treatments-xml/data/F7/68/69/F76869577D01FFC43595FC7D884A6FBA.xml

<document id="CC30868D426CA9DA38A27A32EE40BD75" ID-CLB-Dataset="30139" ID-DOI="10.11646/zootaxa.4415.3.7" ID-GBIF-Dataset="d059e3a0-4309-42fd-8ee7-5d6d92aa1d5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242209" ID-ZooBank="8E3AF386-3A5A-41CD-956A-48055DD82B89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525600796046" checkinUser="plazi" docAuthor="Olthof, Gabriël, Becking, Leontine E. &amp; Fransen, Charles H. J. M." docDate="2018" docId="F76869577D01FFC43595FC7D884A6FBA" docLanguage="en" docName="zootaxa.4415.3.7.pdf" docOrigin="Zootaxa 4415 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocoutierea acutidorsata Olthof &amp; Becking &amp; Fransen 2018, sp. nov." docType="treatment" docVersion="8" lastPageNumber="544" masterDocId="0B51112F7D04FFC83502FFD4896B6F6F" masterDocTitle="On a collection of deep-water shrimp (Crustacea, Decapoda) from the Dutch Caribbean, with the description of a new species of Pseudocoutierea" masterLastPageNumber="548" masterPageNumber="533" pageNumber="538" updateTime="1698541956158" updateUser="ExternalLinkService">
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<mods:title id="E479A4168CCF728FD0785041A48C9397">On a collection of deep-water shrimp (Crustacea, Decapoda) from the Dutch Caribbean, with the description of a new species of Pseudocoutierea</mods:title>
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<mods:namePart id="A8CB019DA6EFC86A065DB51F0BD71A0E">Olthof, Gabriël</mods:namePart>
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<mods:namePart id="F31B51BF5792FE74070AABD4B660F6EC">Becking, Leontine E.</mods:namePart>
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<treatment id="F76869577D01FFC43595FC7D884A6FBA" ID-DOI="http://doi.org/10.5281/zenodo.5971376" ID-GBIF-Taxon="143811468" ID-Zenodo-Dep="5971376" LSID="urn:lsid:plazi:treatment:F76869577D01FFC43595FC7D884A6FBA" httpUri="http://treatment.plazi.org/id/F76869577D01FFC43595FC7D884A6FBA" lastPageId="12" lastPageNumber="544" pageId="5" pageNumber="538">
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<paragraph id="7F7ED8417D01FFCD3595FC7D8B0A6CAC" blockId="5.[151,609,937,997]" box="[151,609,937,963]" pageId="5" pageNumber="538">
<heading id="24366F2D7D01FFCD3595FC7D8B0A6CAC" bold="true" box="[151,609,937,963]" fontSize="11" level="1" pageId="5" pageNumber="538" reason="1">
<emphasis id="4DB504537D01FFCD3595FC7D8B0A6CAC" bold="true" box="[151,609,937,963]" pageId="5" pageNumber="538">
<taxonomicName id="B8C1A3C27D01FFCD3595FC7D88906CAC" ID-CoL="e5006ec6-6193-4584-9d15-3d0a694bc549" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[151,507,937,963]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D01FFCD3595FC7D88906CAC" bold="true" box="[151,507,937,963]" italics="true" pageId="5" pageNumber="538">Pseudocoutierea acutidorsata</emphasis>
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</emphasis>
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(
<figureCitation id="E7FAC4C47D01FFCD359DFC1888626C8A" box="[159,265,972,997]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE5" captionStart-5="FIGURE 6. 50" captionStartId-0="6.[151,250,1913,1935]" captionStartId-1="7.[151,250,1514,1536]" captionStartId-2="10.[151,250,1914,1936]" captionStartId-3="11.[151,250,1425,1447]" captionStartId-4="12.[151,250,923,945]" captionStartId-5="12.[151,250,1765,1787]" captionTargetBox-0="[180,1396,657,1855]" captionTargetBox-1="[164,1427,210,1468]" captionTargetBox-2="[238,1384,214,1885]" captionTargetBox-3="[305,1275,201,1383]" captionTargetBox-4="[196,1417,286,867]" captionTargetBox-5="[394,1212,1162,1694]" captionTargetId-0="figure@6.[151,1436,645,1879]" captionTargetId-1="figure@7.[151,1436,193,1493]" captionTargetId-2="figure@10.[193,1394,194,1895]" captionTargetId-3="figure@11.[296,1290,193,1403]" captionTargetId-4="figure@12.[151,1436,269,885]" captionTargetId-5="figure@12.[340,1247,1134,1734]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="12" captionText-0="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." captionText-1="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." captionText-2="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." captionText-3="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." captionText-4="FIGURE5. Pseudocoutierea acutidorsata sp. nov. A, right second pleopod, plumosehairs on blades omitted,female paratype, RMNH.CRUS.D.57273; B, variation in epigastric spine female paratypes RMNH.CRUS.D.57273; B1, ovigerous female pocl. 2.25mm; B2, ovigerous female pocl. 2.1mm; B3, female pocl. 1.75mm; B4, ovigerous female pocl. 2.75mm. Pseudocoutierea conchae Criales, 1981, male holotype, ZMA DE 103.237. Pleonal segments 3 to 5. C, pleura on right side in lateral view; D, pleura on left side in lateral view.Scale bar A, B = 1mm; C, D = 1.5mm." captionText-5="FIGURE 6. 50% majority rule consensus tree of Pseudocoutierea species; outgroup Coutierea agassizi (Coutière, 1901); tree length = 28, Consistency Index (CI) = 0.6429." httpUri-0="https://zenodo.org/record/1242211/files/figure.png" httpUri-1="https://zenodo.org/record/1242213/files/figure.png" httpUri-2="https://zenodo.org/record/1242215/files/figure.png" httpUri-3="https://zenodo.org/record/1242217/files/figure.png" httpUri-4="https://zenodo.org/record/1242219/files/figure.png" httpUri-5="https://zenodo.org/record/1242221/files/figure.png" pageId="5" pageNumber="538">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="7F7ED8417D01FFCD3595FBC088816B8F" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">
<emphasis id="4DB504537D01FFCD3595FBC088C96B42" bold="true" box="[151,418,1044,1069]" pageId="5" pageNumber="538">Material examined.</emphasis>
<collectionCode id="19D040847D01FFCD34CEFBC08B4D6B42" box="[460,550,1044,1069]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="538">RMNH</collectionCode>
. 
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.D.57272: 1 ovigorous female 
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<emphasis id="4DB504537D01FFCD314CFBC08DDF6B42" bold="true" box="[1102,1204,1044,1069]" pageId="5" pageNumber="538">holotype</emphasis>
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, pocl. 2.0mm; 
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. 
<collectionCode id="19D040847D01FFCD35F7FBE388296B3F" box="[245,322,1079,1104]" pageId="5" pageNumber="538">CRUS</collectionCode>
.D.57273: 5 ovigerous female 
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<emphasis id="4DB504537D01FFCD378CFBE38A6E6B3F" bold="true" box="[654,773,1079,1104]" pageId="5" pageNumber="538">paratypes</emphasis>
</typeStatus>
(pocl. 
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, eggs 
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in width and 
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in length), 2 small non-ovigerous female 
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<emphasis id="4DB504537D01FFCD3760FB888BA66B1A" bold="true" box="[610,717,1116,1141]" pageId="5" pageNumber="538">paratype</emphasis>
</typeStatus>
s (pocl. 1.75 and 
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): stn 
<collectionCode id="19D040847D01FFCD314BFB888DE26B1B" box="[1097,1161,1116,1140]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12673" name="Bolton Museum" pageId="5" pageNumber="538">BON</collectionCode>
3, 
<collectingCountry id="07D698D17D01FFCD31A4FB888C6A6B1A" box="[1190,1281,1116,1141]" name="Bonaire, Sint Eustatius and Saba" pageId="5" pageNumber="538">Bonaire</collectingCountry>
, 
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, 
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, depth 
<quantity id="B83975A47D01FFCD3482FB5488A86BF7" box="[384,451,1152,1176]" metricMagnitude="2" metricUnit="m" metricValue="1.38" pageId="5" pageNumber="538" unit="m" value="138.0">138m</quantity>
, 
<date id="0B7FFE817D01FFCD34CCFB548B546BF7" box="[462,575,1152,1176]" pageId="5" pageNumber="538" value="2013-05-31">31.v.2013</date>
, dive with submersible Curasub, from octocoral 
<taxonomicName id="B8C1A3C27D01FFCD3166FB5488AB6BD2" authority="(Milne Edwards &amp; Haime, 1857)" baseAuthorityName="Milne Edwards &amp; Haime" baseAuthorityYear="1857" class="Anthozoa" family="Primnoidae" genus="Callogorgia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="538" phylum="Cnidaria" rank="species" species="gracilis">
<emphasis id="4DB504537D01FFCD3166FB548C236BF7" box="[1124,1352,1152,1176]" italics="true" pageId="5" pageNumber="538">Callogorgia gracilis</emphasis>
(Milne Edwards &amp; Haime, 1857)
</taxonomicName>
identified by Bastian T. Reijnen (
<date id="0B7FFE817D01FFCD364FFB708AA26BD2" box="[845,969,1188,1213]" pageId="5" pageNumber="538" value="2013-06-31">31.vi.2013</date>
), collected by L.E. Becking and H.W.G. Meesters, sample nr BDR041.
</paragraph>
<paragraph id="7F7ED8417D01FFCD35C5FB388A1E6A1F" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">
<emphasis id="4DB504537D01FFCD35C5FB3888B66A6A" bold="true" box="[199,477,1260,1285]" pageId="5" pageNumber="538">Comparative material.</emphasis>
<taxonomicName id="B8C1A3C27D01FFCD34EBFB388AAC6A6B" authority="Chace, 1972" authorityName="Chace" authorityYear="1972" box="[489,967,1260,1285]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="antillensis">
<emphasis id="4DB504537D01FFCD34EBFB388A4F6A6B" box="[489,804,1260,1284]" italics="true" pageId="5" pageNumber="538">Pseudocoutierea antillensis</emphasis>
<bibRefCitation id="1B50A5B07D01FFCD3633FB388AAC6A6B" box="[817,967,1260,1285]" pageId="5" pageNumber="538" refId="ref9033" refString="Chace, F. A. (1972) The shrimps of the Smithsonian-Bredin Caribbean Expeditions with a summary of the West Indian shallowwater species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. https: // doi. org / 10.5479 / si. 00810282.98" type="journal article">Chace, 1972</bibRefCitation>
</taxonomicName>
: RMNH. 
