622CD4D545FF60" blockId="5.[151,1437,151,1257]" box="[199,1053,223,249]" pageId="5" pageNumber="466">Ozopore starting at ring 6, located on suture between meso- and metazonite.</paragraph>
<paragraph id="8B820416FFD1FFB8FF622D08D478FEFC" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Legs in male on coxae 3 and 4 with short conical process (
<figureCitation id="13061893FFD1FFB8FCD62D08D294FE84" box="[883,972,259,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1B</figureCitation>
), coxae 5–7 normal, tarsal pads present, starting at leg 3 (
<figureCitation id="13061893FFD1FFB8FEF62D2CD0F5FED8" box="[339,429,295,321]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1G</figureCitation>
). Legs 8+ with femur cylindrical, as long as tarsus, podomere length 1&lt;2&lt;3&gt;4=5&lt;6. Each podomere apically with one or two spines, tarsus with wide claw and single short apical spine (
<figureCitation id="13061893FFD1FFB8FB652D40D44FFEFC" box="[1216,1303,331,357]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1H</figureCitation>
)
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622D64D5DFFE34" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Telson: preanal scale very slightly projecting in well-rounded process, not projecting beyond anal valves (
<figureCitation id="13061893FFD1FFB8FACA2D64D1E2FE34" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1C</figureCitation>
). Anal valves with well-developed lips and large groove. Subanal scale inconspicuous.
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622DBCD0F1FDA4" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Anterior gonopod sternite elongated into a wide, broadly rounded-triangular lobe (
<figureCitation id="13061893FFD1FFB8FBC82DBCD59FFE48" box="[1133,1223,439,465]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1D</figureCitation>
). Mesal process of coxite weakly developed, protruding into a short, wide lobe with a rounded tip (
<figureCitation id="13061893FFD1FFB8FB802DD0D527FE6C" box="[1061,1151,475,501]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1D</figureCitation>
). Telopodite on posterior side basally with a circular groove (
<figureCitation id="13061893FFD1FFB8FD912DF4D3D4FD80" box="[564,652,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1E</figureCitation>
). Telopodite process long and well-rounded (
<figureCitation id="13061893FFD1FFB8FB312DF4D5B3FD80" box="[1172,1259,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1E</figureCitation>
), mesal margin laterally sharp (
<figureCitation id="13061893FFD1FFB8FEE32E28D0C3FDA4" box="[326,411,547,573]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622E4CD224FCA0" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
Posterior gonopods telopodites positioned parallel to one another. Coxite relatively short, wide, stout, less than half as long than lateral branch of telopodite (
<figureCitation id="13061893FFD1FFB8FD462E60D205FD1C" box="[739,861,619,645]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F, G</figureCitation>
). Latter slightly curved, basally wide, at midpoint tapering, tip again wide and swollen (
<figureCitation id="13061893FFD1FFB8FD9B2E84D3CBFD30" box="[574,659,655,681]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F</figureCitation>
). Central membranous area developed as a short, wide, well-rounded lobe (
<figureCitation id="13061893FFD1FFB8FF7D2EB8D076FD54" box="[216,302,691,717]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F</figureCitation>
). Mesal main branch short, basal part running parallel to lateral branch. Apical part of main branch slender, extending towards and touching lateral branch (
<figureCitation id="13061893FFD1FFB8FCA42EDCD201FD68" box="[769,857,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1G</figureCitation>
). Mesal margin of main branch with a well-rounded, overlapping membranous fold (
<figureCitation id="13061893FFD1FFB8FE592EF0D309FC8C" box="[508,593,763,789]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F</figureCitation>
). Main branch laterally with a small, irregular-shaped membrane (
<figureCitation id="13061893FFD1FFB8FAE12EF0D1F7FCA0" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Fig. 1F, G</figureCitation>
). Lateral branch apically strongly tapered, slender (
<figureCitation id="13061893FFD1FFB8FD4A2F14D228FCA0" box="[751,880,799,825]" captionStart="FIGURE 1" captionStartId="4.[151,250,1248,1272]" captionTargetBox="[417,1171,507,1224]" captionTargetId="figure-122@4.[416,1171,507,1224]" captionTargetPageId="4" captionText="FIGURE 1. Colossobolus olympiacus (Karsch, 1881), n. comb., topotype (CASENT 9005459). A. Multi-layer photograph, male anterior body, lateral view; B. Multi-layer photograph, male leg pairs 1–7, ventral view; C. Multi-layer photograph, male posterior body end with telson; D. Right anterior gonopod, anterior view; E. Right anterior gonopod, posterior view; F. Left posterior gonopod, anterior view; G. Left posterior gonopod, posterior view; H. Male right midbody leg, posterior view. Abbreviations: Apo = apodeme; av = paraprocts; bb = basal branch of posterior gonopod; Co = collum (ring 1); Cx = coxite; Fe = femur; fs-p = finger-shaped process; Gn = gnathochilarium; Gr = groove; ma = membranous area on posterior gonopod; Md = basal joints of mandible; oz = ozopore; Po = postfemur; Pre = epiproct; sf = setiferous foveolae; St = sternite; sub = hypoproct; T = telopodite; T-Lbr = lateral branch of telopodite; T-mbr = main branch of telopodite; Ta = tarsus; ta-p = tarsal pad; Ti = tibia; numbers refer to leg pair number. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14022750" httpUri="https://zenodo.org/record/14022750/files/figure.png" pageId="5" pageNumber="466">Figs 1 F, G</figureCitation>
).
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622F49D306FC3C" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622F49D080FCC5" bold="true" box="[199,472,834,860]" pageId="5" pageNumber="466">Intraspecific variation.</emphasis>
Specimens from the Galoko Mountain Chain have only 52–54 body rings and are with a length of
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, a width of
<quantity id="4CC5A9F3FFD1FFB8FE512F6DD33CFC18" box="[500,612,870,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.135" pageId="5" pageNumber="466" unit="mm" value="11.35">11.35 mm</quantity>
in the male and a length of
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length with a width of
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in the female shorter than those from Nosy Be.
</paragraph>
</subSubSection>
<subSubSection id="C327579DFFD1FFB8FF622FA4D0CDFB70" pageId="5" pageNumber="466" type="discussion">
<paragraph id="8B820416FFD1FFB8FF622FA4D419FC50" blockId="5.[151,1437,151,1257]" box="[199,1345,943,969]" pageId="5" pageNumber="466">
<emphasis id="B949D804FFD1FFB8FF622FA4D063FC50" bold="true" box="[199,315,943,969]" pageId="5" pageNumber="466">Remarks.</emphasis>
At both localities found in sympatry with
<taxonomicName id="4C3D7F95FFD1FFB8FCB02FA4D464FC50" authority="(Pocock, 1893)" baseAuthorityName="Pocock" baseAuthorityYear="1893" box="[789,1340,943,969]" class="Diplopoda" family="Pachybolidae" genus="Aphistogoniulus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="erythrocephalus">
<emphasis id="B949D804FFD1FFB8FCB02FA4D5DFFC50" box="[789,1159,943,969]" italics="true" pageId="5" pageNumber="466">Aphistogoniulus erythrocephalus</emphasis>
(
<bibRefCitation id="EFAC79E7FFD1FFB8FB332FA4D46CFC50" author="Pocock, R. I." box="[1174,1332,943,969]" pageId="5" pageNumber="466" pagination="248 - 254" refId="ref14311" refString="Pocock, R. I. (1893) Upon the identity of some of the types of Diplopoda contained in the collection of the British Museum, together with description of some new species of exotic Julidae. Annals and Magazine of Natural History, Series 6, 11, 248 - 254. https: // doi. org / 10.1080 / 00222939308677511" type="journal article" year="1893">Pocock, 1893</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B820416FFD1FFB8FF622FD8D0CDFB70" blockId="5.[151,1437,151,1257]" pageId="5" pageNumber="466">
<taxonomicName id="4C3D7F95FFD1FFB8FF622FD8D335FC74" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[199,621,979,1005]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="olympiacus">
<emphasis id="B949D804FFD1FFB8FF622FD8D092FC74" box="[199,458,979,1005]" italics="true" pageId="5" pageNumber="466">Spirobolus olympiacus</emphasis>
<bibRefCitation id="EFAC79E7FFD1FFB8FE712FD8D335FC74" author="Karsch, F." box="[468,621,979,1005]" pageId="5" pageNumber="466" pagination="1 - 79" refId="ref13686" refString="Karsch, F. (1881) Neue Juliden des Berliner Museums, als Prodromus einer Juliden-Monographie. Zeitschrift fur die gesammten Naturwissenschaftem, 54, 1 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
</taxonomicName>
was one of the few remaining
<emphasis id="B949D804FFD1FFB8FC442FD8D527FC74" box="[993,1151,979,1005]" italics="true" pageId="5" pageNumber="466">nomina dubia</emphasis>
of the
<taxonomicName id="4C3D7F95FFD1FFB8FB7D2FD8D403FC74" authorityName="Bollman" authorityYear="1893" box="[1240,1371,979,1005]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
from