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<collectionCode id="19D040847D01FFCD3137FB388DE96A6A" box="[1077,1154,1260,1285]" pageId="5" pageNumber="538">CRUS</collectionCode>
.D.51635, 
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, 
<collectingCountry id="07D698D17D01FFCD3595FAC4899C6A47" box="[151,247,1296,1320]" name="Curaçao" pageId="5" pageNumber="538">Curaçao</collectingCountry>
, 
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, 
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, 
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, depth: 
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</elevation>
, 
<collectingDate id="1B3B07697D01FFCD3696FADB8D656A47" box="[916,1038,1295,1320]" pageId="5" pageNumber="538" value="2005-06-18">
<date id="0B7FFE817D01FFCD3696FADB8D656A47" box="[916,1038,1295,1320]" pageId="5" pageNumber="538" value="2005-06-18">18.vi.2005</date>
</collectingDate>
, from 
<taxonomicName id="B8C1A3C27D01FFCD316AFAC488376A23" authority="(Houttuyn, 1772)" baseAuthorityName="Houttuyn" baseAuthorityYear="1772" class="Anthozoa" family="Plexauridae" genus="Pseudoplexaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="538" phylum="Cnidaria" rank="species" species="flagellosa">
<emphasis id="4DB504537D01FFCD316AFAC48CF06A47" box="[1128,1435,1296,1320]" italics="true" pageId="5" pageNumber="538">Pseudoplexaura flagellosa</emphasis>
(Houttuyn, 1772)
</taxonomicName>
, collected by 
<collectorName id="D234BD977D01FFCD34FAFAE18B1C6A22" box="[504,631,1332,1357]" pageId="5" pageNumber="538">N. Snijders</collectorName>
G.127
</materialsCitation>
. 
<materialsCitation id="CFA9D21C7D01FFCD37CDFAE08A1E6A1F" ID-GBIF-Occurrence="1841141218" collectingDate="1975-11-14" collectionCode="ZMA" collectorName="Bayer" country="Curaçao" elevation="30" location="Curacao" pageId="5" pageNumber="538" specimenCode="DE 103.525" specimenCount="8">
<collectionCode id="19D040847D01FFCD37CDFAE08A7A6A23" box="[719,785,1332,1356]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="5" pageNumber="538">ZMA</collectionCode>
: 
<specimenCode id="2F67703A7D01FFCD361EFAE18ACC6A22" box="[796,935,1332,1357]" collectionCode="ZMA" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="5" pageNumber="538">DE 103.525</specimenCode>
, 
<specimenCount id="69C713C87D01FFCD36B0FAE08D566A22" box="[946,1085,1332,1357]" pageId="5" pageNumber="538" type="generic">8 specimens</specimenCount>
from 
<collectingCountry id="07D698D17D01FFCD3180FAE08D896A23" box="[1154,1250,1332,1356]" name="Curaçao" pageId="5" pageNumber="538">Curaçao</collectingCountry>
, from a depth of around 
<elevation id="F4EC3F727D01FFCD35EEFA8C88746A1F" box="[236,287,1368,1392]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="538" unit="m" value="30.0">
<quantity id="B83975A47D01FFCD35EEFA8C88746A1F" box="[236,287,1368,1392]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="538" unit="m" value="30.0">30m</quantity>
</elevation>
, 
<collectingDate id="1B3B07697D01FFCD3429FA8388CE6A1F" box="[299,421,1367,1392]" pageId="5" pageNumber="538" value="1975-11-14">
<date id="0B7FFE817D01FFCD3429FA8388CE6A1F" box="[299,421,1367,1392]" pageId="5" pageNumber="538" value="1975-11-14">14.xi.1975</date>
</collectingDate>
, from 
<taxonomicName id="B8C1A3C27D01FFCD34ECFA8C8B216A00" baseAuthorityName="Ellis &amp; Solander" baseAuthorityYear="1786" box="[494,586,1368,1391]" class="Anthozoa" family="Plexauridae" genus="Eunicea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="538" phylum="Cnidaria" rank="genus">
<emphasis id="4DB504537D01FFCD34ECFA8C8B216A00" box="[494,586,1368,1391]" italics="true" pageId="5" pageNumber="538">Eunicea</emphasis>
</taxonomicName>
cf. 
<emphasis id="4DB504537D01FFCD3777FA8C8BB46A1F" box="[629,735,1368,1392]" italics="true" pageId="5" pageNumber="538">clavigera</emphasis>
<collectorName id="D234BD977D01FFCD37E4FA8C8A406A1F" box="[742,811,1368,1392]" pageId="5" pageNumber="538">Bayer</collectorName>
, 1961.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="37DB8BCA7D01FFCD35C5FAA888A569B7" pageId="5" pageNumber="538" type="reference_group">
<paragraph id="7F7ED8417D01FFCD35C5FAA88AF26AD7" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">
<taxonomicName id="B8C1A3C27D01FFCD35C5FAA88BF86AFB" ID-CoL="0500beb1-cd24-4737-8e7d-b8c49af12e59" authority="Criales, 1981" authorityName="Criales" authorityYear="1981" box="[199,659,1404,1429]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="conchae">
<emphasis id="4DB504537D01FFCD35C5FAA888816AFB" box="[199,490,1404,1428]" italics="true" pageId="5" pageNumber="538">Pseudocoutierea conchae</emphasis>
<bibRefCitation id="1B50A5B07D01FFCD34F7FAA88BF86AFB" box="[501,659,1404,1429]" pageId="5" pageNumber="538" refId="ref9328" refString="Criales, M. M. (1981) Two new species of Pseudocoutierea (Decapoda Natantia, Palaemonidae) from the Colombian Caribbean. Crustaceana, 41 (2), 167 - 181. https: // doi. org / 10.1163 / 156854081 X 00219" type="journal article">Criales, 1981</bibRefCitation>
</taxonomicName>
: ZMA DE 103.237, holotype, male, Colombia, 11°18'N, 74°10'W, depth 
<quantity id="B83975A47D01FFCD35DEFA4B887B6AD7" box="[220,272,1439,1464]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="538" unit="m" value="15.0">15m</quantity>
, 
<date id="0B7FFE817D01FFCD341EFA4B88FB6AD7" box="[284,400,1439,1464]" pageId="5" pageNumber="538" value="1976-02-15">15.ii.1976</date>
, from 
<taxonomicName id="B8C1A3C27D01FFCD34DBFA748AFE6AD7" ID-CoL="3TQ92" authority="(Lamarck, 1815)" baseAuthorityName="Lamarck" baseAuthorityYear="1815" box="[473,917,1439,1464]" class="Anthozoa" family="Gorgoniidae" genus="Leptogorgia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="538" phylum="Cnidaria" rank="species" species="virgulata">
<emphasis id="4DB504537D01FFCD34DBFA748BA56AD7" box="[473,718,1440,1464]" italics="true" pageId="5" pageNumber="538">Leptogorgia virgulata</emphasis>
(Lamarck, 1815)
</taxonomicName>
.
</paragraph>
<paragraph id="7F7ED8417D01FFCD35C5FA108B486903" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">
<taxonomicName id="B8C1A3C27D01FFCD35C5FA108BE36AB3" ID-CoL="3e033cc0-8af9-4b7a-817b-f52a49f0790a" authority="Criales, 1981" authorityName="Criales" authorityYear="1981" box="[199,648,1476,1501]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="edentata">
<emphasis id="4DB504537D01FFCD35C5FA1088826AB3" box="[199,489,1476,1500]" italics="true" pageId="5" pageNumber="538">Pseudocoutierea edentata</emphasis>
<bibRefCitation id="1B50A5B07D01FFCD34EDFA108BE36AB3" box="[495,648,1476,1501]" pageId="5" pageNumber="538" refId="ref9328" refString="Criales, M. M. (1981) Two new species of Pseudocoutierea (Decapoda Natantia, Palaemonidae) from the Colombian Caribbean. Crustaceana, 41 (2), 167 - 181. https: // doi. org / 10.1163 / 156854081 X 00219" type="journal article">Criales, 1981</bibRefCitation>
</taxonomicName>
: ZMA DE 103.238, 1 juvenile holotype, Colombia, 11°18'N, 74°10'W, depth 
<quantity id="B83975A47D01FFCD35DCFA3C8878696F" box="[222,275,1512,1536]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="538" unit="m" value="18.0">18m</quantity>
, 
<date id="0B7FFE817D01FFCD3423FA3388F2696F" box="[289,409,1511,1536]" pageId="5" pageNumber="538" value="1977-04-17">17.iv.1977</date>
, from 
<taxonomicName id="B8C1A3C27D01FFCD34E5FA3C8AC1696F" ID-CoL="3TQ92" authority="(Lamarck, 1815)" baseAuthorityName="Lamarck" baseAuthorityYear="1815" box="[487,938,1511,1536]" class="Anthozoa" family="Gorgoniidae" genus="Leptogorgia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="538" phylum="Cnidaria" rank="species" species="virgulata">
<emphasis id="4DB504537D01FFCD34E5FA3C8B8B696F" box="[487,736,1512,1536]" italics="true" pageId="5" pageNumber="538">Leptogorgia virgulata</emphasis>
(Lamarck, 1815)
</taxonomicName>
; ZMA: DE 103.239, 1 ovigerous female, 6 non-ovigerous female paratypes, Colombia, 11°18'N, 74°10'W, 
<date id="0B7FFE817D01FFCD3664F9D88AB5694A" box="[870,990,1548,1573]" pageId="5" pageNumber="538" value="1977-04-17">17.iv.1977</date>
, from 
<taxonomicName id="B8C1A3C27D01FFCD312BF9D989B06927" ID-CoL="3TQ92" authority="(Lamarck, 1815)" baseAuthorityName="Lamarck" baseAuthorityYear="1815" class="Anthozoa" family="Gorgoniidae" genus="Leptogorgia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="5" pageNumber="538" phylum="Cnidaria" rank="species" species="virgulata">
<emphasis id="4DB504537D01FFCD312BF9D98C4B694B" box="[1065,1312,1548,1572]" italics="true" pageId="5" pageNumber="538">Leptogorgia virgulata</emphasis>
(Lamarck, 1815)
</taxonomicName>
; RMNH.CRUS.D.51368, 1 ovigerous female, Fort Lauderdale, Florida, USA, depth 
<quantity id="B83975A47D01FFCD31ACF9FB8D896927" box="[1198,1250,1583,1608]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="538" unit="m" value="50.0">50m</quantity>
, on 
<taxonomicName id="B8C1A3C27D01FFCD3017F9E48C016927" ID-CoL="62SCB" authorityName="Soika" authorityYear="1974" box="[1301,1386,1584,1608]" class="Insecta" family="Eumenidae" genus="Elisella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D01FFCD3017F9E48C016927" box="[1301,1386,1584,1608]" italics="true" pageId="5" pageNumber="538">Elisella</emphasis>
</taxonomicName>
sp., spring 1975, collected by R. Guest.
</paragraph>
<paragraph id="7F7ED8417D01FFCD35C5F9AC88A569B7" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">
<taxonomicName id="B8C1A3C27D01FFCD35C5F9AC8BF069FF" ID-CoL="4dcc6628-2e21-4a14-91b5-439e5c489054" authority="Holthuis, 1951" authorityName="Holthuis" authorityYear="1951" box="[199,667,1655,1680]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="538" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="4DB504537D01FFCD35C5F9AC888869FF" box="[199,483,1656,1680]" italics="true" pageId="5" pageNumber="538">Pseudocoutierea elegans</emphasis>
Holthuis, 1951
</taxonomicName>
: RMNH. 