<collectingCountry id="F32A4486FFD1FFB8FF322FFCD046FB88" box="[151,286,1015,1041]" name="Madagascar" pageId="5" pageNumber="466">Madagascar</collectingCountry>
(
<bibRefCitation id="EFAC79E7FFD1FFB8FE8B2FFCD37BFB88" author="Wesener T. &amp; Enghoff H. &amp; Sierwald P." box="[302,547,1015,1041]" pageId="5" pageNumber="466" pagination="1 - 128" refId="ref15399" refString="Wesener T., Enghoff H., Sierwald P. (2009 a) Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda). ZooKeys, 19, 1 - 128. https: // doi. org / 10.3897 / zookeys. 19.221" type="journal article" year="2009">
Wesener
<emphasis id="B949D804FFD1FFB8FE3D2FFCD08AFB88" box="[408,466,1015,1041]" italics="true" pageId="5" pageNumber="466">et al.</emphasis>
2009a
</bibRefCitation>
). The short first description is the only known record, based on a female from Nosy Be, and mentions few characters aside from the large size (
<quantity id="4CC5A9F3FFD1FFB8FC3F2811D2A5FBAC" box="[922,1021,1050,1077]" metricMagnitude="-1" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="466" unit="mm" value="215.0">215 mm</quantity>
) and the presence of 57 rings. The
<typeStatus id="5486BAB4FFD1FFB8FF322834D1A1FBC0" box="[151,249,1087,1113]" pageId="5" pageNumber="466" type="holotype">holotype</typeStatus>
(and only known specimen) seems unfortunately lost. It is no longer in the collections of the museum in Berlin (Mark Judson, pers. comm.), and it is also one of the few not listed in the first Berlin Myriapoda type catalogues, neither under
<taxonomicName id="4C3D7F95FFD1FFB8FE11288CD36FFB38" authorityName="Bollman" authorityYear="1893" box="[436,567,1159,1185]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
nor
<taxonomicName id="4C3D7F95FFD1FFB8FDCD288CD25DFB38" authorityName="Brandt" authorityYear="1833" box="[616,773,1159,1185]" class="Diplopoda" kingdom="Animalia" order="Spirostreptida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
(
<bibRefCitation id="EFAC79E7FFD1FFB8FCB6288CD54EFB38" author="Moritz, M. &amp; Fischer, S. C." box="[787,1046,1159,1185]" pageId="5" pageNumber="466" pagination="323 - 375" refId="ref13815" refString="Moritz, M. &amp; Fischer, S. C. (1974) Typen der Myriapoden-Sammlung des Zoologischen Museums Berlin. I. Diplopoda. Teil 2: Craspedosomatida, Stemmiulida, Spirostreptida. Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 50 (1), 323 - 375. https: // doi. org / 10.1002 / mmnz. 19750510206" type="journal article" year="1974">Moritz &amp; Fischer 1974</bibRefCitation>
,
<bibRefCitation id="EFAC79E7FFD1FFB8FB85288DD504FB38" author="Moritz, M. &amp; Fischer, S. C." box="[1056,1116,1158,1185]" pageId="5" pageNumber="466" pagination="199 - 255" refId="ref13890" refString="Moritz, M. &amp; Fischer, S. C. (1975) Die Typen der Myriapoden-Sammlung des Zoologischen Museums Berlin. I. Diplopoda. Teil 3: Julida, Spirobolida. Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, Berlin, 51 (2), 199 - 255. https: // doi. org / 10.1002 / mmnz. 19750510206" type="journal article" year="1975">1975</bibRefCitation>
). The discovery of a massive
<taxonomicName id="4C3D7F95FFD1FFB8FF3228A0D042FB5C" authorityName="Bollman" authorityYear="1893" box="[151,282,1195,1221]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="466" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
from Nosy Be (albeit “only”
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in length) with 56 rings plus telson allows here a redescription of this enigmatic species.
</paragraph>
</subSubSection>
</treatment>
</document>
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