<materialsCitation id="CFA9D21C7D01FFCD3605F9AC8D8769DB" ID-GBIF-Occurrence="1841141212" collectingDate="1934-03-02" collectionCode="CRUS" country="Mexico" elevation="45" location="White Friars" municipality="Don" pageId="5" pageNumber="538" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero" typeStatus="paratype">
<collectionCode id="19D040847D01FFCD3605F9AC8A3F69FF" box="[775,852,1656,1680]" pageId="5" pageNumber="538">CRUS</collectionCode>
.D.9194, 
<specimenCount id="69C713C87D01FFCD36BBF9AC8D7569FF" box="[953,1054,1655,1680]" pageId="5" pageNumber="538" type="female">1 female</specimenCount>
<typeStatus id="A07A66E37D01FFCD312BF9AC8DE769FF" box="[1065,1164,1656,1680]" pageId="5" pageNumber="538" type="paratype">paratype</typeStatus>
, south of 
<location id="7A1E8E9A7D01FFCD3007F9A38CF369FF" LSID="urn:lsid:plazi:treatment:F76869577D01FFC43595FC7D884A6FBA:7A1E8E9A7D01FFCD3007F9A38CF369FF" box="[1285,1432,1655,1680]" country="Mexico" municipality="Don" name="White Friars" pageId="5" pageNumber="538" stateProvince="Guerrero">White Friars</location>
, 
<collectingRegion id="BD0516A37D01FFCD3595F948899469DB" box="[151,255,1692,1716]" country="Mexico" name="Guerrero" pageId="5" pageNumber="538">Guerrero</collectingRegion>
, West 
<collectingCountry id="07D698D17D01FFCD344BF94888CA69DA" box="[329,417,1692,1717]" name="Mexico" pageId="5" pageNumber="538">Mexico</collectingCountry>
, depth 
<elevation id="F4EC3F727D01FFCD34F0F9488B4C69DA" box="[498,551,1692,1717]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="538" unit="m" value="45.0">
<quantity id="B83975A47D01FFCD34F0F9488B4C69DA" box="[498,551,1692,1717]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="538" unit="m" value="45.0">45m</quantity>
</elevation>
, 
<collectingDate id="1B3B07697D01FFCD3730F9488BCB69DA" box="[562,672,1692,1717]" pageId="5" pageNumber="538" value="1934-03-02">
<date id="0B7FFE817D01FFCD3730F9488BCB69DA" box="[562,672,1692,1717]" pageId="5" pageNumber="538" value="1934-03-02">2.iii.1934</date>
</collectingDate>
, rock, gorgonids, Allan Hancock Exp. 264–34. 
<collectingMunicipality id="9F1A423B7D01FFCD31BAF9498D8769DB" box="[1208,1260,1693,1716]" pageId="5" pageNumber="538">Don</collectingMunicipality>
</materialsCitation>
. U.S.N.M., reg. no. 90275, acc. no. 128938.
</paragraph>
</subSubSection>
<subSubSection id="37DB8BCA7D01FFCD35C5F9308A4568BA" pageId="5" pageNumber="538" type="diagnosis">
<paragraph id="7F7ED8417D01FFCD35C5F9308DFD6807" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">
<emphasis id="4DB504537D01FFCD35C5F930882A6992" bold="true" box="[199,321,1764,1789]" pageId="5" pageNumber="538">Diagnosis.</emphasis>
Body dorsoventrally depressed, wing-like expansions of rostrum quadrate with anteriorly sharp supra-ocular tooth. No hepatic protuberance, but with epigastric tooth in extension of rostrum. Third to fifth pleonal pleura with posterior tooth. Triangular spine on posterior margin of sixth pleonite. Scaphocerite 2.8 times as long as wide, outer border slightly concave. Mandible with four teeth on incisor process.
</paragraph>
<paragraph id="7F7ED8417D01FFCD35C5F8A08A4568BA" blockId="5.[151,1437,1044,2005]" pageId="5" pageNumber="538">Second maxilliped without exopod, epipod rounded and directed upwards, not reaching end of merus. Third to fifth pereiopod with well-developed meral swelling and infero-distal spine on propodus. Exopod of uropod with small moveable spine between blade and distolateral spine.</paragraph>
</subSubSection>
<subSubSection id="37DB8BCA7D02FFC135C5FF43886F6F97" lastPageId="9" lastPageNumber="542" pageId="6" pageNumber="539" type="description">
<paragraph id="7F7ED8417D02FFCE35C5FF438B7D6E0B" blockId="6.[151,1436,151,571]" pageId="6" pageNumber="539">
<emphasis id="4DB504537D02FFCE35C5FF4388326FDF" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="539">Description.</emphasis>
Carapace broad (
<figureCitation id="E7FAC4C47D02FFCE371CFF438BF36FDF" box="[542,664,151,176]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A–C</figureCitation>
), slightly depressed. Sharp antennal tooth present on anterior margin of carapace with acuminate lobe just below, giving bifid appearance to antennal tooth in lateral view (
<figureCitation id="E7FAC4C47D02FFCE3031FF688CE56FBB" box="[1331,1422,188,213]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
). Well-developed pterygostomial sinus present on anterior margin of carapace below antennal tooth (
<figureCitation id="E7FAC4C47D02FFCE31F3FF0B8C216F97" box="[1265,1354,223,248]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
). Short ridge from antennal tooth upwards on carapace terminating behind orbit (
<figureCitation id="E7FAC4C47D02FFCE36EFFED08D226E73" box="[1005,1097,260,285]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
). No hepatic protuberance or tooth present. Longitudinal groove running near lateral margin of carapace extending from pterygostomial sinus to near posterior margin of carapace.
</paragraph>
<paragraph id="7F7ED8417D02FFCE35C5FEA48B606D54" blockId="6.[151,1436,151,571]" pageId="6" pageNumber="539">
Rostrum long, unarmed, reaching beyond bifurcation of upper antennular flagellum (
<figureCitation id="E7FAC4C47D02FFCE3196FEBB8D8E6EE7" box="[1172,1253,367,392]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig 1B</figureCitation>
), round, curved slightly upwards distally (
<figureCitation id="E7FAC4C47D02FFCE34BAFE478B7A6EC3" box="[440,529,403,428]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
), proximally on both sides expanded into supra-orbital eave with anteriorly sharp supra-ocular tooth (
<figureCitation id="E7FAC4C47D02FFCE3476FE6388856EBF" box="[372,494,439,464]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1B, C</figureCitation>
). Expansions continue as small ridge downwards on carapace and terminate behind orbit close to end of ridge from antennal tooth (
<figureCitation id="E7FAC4C47D02FFCE37AAFE0F8A6A6E9C" box="[680,769,475,500]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
). Rostrum continues as ridge on carapace; rostrum elevated above eyes (
<figureCitation id="E7FAC4C47D02FFCE3424FE2A88EA6D78" box="[294,385,510,535]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
), epigastric tooth present behind elevation above point where ridges from supraorbital eave and antennal tooth end (
<figureCitation id="E7FAC4C47D02FFCE34A7FDF688966D54" box="[421,509,546,571]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539">Fig. 1A</figureCitation>
).
</paragraph>
<caption id="2BBE88C97D02FFCE3595F8AD8D1368A3" httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="6" pageNumber="539" startId="6.[151,250,1913,1935]" targetBox="[180,1396,657,1855]" targetPageId="6">
<paragraph id="7F7ED8417D02FFCE3595F8AD8D1368A3" blockId="6.[151,1435,1913,1996]" pageId="6" pageNumber="539">
<emphasis id="4DB504537D02FFCE3595F8AD887E68E1" bold="true" box="[151,277,1913,1935]" pageId="6" pageNumber="539">FIGURE 1.</emphasis>
<taxonomicName id="B8C1A3C27D02FFCE341FF8AD8B2468E0" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[285,591,1913,1935]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="539" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D02FFCE341FF8AD8B2468E0" box="[285,591,1913,1935]" italics="true" pageId="6" pageNumber="539">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D02FFCE3755F8AD8BC168E1" box="[599,682,1913,1934]" pageId="6" pageNumber="539" rank="species">
<emphasis id="4DB504537D02FFCE3755F8AD8BC168E1" bold="true" box="[599,682,1913,1934]" pageId="6" pageNumber="539">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna. Scale bar A–C = 2.0mm; D = 1.5mm; E = 0.4mm; F, G = 0.6mm.
</paragraph>
</caption>
<caption id="2BBE88C97D03FFCF3595FA3E8D926970" httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="7" pageNumber="540" startId="7.[151,250,1514,1536]" targetBox="[164,1427,210,1468]" targetPageId="7">
<paragraph id="7F7ED8417D03FFCF3595FA3E8D926970" blockId="7.[151,1435,1514,1567]" pageId="7" pageNumber="540">
<emphasis id="4DB504537D03FFCF3595FA3E887F6A90" bold="true" box="[151,276,1514,1536]" pageId="7" pageNumber="540">FIGURE 2.</emphasis>
<taxonomicName id="B8C1A3C27D03FFCF341EFA3E8B26696F" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[284,589,1514,1536]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="540" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D03FFCF341EFA3E8B26696F" box="[284,589,1514,1536]" italics="true" pageId="7" pageNumber="540">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D03FFCF3757FA3F8BCC696F" box="[597,679,1515,1536]" pageId="7" pageNumber="540" rank="species">
<emphasis id="4DB504537D03FFCF3757FA3F8BCC696F" bold="true" box="[597,679,1515,1536]" pageId="7" pageNumber="540">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm.
</paragraph>
</caption>
<paragraph id="7F7ED8417D03FFCF35C5F99F8B4C69A0" blockId="7.[151,1437,1611,2030]" pageId="7" pageNumber="540">
First two pleonal segments with pleura rounded. Third pleuron with small posteroventral tooth (
<figureCitation id="E7FAC4C47D03FFCF3033F99F8CE5690B" box="[1329,1422,1611,1636]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="7" pageNumber="540">Fig. 1A</figureCitation>
). Fourth and fifth pleura ending in sharp, slender point. Posterolateral angle of sixth pleonite sharply triangular (
<figureCitation id="E7FAC4C47D03FFCF306DF9BB89D069C3" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="7" pageNumber="540">Fig. 1A</figureCitation>
); posteroventral angle small, acuminate. Sixth pleonite two times as long as fifth, 0.9 times as long as telson measured dorsally (without spines).
</paragraph>
<paragraph id="7F7ED8417D03FFCF35C5F90E88806831" blockId="7.[151,1437,1611,2030]" pageId="7" pageNumber="540">
Telson with two pairs of dorsolateral spines (about 0.05 times as long as telson), proximal pair at 0.56 of telson length, distal pair at 0.87 to 0.9 of telson length. Tip with three pairs of cuspidate setae, outer pair 0.31 times as long as intermediate pair, intermediate pair 0.20 times as long as telson, medial spines 0.74 times as long as intermediate pair (
<figureCitation id="E7FAC4C47D03FFCF3467F89188B66831" box="[357,477,1861,1886]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="7" pageNumber="540">Fig. 1D, E</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D03FFCF35C5F8BD886F68C9" blockId="7.[151,1437,1611,2030]" pageId="7" pageNumber="540">
Eyestalks large, broad, cornea not much broader than eyestalks, small spine dorsally on eyestalk near cornea (
<figureCitation id="E7FAC4C47D03FFCF359DF859899D68C9" box="[159,246,1933,1958]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="7" pageNumber="540">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D03FFC035C5F8658C3A6F97" blockId="7.[151,1437,1611,2030]" lastBlockId="8.[151,1437,151,2003]" lastPageId="8" lastPageNumber="541" pageId="7" pageNumber="540">
Antennular peduncle with short stylocerite measuring about 0.4 of length of basal segment (without distolateral spine), distolateral spine well developed reaching to about end of intermediate segment, acute tooth present on ventral side near inner margin (
<figureCitation id="E7FAC4C47D0CFFC03789FF438B886FDF" box="[651,739,151,176]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="8" pageNumber="541">Fig. 1F</figureCitation>
). Ultimate segment about 1.9 times as long as intermediate segment, intermediate segment 0.23 times as long as basal segment. Upper/outer antennular flagellum fused for three segments; short inner ramus two-segmented; longer outer ramus about as long as antennular peduncle.
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FED78D686E50" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Scaphocerite overreaching antennular peduncle, about 2.8 times as long as wide, outer margin slightly concave, distolateral tooth not overreaching distal margin of lamina (
<figureCitation id="E7FAC4C47D0CFFC0369EFEF28A9E6E50" box="[924,1013,294,319]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="8" pageNumber="541">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FE9E89A16EE8" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Molar process of mandible with large blunt teeth and setal brushes; incisor process with 4 distal teeth (
<figureCitation id="E7FAC4C47D0CFFC0306DFE9E89D66EE8" captionStart="FIGURE 2" captionStartId="7.[151,250,1514,1536]" captionTargetBox="[164,1427,210,1468]" captionTargetId="figure@7.[151,1436,193,1493]" captionTargetPageId="7" captionText="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="8" pageNumber="541">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FE468B976EA1" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Maxillula with hook-like process on palp, hook directed upwards. Lower lacinia lost during dissection, upper lacinia with both plumose and simple spines (
<figureCitation id="E7FAC4C47D0CFFC03795FE618B856EA1" box="[663,750,437,462]" captionStart="FIGURE 2" captionStartId="7.[151,250,1514,1536]" captionTargetBox="[164,1427,210,1468]" captionTargetId="figure@7.[151,1436,193,1493]" captionTargetPageId="7" captionText="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="8" pageNumber="541">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FE0D8C216D79" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Maxilla with endite cleft, apical setae serrate on distal lobe, simple setae on proximal lobe, lobes subequal in length. Scaphognathite well developed, outer margin slightly convex. Palp distinct, without setae (
<figureCitation id="E7FAC4C47D0CFFC031E6FE298C566D79" box="[1252,1341,509,534]" captionStart="FIGURE 2" captionStartId="7.[151,250,1514,1536]" captionTargetBox="[164,1427,210,1468]" captionTargetId="figure@7.[151,1436,193,1493]" captionTargetPageId="7" captionText="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="8" pageNumber="541">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FDF58A5B6D32" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
First maxilliped without flagellum on caridean lobe; palp short; epipod well developed, bilobed. Outer margins of basal endite and caridean lobe slightly convex (
<figureCitation id="E7FAC4C47D0CFFC037CBFD908A4F6D32" box="[713,804,580,605]" captionStart="FIGURE 2" captionStartId="7.[151,250,1514,1536]" captionTargetBox="[164,1427,210,1468]" captionTargetId="figure@7.[151,1436,193,1493]" captionTargetPageId="7" captionText="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="8" pageNumber="541">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FDBC88C56DCA" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Second maxilliped without exopod, epipod consisting of one rounded lobe directed upwards, not reaching meral segment (
<figureCitation id="E7FAC4C47D0CFFC03448FD5888C96DCA" box="[330,418,652,677]" captionStart="FIGURE 2" captionStartId="7.[151,250,1514,1536]" captionTargetBox="[164,1427,210,1468]" captionTargetId="figure@7.[151,1436,193,1493]" captionTargetPageId="7" captionText="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="8" pageNumber="541">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FD7B889A6C7F" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Third maxilliped without exopod, overreaching basicerite, without arthrobranch but with small knob there. Ultimate segment about as long as penultimate segment, antepenultimate segment more than twice as long as penultimate segment (
<figureCitation id="E7FAC4C47D0CFFC0348DFD23888F6C7F" box="[399,484,759,784]" captionStart="FIGURE 2" captionStartId="7.[151,250,1514,1536]" captionTargetBox="[164,1427,210,1468]" captionTargetId="figure@7.[151,1436,193,1493]" captionTargetPageId="7" captionText="FIGURE 2. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, mandible; B, maxillula; C, maxilla; D, first maxilliped; E, second maxilliped; F, third maxilliped. Scale bar A, B = 0.3mm; C–F = 0.5mm." httpUri="https://zenodo.org/record/1242213/files/figure.png" pageId="8" pageNumber="541">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FCCF8B7E6C14" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
First pereiopod reaching just beyond scaphocerite. Chela slender with fingers 0.5 times as long as palm. Carpus 0.9 times as long as chela and 0.9 times as long as merus (
<figureCitation id="E7FAC4C47D0CFFC0367CFCEA8A936C38" box="[894,1016,830,855]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Fig. 3A, B</figureCitation>
). Cutting edge of chela entire, tips of fingers strongly hooked (
<figureCitation id="E7FAC4C47D0CFFC034B2FCB68B636C14" box="[432,520,866,891]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FC528C016B3E" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Second pereiopods strongly unequal: major second pereiopod about 2.8 times as long as minor second pereiopod. Especially palm more robust in major second pereiopod (fig. 2F, G), being about 4.5 times as broad as palm of minor second pereiopod (fig. 3D) at broadest point. Major second pereiopod with merus reaching to about end of scaphocerite, with tubercles on surface of merus and propodus (
<figureCitation id="E7FAC4C47D0CFFC036D2FC258D276B65" box="[976,1100,1009,1034]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Fig. 3F, G</figureCitation>
). Merus 1.3 times as long as ischium, 3.2 times as long as carpus. Carpus 0.2 times as long as palm, fingers 0.39 times as long as palm. Dactylus with small tooth on cutting edge, with indent in fixed finger to accommodate movement of dactylus (
<figureCitation id="E7FAC4C47D0CFFC03001FBED8C366B3E" box="[1283,1373,1081,1106]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Fig. 3H</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FB888B3F6BD2" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Minor second pereiopod with merus 2.8 times as long as ischium, 1.6 times as long as carpus (
<figureCitation id="E7FAC4C47D0CFFC031E2FB888C516B1A" box="[1248,1338,1116,1141]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Fig. 3D</figureCitation>
). Carpus 0.6 times as long as palm, chela with long fingers (0,63 times as long as palm), no tubercles on merus or propodus. Cutting edge of fingers entire (
<figureCitation id="E7FAC4C47D0CFFC034F2FB708B2C6BD2" box="[496,583,1188,1213]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FB138D136A00" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Third pereiopod robust; merus slightly curved, about 1.8 times as long as ischium (
<figureCitation id="E7FAC4C47D0CFFC03174FB138DAE6B8F" box="[1142,1221,1223,1248]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">
Fig. 
<date id="0B7FFE817D0CFFC031AEFB1C8DAE6B8F" box="[1196,1221,1224,1248]" pageId="8" pageNumber="541">3I</date>
</figureCitation>
). Carpus about 0.4 times as long as merus; propodus curved, slightly longer than merus. Merus with swelling on distal part of flexor margin (
<figureCitation id="E7FAC4C47D0CFFC035F6FADB88276A47" box="[244,332,1295,1320]" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">
Figs. 
<date id="0B7FFE817D0CFFC03431FAC488276A47" box="[307,332,1296,1320]" pageId="8" pageNumber="541">3I</date>
</figureCitation>
, 
<figureCitation id="E7FAC4C47D0CFFC03455FAC488FC6A47" box="[343,407,1296,1320]" captionStart="FIGURE 4" captionStartId="11.[151,250,1425,1447]" captionTargetBox="[305,1275,201,1383]" captionTargetId="figure@11.[296,1290,193,1403]" captionTargetPageId="11" captionText="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="8" pageNumber="541">4E, F</figureCitation>
). Two small spines present in distoventral part of propodus together with several setae (
<figureCitation id="E7FAC4C47D0CFFC03066FADB89D96A24" captionStart="FIGURE 3" captionStartId="10.[151,250,1914,1936]" captionTargetBox="[238,1384,214,1885]" captionTargetId="figure@10.[193,1394,194,1895]" captionTargetPageId="10" captionText="FIGURE 3. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod;E, idem, chela; F,major rightsecond pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar:A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm." httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="8" pageNumber="541">Figs. 3J</figureCitation>
, 
<figureCitation id="E7FAC4C47D0CFFC035BFFAE788696A24" box="[189,258,1331,1355]" captionStart="FIGURE 4" captionStartId="11.[151,250,1425,1447]" captionTargetBox="[305,1275,201,1383]" captionTargetId="figure@11.[296,1290,193,1403]" captionTargetPageId="11" captionText="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="8" pageNumber="541">4B, C</figureCitation>
). Dactylus less than half as long as propodus, curved, with small decalcified protuberance in basal part of flexor margin (
<figureCitation id="E7FAC4C47D0CFFC0343DFA8288F16A00" box="[319,410,1366,1391]" captionStart="FIGURE 4" captionStartId="11.[151,250,1425,1447]" captionTargetBox="[305,1275,201,1383]" captionTargetId="figure@11.[296,1290,193,1403]" captionTargetPageId="11" captionText="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="8" pageNumber="541">Fig. 4A</figureCitation>
). Suture visible between corpus and unguis of dactylus (
<figureCitation id="E7FAC4C47D0CFFC03112FA828D006A00" box="[1040,1131,1366,1391]" captionStart="FIGURE 4" captionStartId="11.[151,250,1425,1447]" captionTargetBox="[305,1275,201,1383]" captionTargetId="figure@11.[296,1290,193,1403]" captionTargetPageId="11" captionText="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="8" pageNumber="541">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FAAE886D6AD8" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Pleopods with endopod slightly shorter than exopod, second to fifth pleopods of female with appendix interna (
<figureCitation id="E7FAC4C47D0CFFC0359DFA4A89916AD8" box="[159,250,1438,1463]" captionStart="FIGURE5" captionStartId="12.[151,250,923,945]" captionTargetBox="[196,1417,286,867]" captionTargetId="figure@12.[151,1436,269,885]" captionTargetPageId="12" captionText="FIGURE5. Pseudocoutierea acutidorsata sp. nov. A, right second pleopod, plumosehairs on blades omitted,female paratype, RMNH.CRUS.D.57273; B, variation in epigastric spine female paratypes RMNH.CRUS.D.57273; B1, ovigerous female pocl. 2.25mm; B2, ovigerous female pocl. 2.1mm; B3, female pocl. 1.75mm; B4, ovigerous female pocl. 2.75mm. Pseudocoutierea conchae Criales, 1981, male holotype, ZMA DE 103.237. Pleonal segments 3 to 5. C, pleura on right side in lateral view; D, pleura on left side in lateral view.Scale bar A, B = 1mm; C, D = 1.5mm." httpUri="https://zenodo.org/record/1242219/files/figure.png" pageId="8" pageNumber="541">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FA168DA96AB4" blockId="8.[151,1437,151,2003]" box="[199,1218,1474,1499]" pageId="8" pageNumber="541">
Exopod of uropod with tiny moveable spine between distolateral tooth and blade (
<figureCitation id="E7FAC4C47D0CFFC0315DFA168DDF6AB4" box="[1119,1204,1474,1499]" captionStart="FIGURE 1" captionStartId="6.[151,250,1913,1935]" captionTargetBox="[180,1396,657,1855]" captionTargetId="figure@6.[151,1436,645,1879]" captionTargetPageId="6" captionText="FIGURE 1. Pseudocoutierea acutidorsata sp. nov., female paratype, RMNH.CRUS.D.57273. A, habitus in lateral view; B, carapace and anterior appendages, dorsal view; C, carapace, dorsal view, eyes omitted; D, telson and uropods; E, distal part telson; F, antennula; G, antenna.Scale bar A–C = 2.0mm; D = 1.5mm; E= 0.4mm; F, G = 0.6mm." httpUri="https://zenodo.org/record/1242211/files/figure.png" pageId="8" pageNumber="541">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5FA32888B6870" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
<emphasis id="4DB504537D0CFFC035C5FA3288F46A90" bold="true" box="[199,415,1510,1535]" pageId="8" pageNumber="541">Special structures</emphasis>
. Species of the genus 
<taxonomicName id="B8C1A3C27D0CFFC037ABFA328A0F6A91" authorityName="Holthuis" authorityYear="1951" box="[681,868,1510,1534]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D0CFFC037ABFA328A0F6A91" box="[681,868,1510,1534]" italics="true" pageId="8" pageNumber="541">Pseudocoutierea</emphasis>
</taxonomicName>
all possess a swelling (either triangular, or hook shaped in 
<taxonomicName id="B8C1A3C27D0CFFC03413F9DF8804694C" authorityName="De Grave" authorityYear="2007" box="[273,367,1547,1571]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" species="dotae">
<emphasis id="4DB504537D0CFFC03413F9DF8804694C" box="[273,367,1547,1571]" italics="true" pageId="8" pageNumber="541">P. dotae</emphasis>
</taxonomicName>
only) on the merus of the third pereiopod and a protuberance on the flexor margin of the dactylus. d’Udekem d’Acoz (2000) first examined these protuberances in more detail for 
<taxonomicName id="B8C1A3C27D0CFFC03191F9FB8D9B6929" authorityName="d'Udekem d'Acoz" authorityYear="2000" box="[1171,1264,1583,1606]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" species="wirtzi">
<emphasis id="4DB504537D0CFFC03191F9FB8D9B6929" box="[1171,1264,1583,1606]" italics="true" pageId="8" pageNumber="541">P. wirtzi</emphasis>
</taxonomicName>
. He found that the meral swelling had a decalcified part and hypothesized that under these swellings exocrine glands could be present which would possibly secrete through these decalcifications or via the setae beside it. To see if these decalcified parts were also present in the new species, the third pereiopod was photographed with a scanning electron microscope. No decalcifications were present and only some indents could be seen on the meral swelling (
<figureCitation id="E7FAC4C47D0CFFC0359DF936887C6994" box="[159,279,1762,1787]" captionStart="FIGURE 4" captionStartId="11.[151,250,1425,1447]" captionTargetBox="[305,1275,201,1383]" captionTargetId="figure@11.[296,1290,193,1403]" captionTargetPageId="11" captionText="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="8" pageNumber="541">Fig. 4E, F</figureCitation>
). If an exocrine gland is present in the meral swelling of this species, it can only secrete through the setae present on the swelling.
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5F8FE8D0168E4" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
Another interesting structure on the third pereiopod is the decalcified part on the flexor border of the dactylus of the third pereiopod. That structure was also first described for this genus in 
<taxonomicName id="B8C1A3C27D0CFFC0313DF89B8DF46809" authorityName="d'Udekem d'Acoz" authorityYear="2000" box="[1087,1183,1871,1894]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" species="wirtzi">
<emphasis id="4DB504537D0CFFC0313DF89B8DF46809" box="[1087,1183,1871,1894]" italics="true" pageId="8" pageNumber="541">P. wirtzi</emphasis>
</taxonomicName>
. We also made some photographs of the third pereiopod and found the exact same decalcified part (
<figureCitation id="E7FAC4C47D0CFFC03101F8A68D3568E4" box="[1027,1118,1906,1931]" captionStart="FIGURE 4" captionStartId="11.[151,250,1425,1447]" captionTargetBox="[305,1275,201,1383]" captionTargetId="figure@11.[296,1290,193,1403]" captionTargetPageId="11" captionText="FIGURE 4. Pseudocoutierea acutidorsata sp. nov. female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus;B, flexor border between propodus and dactylus, showingthe two infero-distal spines,ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view." httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="8" pageNumber="541">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="7F7ED8417D0CFFC035C5F8428CF768BC" blockId="8.[151,1437,151,2003]" pageId="8" pageNumber="541">
The epigastric tooth is a relatively new character for this genus that was first described for 
<taxonomicName id="B8C1A3C27D0CFFC031D7F8438C1368C1" authorityName="Ramos-Tafur &amp; Lemaitre" authorityYear="2016" box="[1237,1400,1942,1966]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" species="stephanieae">
<emphasis id="4DB504537D0CFFC031D7F8438C1368C1" box="[1237,1400,1942,1966]" italics="true" pageId="8" pageNumber="541">P. stephanieae</emphasis>
</taxonomicName>
by Ramos-Tafur &amp; Lemaitre (2016). Although all eight currently reported specimens of 
<taxonomicName id="B8C1A3C27D0CFFC03186F86F8C5A68BC" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[1156,1329,1979,2003]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="541" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0CFFC03186F86F8C5A68BC" box="[1156,1329,1979,2003]" italics="true" pageId="8" pageNumber="541">P. acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0CFFC0303FF86E8CF768BC" box="[1341,1436,1978,2003]" pageId="8" pageNumber="541" rank="species">
<emphasis id="4DB504537D0CFFC0303FF86E8CF768BC" bold="true" box="[1341,1436,1978,2003]" pageId="8" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="7F7ED8417D0DFFC13595FF4C886F6F97" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
possess the aforementioned tooth, there seems to be extensive variability in the exact form of this tooth. In some specimens the tooth is nothing more than a small knob while in other specimens it is elongated and almost rod-like (
<figureCitation id="E7FAC4C47D0DFFC1359DFF0B899C6F97" box="[159,247,223,248]" captionStart="FIGURE5" captionStartId="12.[151,250,923,945]" captionTargetBox="[196,1417,286,867]" captionTargetId="figure@12.[151,1436,269,885]" captionTargetPageId="12" captionText="FIGURE5. Pseudocoutierea acutidorsata sp. nov. A, right second pleopod, plumosehairs on blades omitted,female paratype, RMNH.CRUS.D.57273; B, variation in epigastric spine female paratypes RMNH.CRUS.D.57273; B1, ovigerous female pocl. 2.25mm; B2, ovigerous female pocl. 2.1mm; B3, female pocl. 1.75mm; B4, ovigerous female pocl. 2.75mm. Pseudocoutierea conchae Criales, 1981, male holotype, ZMA DE 103.237. Pleonal segments 3 to 5. C, pleura on right side in lateral view; D, pleura on left side in lateral view.Scale bar A, B = 1mm; C, D = 1.5mm." httpUri="https://zenodo.org/record/1242219/files/figure.png" pageId="9" pageNumber="542">Fig. 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="37DB8BCA7D0DFFC135C5FED088466D77" pageId="9" pageNumber="542" type="etymology">
<paragraph id="7F7ED8417D0DFFC135C5FED08A706E0B" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
<emphasis id="4DB504537D0DFFC135C5FED0882D6E72" bold="true" box="[199,326,260,285]" pageId="9" pageNumber="542">Etymology</emphasis>
. This species is named after the epigastric tooth on the carapace which separates it from almost all other species of 
<taxonomicName id="B8C1A3C27D0DFFC13459FEFC8B7D6E2F" authorityName="Holthuis" authorityYear="1951" box="[347,534,296,320]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D0DFFC13459FEFC8B7D6E2F" box="[347,534,296,320]" italics="true" pageId="9" pageNumber="542">Pseudocoutierea</emphasis>
</taxonomicName>
. The name is derived from the latin 
<emphasis id="4DB504537D0DFFC136D1FEFD8D776E2F" box="[979,1052,297,320]" italics="true" pageId="9" pageNumber="542">acutus</emphasis>
-sharp and 
<emphasis id="4DB504537D0DFFC131A0FEFC8D9D6E2F" box="[1186,1270,296,320]" italics="true" pageId="9" pageNumber="542">dorsum</emphasis>
-back, 
<emphasis id="4DB504537D0DFFC13044FEFC8CF06E2F" box="[1350,1435,296,320]" italics="true" pageId="9" pageNumber="542">dorsata</emphasis>
makes it an adjective suited for the feminine genus name.
</paragraph>
<paragraph id="7F7ED8417D0DFFC135C5FEBB88466D77" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
<emphasis id="4DB504537D0DFFC135C5FEBB88486EE7" bold="true" box="[199,291,367,392]" pageId="9" pageNumber="542">Habitat</emphasis>
. Eight specimens were collected from a colony of 
<taxonomicName id="B8C1A3C27D0DFFC13675FEA489B36EC2" authority="(Milne Edwards &amp; Haime, 1857)" baseAuthorityName="Milne Edwards &amp; Haime" baseAuthorityYear="1857" class="Anthozoa" family="Primnoidae" genus="Callogorgia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="9" pageNumber="542" phylum="Cnidaria" rank="species" species="gracilis">
<emphasis id="4DB504537D0DFFC13675FEA48D356EE7" box="[887,1118,368,392]" italics="true" pageId="9" pageNumber="542">Callogorgia gracilis</emphasis>
(Milne Edwards &amp; Haime, 1857)
</taxonomicName>
(Anthozoa, 
<taxonomicName id="B8C1A3C27D0DFFC13460FE4088836EC2" box="[354,488,404,429]" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="9" pageNumber="542" phylum="Cnidaria" rank="order">Alcyonacea</taxonomicName>
) at a depth of 138 meters on the deep reef of 
<collectingCountry id="07D698D17D0DFFC136DCFE408D5C6EC2" box="[990,1079,404,429]" name="Bonaire, Sint Eustatius and Saba" pageId="9" pageNumber="542">Bonaire</collectingCountry>
(Dutch Caribbean). No real-life colors were recorded for this shrimp before it was fixed in ethanol. The three last pereiopods were curved inwards as described by d’Udekem d’Acoz (2000) and we agree with his explanation of this position by the lifestyle of these shrimp.
</paragraph>
</subSubSection>
<subSubSection id="37DB8BCA7D0DFFC435C5FDF0884A6FBA" lastPageId="12" lastPageNumber="545" pageId="9" pageNumber="542" type="discussion">
<paragraph id="7F7ED8417D0DFFC135C5FDF08B886D9F" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
<emphasis id="4DB504537D0DFFC135C5FDF0885F6D52" bold="true" box="[199,308,548,573]" pageId="9" pageNumber="542">Remarks</emphasis>
. 
<taxonomicName id="B8C1A3C27D0DFFC1344AFDF08BF56D53" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[328,670,548,572]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0DFFC1344AFDF08BF56D53" box="[328,670,548,572]" italics="true" pageId="9" pageNumber="542">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0DFFC137A9FDF08A606D52" box="[683,779,548,573]" pageId="9" pageNumber="542" rank="species">
<emphasis id="4DB504537D0DFFC137A9FDF08A606D52" bold="true" box="[683,779,548,573]" pageId="9" pageNumber="542">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from most other species in the presence of an epigastric tooth which it shares with only the recently described 
<taxonomicName id="B8C1A3C27D0DFFC13686FD9D8D436D0F" authorityName="Ramos-Tafur &amp; Lemaitre" authorityYear="2016" box="[900,1064,584,608]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="stephanieae">
<emphasis id="4DB504537D0DFFC13686FD9D8D436D0F" box="[900,1064,584,608]" italics="true" pageId="9" pageNumber="542">P. stephanieae</emphasis>
</taxonomicName>
. It can be distinguished from 
<taxonomicName id="B8C1A3C27D0DFFC13085FD9D88766DEB" authorityName="Ramos-Tafur &amp; Lemaitre" authorityYear="2016" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="stephanieae">
<emphasis id="4DB504537D0DFFC13085FD9D88766DEB" italics="true" pageId="9" pageNumber="542">P. stephanieae</emphasis>
</taxonomicName>
by the presence of a strong pterygostomial sinus (which is only very shallow in 
<taxonomicName id="B8C1A3C27D0DFFC13198FDB98C506DEB" authorityName="Ramos-Tafur &amp; Lemaitre" authorityYear="2016" box="[1178,1339,620,644]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="stephanieae">
<emphasis id="4DB504537D0DFFC13198FDB98C506DEB" box="[1178,1339,620,644]" italics="true" pageId="9" pageNumber="542">P. stephanieae</emphasis>
</taxonomicName>
) and the length and curvature of the rostrum. In 
<taxonomicName id="B8C1A3C27D0DFFC13763FD458A676DC7" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[609,780,656,680]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0DFFC13763FD458A676DC7" box="[609,780,656,680]" italics="true" pageId="9" pageNumber="542">P. acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0DFFC13614FD5B8A196DC7" box="[790,882,655,680]" pageId="9" pageNumber="542" rank="species">
<emphasis id="4DB504537D0DFFC13614FD5B8A196DC7" bold="true" box="[790,882,655,680]" pageId="9" pageNumber="542">sp. nov.</emphasis>
</taxonomicNameLabel>
, the rostrum is curved upwards and reaches the bifurcation of the upper antennular flagellum, while in 
<taxonomicName id="B8C1A3C27D0DFFC13620FD618AA36DA3" authorityName="Ramos-Tafur &amp; Lemaitre" authorityYear="2016" box="[802,968,692,716]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="stephanieae">
<emphasis id="4DB504537D0DFFC13620FD618AA36DA3" box="[802,968,692,716]" italics="true" pageId="9" pageNumber="542">P. stephanieae</emphasis>
</taxonomicName>
the rostrum just reaches the end of the antennular peduncle and curves slightly downwards.
</paragraph>
<paragraph id="7F7ED8417D0DFFC135C5FD288C196C33" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
<taxonomicName id="B8C1A3C27D0DFFC135C5FD288B726C7B" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[199,537,764,788]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0DFFC135C5FD288B726C7B" box="[199,537,764,788]" italics="true" pageId="9" pageNumber="542">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0DFFC13720FD288B156C7A" box="[546,638,764,789]" pageId="9" pageNumber="542" rank="species">
<emphasis id="4DB504537D0DFFC13720FD288B156C7A" bold="true" box="[546,638,764,789]" pageId="9" pageNumber="542">sp. nov.</emphasis>
</taxonomicNameLabel>
is closely related to the East-Pacific species 
<taxonomicName id="B8C1A3C27D0DFFC13185FD298D916C7B" authorityName="Holthuis" authorityYear="1951" box="[1159,1274,764,788]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="4DB504537D0DFFC13185FD298D916C7B" box="[1159,1274,764,788]" italics="true" pageId="9" pageNumber="542">P. elegans</emphasis>
</taxonomicName>
with which it shares a tiny moveable spine between the distolateral tooth and the blade of the exopod of the uropod, the absence of an exopodal flagellum on the caridean lobe of the first maxilliped and a long rostrum that distally curves up.
</paragraph>
<paragraph id="7F7ED8417D0DFFC135C5FCBC8C756B5A" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
<bibRefCitation id="1B50A5B07D0DFFC135C5FCBC88EF6CEF" box="[199,388,872,896]" pageId="9" pageNumber="542" refId="ref9542" refString="De Grave, S. (2007) A new species of Pseudocoutierea Holthuis from the Caribbean coast of Panama (Crustacea, Decapoda, Palaemonidae), with a key to the genus. Zootaxa, 37 (1397), 29 - 37. https: // doi. org / 10.11646 / zootaxa. 1397.4" type="journal article">De Grave (2007)</bibRefCitation>
and Ramos &amp; Lemaitre (2016) used the placement of the dorsal tooth on the third pleonite (in midlength or distally) to distinguish 
<taxonomicName id="B8C1A3C27D0DFFC13733FC598BC86CCB" authorityName="Holthuis" authorityYear="1951" box="[561,675,908,932]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="4DB504537D0DFFC13733FC598BC86CCB" box="[561,675,908,932]" italics="true" pageId="9" pageNumber="542">P. elegans</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8C1A3C27D0DFFC137D8FC598A396CCB" authorityName="Criales" authorityYear="1981" box="[730,850,908,932]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="conchae">
<emphasis id="4DB504537D0DFFC137D8FC598A396CCB" box="[730,850,908,932]" italics="true" pageId="9" pageNumber="542">P. conchae</emphasis>
</taxonomicName>
. This character seems only based on the drawing of the pleonal somites in the species description of 
<taxonomicName id="B8C1A3C27D0DFFC137C2FC658A516CA7" authorityName="Criales" authorityYear="1981" box="[704,826,944,968]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="conchae">
<emphasis id="4DB504537D0DFFC137C2FC658A516CA7" box="[704,826,944,968]" italics="true" pageId="9" pageNumber="542">P. conchae</emphasis>
</taxonomicName>
(
<bibRefCitation id="1B50A5B07D0DFFC13649FC7B8D206CA7" box="[843,1099,943,968]" pageId="9" pageNumber="542" refId="ref9328" refString="Criales, M. M. (1981) Two new species of Pseudocoutierea (Decapoda Natantia, Palaemonidae) from the Colombian Caribbean. Crustaceana, 41 (2), 167 - 181. https: // doi. org / 10.1163 / 156854081 X 00219" type="journal article">
Criales, 1981, 
<figureCitation id="E7FAC4C47D0DFFC136F0FC7B8D206CA7" box="[1010,1099,943,968]" captionStart="FIGURE 6. 50" captionStartId="12.[151,250,1765,1787]" captionTargetBox="[394,1212,1162,1694]" captionTargetId="figure@12.[340,1247,1134,1734]" captionTargetPageId="12" captionText="FIGURE 6. 50% majority rule consensus tree of Pseudocoutierea species; outgroup Coutierea agassizi (Coutière, 1901); tree length = 28, Consistency Index (CI) = 0.6429." httpUri="https://zenodo.org/record/1242221/files/figure.png" pageId="9" pageNumber="542">Fig. 6C</figureCitation>
</bibRefCitation>
). Upon closer examination of the 
<typeStatus id="A07A66E37D0DFFC135C3FC0088486C82" box="[193,291,980,1005]" pageId="9" pageNumber="542" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="B8C1A3C27D0DFFC13449FC0188AF6C83" authorityName="Criales" authorityYear="1981" box="[331,452,980,1004]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="conchae">
<emphasis id="4DB504537D0DFFC13449FC0188AF6C83" box="[331,452,980,1004]" italics="true" pageId="9" pageNumber="542">P. conchae</emphasis>
</taxonomicName>
, the tooth on the third pleonite was found to be in roughly the same place as the same tooth in other 
<taxonomicName id="B8C1A3C27D0DFFC1343AFC2C88986B7F" authorityName="Holthuis" authorityYear="1951" box="[312,499,1016,1040]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D0DFFC1343AFC2C88986B7F" box="[312,499,1016,1040]" italics="true" pageId="9" pageNumber="542">Pseudocoutierea</emphasis>
</taxonomicName>
species (
<figureCitation id="E7FAC4C47D0DFFC1375FFC238BB06B7F" box="[605,731,1015,1040]" captionStart="FIGURE5" captionStartId="12.[151,250,923,945]" captionTargetBox="[196,1417,286,867]" captionTargetId="figure@12.[151,1436,269,885]" captionTargetPageId="12" captionText="FIGURE5. Pseudocoutierea acutidorsata sp. nov. A, right second pleopod, plumosehairs on blades omitted,female paratype, RMNH.CRUS.D.57273; B, variation in epigastric spine female paratypes RMNH.CRUS.D.57273; B1, ovigerous female pocl. 2.25mm; B2, ovigerous female pocl. 2.1mm; B3, female pocl. 1.75mm; B4, ovigerous female pocl. 2.75mm. Pseudocoutierea conchae Criales, 1981, male holotype, ZMA DE 103.237. Pleonal segments 3 to 5. C, pleura on right side in lateral view; D, pleura on left side in lateral view.Scale bar A, B = 1mm; C, D = 1.5mm." httpUri="https://zenodo.org/record/1242219/files/figure.png" pageId="9" pageNumber="542">Fig. 5C, D</figureCitation>
). The medial placement of the tooth seems to be an artifact in the drawing by 
<bibRefCitation id="1B50A5B07D0DFFC13444FBC8888D6B5B" box="[326,486,1052,1077]" pageId="9" pageNumber="542" refId="ref9328" refString="Criales, M. M. (1981) Two new species of Pseudocoutierea (Decapoda Natantia, Palaemonidae) from the Colombian Caribbean. Crustaceana, 41 (2), 167 - 181. https: // doi. org / 10.1163 / 156854081 X 00219" type="journal article">Criales (1981)</bibRefCitation>
and should therefore not be considered as a valid deterministic character.
</paragraph>
<paragraph id="7F7ED8417D0DFFC135C5FBEB8BFA6B12" blockId="9.[151,1437,151,1149]" pageId="9" pageNumber="542">
<emphasis id="4DB504537D0DFFC135C5FBEB88BA6B37" bold="true" box="[199,465,1087,1112]" pageId="9" pageNumber="542">Phylogenetic analysis.</emphasis>
Characters and character states (
<tableCitation id="3243EDFA7D0DFFC1365AFBEB8AD66B37" box="[856,957,1087,1112]" captionStart="TABLE 1" captionStartId="9.[151,239,1196,1219]" captionText="TABLE 1. Character analysis and coding of character states of Pseudocoutierea species and outgroup taxon Coutierea agassizi." pageId="9" pageNumber="542">Tables 1</tableCitation>
, 
<tableCitation id="3243EDFA7D0DFFC136D2FB948AB46B37" box="[976,991,1088,1112]" captionStart="TABLE 2" captionStartId="13.[151,239,655,678]" captionText="TABLE 2. Data matrix for Pseudocoutierea species and outgroup taxon Coutierea agassizi used in the phylogenetic analysis." pageId="9" pageNumber="542">2</tableCitation>
) used are derived from the literature, material collected, and comparative material.
</paragraph>
<caption id="2BBE88C97D0DFFC13595FB7889986B8C" pageId="9" pageNumber="542">
<paragraph id="7F7ED8417D0DFFC13595FB7889986B8C" blockId="9.[151,1436,1195,1251]" pageId="9" pageNumber="542">
<emphasis id="4DB504537D0DFFC13595FB7888606BAC" bold="true" box="[151,267,1196,1219]" pageId="9" pageNumber="542">TABLE 1.</emphasis>
Character analysis and coding of character states of 
<taxonomicName id="B8C1A3C27D0DFFC1364BFB7F8A906BAC" authorityName="Holthuis" authorityYear="1951" box="[841,1019,1195,1219]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D0DFFC1364BFB7F8A906BAC" box="[841,1019,1195,1219]" italics="true" pageId="9" pageNumber="542">Pseudocoutierea</emphasis>
</taxonomicName>
species and outgroup taxon 
<taxonomicName id="B8C1A3C27D0DFFC13030FB7F89866B8C" authorityName="Coutiere" authorityYear="1901" class="Malacostraca" family="Palaemonidae" genus="Coutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="542" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="4DB504537D0DFFC13030FB7F89866B8C" italics="true" pageId="9" pageNumber="542">Coutierea agassizi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="7F7ED8417D0DFFC1359DFB2B8AC268BC" pageId="9" pageNumber="542">
<table id="0DC12AE17D0D0037359DFB2A8D3468BC" box="[159,1119,1278,2003]" gridcols="2" gridrows="18" pageId="9" pageNumber="542">
<tr id="C1F1DA037D0D0037359DFB2A8D346A7B" box="[159,1119,1278,1300]" gridrow="0" pageId="9" pageNumber="542">
<th id="8220B37F7D0D0037359DFB2A89AE6A7B" box="[159,197,1278,1300]" gridcol="0" gridrow="0" pageId="9" pageNumber="542">No.</th>
<th id="8220B37F7D0D003735E6FB2A8D346A7B" box="[228,1119,1278,1300]" gridcol="1" gridrow="0" pageId="9" pageNumber="542">Description</th>
</tr>
<tr id="C1F1DA037D0D0037359DFAFC8D346A51" box="[159,1119,1320,1342]" gridrow="1" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DFAFC89AE6A51" box="[159,197,1320,1342]" gridcol="0" gridrow="1" pageId="9" pageNumber="542">1</td>
<td id="8220B37F7D0D003735E6FAFC8D346A51" box="[228,1119,1320,1342]" gridcol="1" gridrow="1" pageId="9" pageNumber="542">Post-hepatic protuberance/spine present (0) or absent (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DFA858D346A08" box="[159,1119,1361,1383]" gridrow="2" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DFA8589AE6A08" box="[159,197,1361,1383]" gridcol="0" gridrow="2" pageId="9" pageNumber="542">2</td>
<td id="8220B37F7D0D003735E6FA858D346A08" box="[228,1119,1361,1383]" gridcol="1" gridrow="2" pageId="9" pageNumber="542">Supra-orbital eaves anterolaterally acute (0) or rounded (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DFAAE8D346AFF" box="[159,1119,1402,1424]" gridrow="3" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DFAAE89AE6AFF" box="[159,197,1402,1424]" gridcol="0" gridrow="3" pageId="9" pageNumber="542">3</td>
<td id="8220B37F7D0D003735E6FAAE8D346AFF" box="[228,1119,1402,1424]" gridcol="1" gridrow="3" pageId="9" pageNumber="542">Pleura of third pleonite posterolaterally acute (0) or rounded (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DFA708D346AD5" box="[159,1119,1444,1466]" gridrow="4" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DFA7089AE6AD5" box="[159,197,1444,1466]" gridcol="0" gridrow="4" pageId="9" pageNumber="542">4</td>
<td id="8220B37F7D0D003735E6FA708D346AD5" box="[228,1119,1444,1466]" gridcol="1" gridrow="4" pageId="9" pageNumber="542">Propodus of third pereiopod with distoventral spine (0) or without distoventral spine (0)</td>
</tr>
<tr id="C1F1DA037D0D0037359DFA198D346A8C" box="[159,1119,1485,1507]" gridrow="5" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DFA1989AE6A8C" box="[159,197,1485,1507]" gridcol="0" gridrow="5" pageId="9" pageNumber="542">5</td>
<td id="8220B37F7D0D003735E6FA198D346A8C" box="[228,1119,1485,1507]" gridcol="1" gridrow="5" pageId="9" pageNumber="542">Epigastric knob or tooth absent (0) or present (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DFA228D346963" box="[159,1119,1526,1548]" gridrow="6" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DFA2289AE6963" box="[159,197,1526,1548]" gridcol="0" gridrow="6" pageId="9" pageNumber="542">6</td>
<td id="8220B37F7D0D003735E6FA228D346963" box="[228,1119,1526,1548]" gridcol="1" gridrow="6" pageId="9" pageNumber="542">Moveable spine between tooth and blade of exopod of uropod absent (0) or present (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DF9F48D346959" box="[159,1119,1568,1590]" gridrow="7" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF9F489AE6959" box="[159,197,1568,1590]" gridcol="0" gridrow="7" pageId="9" pageNumber="542">7</td>
<td id="8220B37F7D0D003735E6F9F48D346959" box="[228,1119,1568,1590]" gridcol="1" gridrow="7" pageId="9" pageNumber="542">Pterygostomial sinus deep (0) or shallow (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DF99D8D346930" box="[159,1119,1609,1631]" gridrow="8" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF99D89AE6930" box="[159,197,1609,1631]" gridcol="0" gridrow="8" pageId="9" pageNumber="542">8</td>
<td id="8220B37F7D0D003735E6F99D8D346930" box="[228,1119,1609,1631]" gridcol="1" gridrow="8" pageId="9" pageNumber="542">Lash on caridean lobe on first maxilliped present (0) or absent (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DF9A68D3469E7" box="[159,1119,1650,1672]" gridrow="9" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF9A689AE69E7" box="[159,197,1650,1672]" gridcol="0" gridrow="9" pageId="9" pageNumber="542">9</td>
<td id="8220B37F7D0D003735E6F9A68D3469E7" box="[228,1119,1650,1672]" gridcol="1" gridrow="9" pageId="9" pageNumber="542">Second maxilliped without exopod (0) or with exopod (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DF9488D3469DD" box="[159,1119,1692,1714]" gridrow="10" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF94889AE69DD" box="[159,197,1692,1714]" gridcol="0" gridrow="10" pageId="9" pageNumber="542">10</td>
<td id="8220B37F7D0D003735E6F9488D3469DD" box="[228,1119,1692,1714]" gridcol="1" gridrow="10" pageId="9" pageNumber="542">Thirs maxilliped without arthrobranch (0) or with arthrobranch (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DF9118D3469B4" box="[159,1119,1733,1755]" gridrow="11" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF91189AE69B4" box="[159,197,1733,1755]" gridcol="0" gridrow="11" pageId="9" pageNumber="542">11</td>
<td id="8220B37F7D0D003735E6F9118D3469B4" box="[228,1119,1733,1755]" gridcol="1" gridrow="11" pageId="9" pageNumber="542">Acuminate lobe immediately below antennal spine absent (0) or present (1)</td>
</tr>
<tr id="C1F1DA037D0D0037359DF93A8D34686B" box="[159,1119,1774,1796]" gridrow="12" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF93A89AE686B" box="[159,197,1774,1796]" gridcol="0" gridrow="12" pageId="9" pageNumber="542">12</td>
<td id="8220B37F7D0D003735E6F93A8D34686B" box="[228,1119,1774,1796]" gridcol="1" gridrow="12" pageId="9" pageNumber="542">Pterygostomial spine present (0) or absent (1);</td>
</tr>
<tr id="C1F1DA037D0D0037359DF8CC8D346841" box="[159,1119,1816,1838]" gridrow="13" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF8CC89AE6841" box="[159,197,1816,1838]" gridcol="0" gridrow="13" pageId="9" pageNumber="542">13</td>
<td id="8220B37F7D0D003735E6F8CC8D346841" box="[228,1119,1816,1838]" gridcol="1" gridrow="13" pageId="9" pageNumber="542">Rostrum curved up (0) or straight (1);</td>
</tr>
<tr id="C1F1DA037D0D0037359DF8958D346838" box="[159,1119,1857,1879]" gridrow="14" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF89589AE6838" box="[159,197,1857,1879]" gridcol="0" gridrow="14" pageId="9" pageNumber="542">14</td>
<td id="8220B37F7D0D003735E6F8958D346838" box="[228,1119,1857,1879]" gridcol="1" gridrow="14" pageId="9" pageNumber="542">Carapace depressed (0) or not depressed (1);</td>
</tr>
<tr id="C1F1DA037D0D0037359DF8BE8D3468EF" box="[159,1119,1898,1920]" gridrow="15" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF8BE89AE68EF" box="[159,197,1898,1920]" gridcol="0" gridrow="15" pageId="9" pageNumber="542">15</td>
<td id="8220B37F7D0D003735E6F8BE8D3468EF" box="[228,1119,1898,1920]" gridcol="1" gridrow="15" pageId="9" pageNumber="542">Prolongation of sixth pleonite elongated (0) or triangular (1);</td>
</tr>
<tr id="C1F1DA037D0D0037359DF8408D3468C5" box="[159,1119,1940,1962]" gridrow="16" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF84089AE68C5" box="[159,197,1940,1962]" gridcol="0" gridrow="16" pageId="9" pageNumber="542">16</td>
<td id="8220B37F7D0D003735E6F8408D3468C5" box="[228,1119,1940,1962]" gridcol="1" gridrow="16" pageId="9" pageNumber="542">Mandibular process with four teeth (0) or three teeth (1);</td>
</tr>
<tr id="C1F1DA037D0D0037359DF8698D3468BC" box="[159,1119,1981,2003]" gridrow="17" pageId="9" pageNumber="542">
<td id="8220B37F7D0D0037359DF86989AE68BC" box="[159,197,1981,2003]" gridcol="0" gridrow="17" pageId="9" pageNumber="542">17</td>
<td id="8220B37F7D0D003735E6F8698D3468BC" box="[228,1119,1981,2003]" gridcol="1" gridrow="17" pageId="9" pageNumber="542">Lateral margin of scaphocerite convex (0), straight (1), or concave (2)</td>
</tr>
</table>
</paragraph>
<caption id="2BBE88C97D0EFFC23595F8AE8A6C6883" httpUri="https://zenodo.org/record/1242215/files/figure.png" pageId="10" pageNumber="543" startId="10.[151,250,1914,1936]" targetBox="[238,1384,214,1885]" targetPageId="10">
<paragraph id="7F7ED8417D0EFFC23595F8AE8A6C6883" blockId="10.[151,1436,1914,2028]" pageId="10" pageNumber="543">
<emphasis id="4DB504537D0EFFC23595F8AE887868E0" bold="true" box="[151,275,1914,1936]" pageId="10" pageNumber="543">FIGURE 3.</emphasis>
<taxonomicName id="B8C1A3C27D0EFFC23419F8AE8B2068FF" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[283,587,1914,1936]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="543" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0EFFC23419F8AE8B2068FF" box="[283,587,1914,1936]" italics="true" pageId="10" pageNumber="543">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0EFFC23750F8AE8BC868E0" box="[594,675,1914,1935]" pageId="10" pageNumber="543" rank="species">
<emphasis id="4DB504537D0EFFC23750F8AE8BC868E0" bold="true" box="[594,675,1914,1935]" pageId="10" pageNumber="543">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype, RMNH.CRUS.D.57273. A, left first pereiopod, anterior view; B, idem, posterior view; C, idem, chela; D, minor left second pereiopod; E, idem, chela; F, major right second pereiopod, ventral view; G, idem, dorsal view; H, idem chela; I, right third pereiopod; J, idem, dactylus. Scale bar: A, B, I = 1mm; C, E = 0.2mm; D, H = 0.6mm; F, G = 2mm; H = 0.5mm; J = 0.3mm.
</paragraph>
</caption>
<caption id="2BBE88C97D0FFFC33595FA458C2B696D" httpUri="https://zenodo.org/record/1242217/files/figure.png" pageId="11" pageNumber="544" startId="11.[151,250,1425,1447]" targetBox="[305,1275,201,1383]" targetPageId="11">
<paragraph id="7F7ED8417D0FFFC33595FA458C2B696D" blockId="11.[151,1436,1425,1538]" pageId="11" pageNumber="544">
<emphasis id="4DB504537D0FFFC33595FA4588656AC9" bold="true" box="[151,270,1425,1447]" pageId="11" pageNumber="544">FIGURE 4</emphasis>
. 
<taxonomicName id="B8C1A3C27D0FFFC3341EFA458B266AC8" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[284,589,1425,1447]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0FFFC3341EFA458B266AC8" box="[284,589,1425,1447]" italics="true" pageId="11" pageNumber="544">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0FFFC33756FA458BCC6AC9" box="[596,679,1425,1446]" pageId="11" pageNumber="544" rank="species">
<emphasis id="4DB504537D0FFFC33756FA458BCC6AC9" bold="true" box="[596,679,1425,1446]" pageId="11" pageNumber="544">sp. nov.</emphasis>
</taxonomicNameLabel>
female paratype, third pereiopod, RMNH.CRUS.D.57273. A, decalcified part on flexor border of dactylus; B, flexor border between propodus and dactylus, showing the two infero-distal spines, ventral view; C, idem, lateral view; D, tip of dactylus with suture between corpus and unguis; E, merus and carpus lateral view, showing meral swelling (arrow), lateral view; F, meral swelling (arrow) on flexor border of merus, ventrolateral view.
</paragraph>
</caption>
<paragraph id="7F7ED8417D0FFFC335C5F99F8D8E69BF" blockId="11.[151,1436,1611,2032]" pageId="11" pageNumber="544">
The species 
<taxonomicName id="B8C1A3C27D0FFFC33469F9988A7B690B" authority="Coutiere, 1901" authorityName="Coutiere" authorityYear="1901" box="[363,784,1611,1636]" class="Malacostraca" family="Palaemonidae" genus="Coutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="agassizi">
<emphasis id="4DB504537D0FFFC33469F9988B23690C" box="[363,584,1612,1635]" italics="true" pageId="11" pageNumber="544">Coutierea agassizi</emphasis>
Coutière, 1901
</taxonomicName>
was selected as the outgroup taxon. It resembles 
<taxonomicName id="B8C1A3C27D0FFFC33595F9BB883969E8" authorityName="Holthuis" authorityYear="1951" box="[151,338,1647,1671]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D0FFFC33595F9BB883969E8" box="[151,338,1647,1671]" italics="true" pageId="11" pageNumber="544">Pseudocoutierea</emphasis>
</taxonomicName>
in the presence of a pterygostomial sinus, the lack of an exopod on the maxillipeds (although an exopod is present on the second maxilliped in 
<taxonomicName id="B8C1A3C27D0FFFC337A2F9408A7269C3" authorityName="Criales" authorityYear="1981" box="[672,793,1684,1708]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="conchae">
<emphasis id="4DB504537D0FFFC337A2F9408A7269C3" box="[672,793,1684,1708]" italics="true" pageId="11" pageNumber="544">P. conchae</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8C1A3C27D0FFFC33652F9408AA769C3" authorityName="Criales" authorityYear="1981" box="[848,972,1684,1708]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="edentata">
<emphasis id="4DB504537D0FFFC33652F9408AA769C3" box="[848,972,1684,1708]" italics="true" pageId="11" pageNumber="544">P. edentata</emphasis>
</taxonomicName>
), the presence of an orbital eave, a meral swelling on the third pereiopod and a strong lateral carina on the carapace near the ventral margin.
</paragraph>
<paragraph id="7F7ED8417D0FFFC435C5F90F884A6FBA" blockId="11.[151,1436,1611,2032]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="545" pageId="11" pageNumber="544">
<figureCitation id="E7FAC4C47D0FFFC335C5F90F884E699B" box="[199,293,1755,1780]" captionStart="FIGURE 6. 50" captionStartId="12.[151,250,1765,1787]" captionTargetBox="[394,1212,1162,1694]" captionTargetId="figure@12.[340,1247,1134,1734]" captionTargetPageId="12" captionText="FIGURE 6. 50% majority rule consensus tree of Pseudocoutierea species; outgroup Coutierea agassizi (Coutière, 1901); tree length = 28, Consistency Index (CI) = 0.6429." httpUri="https://zenodo.org/record/1242221/files/figure.png" pageId="11" pageNumber="544">Figure 6</figureCitation>
shows the 50% majority rule consensus tree, which has the same topology as the strict consensus tree, of the genus 
<taxonomicName id="B8C1A3C27D0FFFC33425F92B88896878" authorityName="Holthuis" authorityYear="1951" box="[295,482,1791,1815]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="genus">
<emphasis id="4DB504537D0FFFC33425F92B88896878" box="[295,482,1791,1815]" italics="true" pageId="11" pageNumber="544">Pseudocoutierea</emphasis>
</taxonomicName>
. The close relationship between 
<taxonomicName id="B8C1A3C27D0FFFC33651F8D48A966878" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[851,1021,1791,1815]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0FFFC33651F8D48A966878" box="[851,1021,1791,1815]" italics="true" pageId="11" pageNumber="544">P. acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0FFFC33106F92B8D366877" box="[1028,1117,1791,1816]" pageId="11" pageNumber="544" rank="species">
<emphasis id="4DB504537D0FFFC33106F92B8D366877" bold="true" box="[1028,1117,1791,1816]" pageId="11" pageNumber="544">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="B8C1A3C27D0FFFC33196F8D48C6F6878" authorityName="Holthuis" authorityYear="1951" box="[1172,1284,1791,1815]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="4DB504537D0FFFC33196F8D48C6F6878" box="[1172,1284,1791,1815]" italics="true" pageId="11" pageNumber="544">P. elegans</emphasis>
</taxonomicName>
is interesting as 
<taxonomicName id="B8C1A3C27D0FFFC335B5F8F088426853" authorityName="Holthuis" authorityYear="1951" box="[183,297,1828,1852]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="4DB504537D0FFFC335B5F8F088426853" box="[183,297,1828,1852]" italics="true" pageId="11" pageNumber="544">P. elegans</emphasis>
</taxonomicName>
is known from the East Pacific and is separated from the Caribbean 
<taxonomicName id="B8C1A3C27D0FFFC3313CF8F08D826853" authority="Olthof &amp; Becking &amp; Fransen, 2018" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[1086,1257,1828,1852]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="acutidorsata" status="sp. nov.">
<emphasis id="4DB504537D0FFFC3313CF8F08D826853" box="[1086,1257,1828,1852]" italics="true" pageId="11" pageNumber="544">P. acutidorsata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5686B9287D0FFFC331F0F8F78C266853" box="[1266,1357,1827,1852]" pageId="11" pageNumber="544" rank="species">
<emphasis id="4DB504537D0FFFC331F0F8F78C266853" bold="true" box="[1266,1357,1827,1852]" pageId="11" pageNumber="544">sp. nov.</emphasis>
</taxonomicNameLabel>
by the 
<collectingRegion id="BD0516A37D0FFFC33595F89C899B6830" box="[151,240,1864,1887]" country="Panama" name="Panama" pageId="11" pageNumber="544">
<collectingCountry id="07D698D17D0FFFC33595F89C899B6830" box="[151,240,1864,1887]" name="Panama" pageId="11" pageNumber="544">Panama</collectingCountry>
</collectingRegion>
Isthmus. 
<taxonomicName id="B8C1A3C27D0FFFC3345FF8938BCD6830" authorityName="Ramos-Tafur &amp; Lemaitre" authorityYear="2016" box="[349,678,1863,1887]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="stephanieae">
<emphasis id="4DB504537D0FFFC3345FF8938BCD6830" box="[349,678,1863,1887]" italics="true" pageId="11" pageNumber="544">Pseudocoutierea stephanieae</emphasis>
</taxonomicName>
stands apart of most other species, partly because of the absence of a deep pterygostomial sinus which is present in all of the other species. 
<taxonomicName id="B8C1A3C27D0FFFC336CCF8B88D4C68EB" authorityName="De Grave" authorityYear="2007" box="[974,1063,1900,1924]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="dotae">
<emphasis id="4DB504537D0FFFC336CCF8B88D4C68EB" box="[974,1063,1900,1924]" italics="true" pageId="11" pageNumber="544">P. dotae</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8C1A3C27D0FFFC33161F8B88DAB68EC" authorityName="d'Udekem d'Acoz" authorityYear="2000" box="[1123,1216,1900,1923]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="wirtzi">
<emphasis id="4DB504537D0FFFC33161F8B88DAB68EC" box="[1123,1216,1900,1923]" italics="true" pageId="11" pageNumber="544">P. wirtzi</emphasis>
</taxonomicName>
seem to be closely related which is in conformation with the remark by 
<bibRefCitation id="1B50A5B07D0FFFC337E5F8448ACE68C7" box="[743,933,1936,1960]" pageId="11" pageNumber="544" refId="ref9542" refString="De Grave, S. (2007) A new species of Pseudocoutierea Holthuis from the Caribbean coast of Panama (Crustacea, Decapoda, Palaemonidae), with a key to the genus. Zootaxa, 37 (1397), 29 - 37. https: // doi. org / 10.11646 / zootaxa. 1397.4" type="journal article">De Grave (2007)</bibRefCitation>
on their close affinity. The two species from Caribbean 
<collectingCountry id="07D698D17D0FFFC33413F867881468A3" box="[273,383,1971,1996]" name="Colombia" pageId="11" pageNumber="544">Colombia</collectingCountry>
(
<taxonomicName id="B8C1A3C27D0FFFC33492F8608B6368A3" authorityName="Criales" authorityYear="1981" box="[400,520,1972,1996]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="conchae">
<emphasis id="4DB504537D0FFFC33492F8608B6368A3" box="[400,520,1972,1996]" italics="true" pageId="11" pageNumber="544">P. conchae</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8C1A3C27D0FFFC3373DF8608BD068A3" authorityName="Criales" authorityYear="1981" box="[575,699,1972,1996]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="544" phylum="Arthropoda" rank="species" species="edentata">
<emphasis id="4DB504537D0FFFC3373DF8608BD068A3" box="[575,699,1972,1996]" italics="true" pageId="11" pageNumber="544">P. edentata</emphasis>
</taxonomicName>
) also seem closely related. From this phylogenetic reconstruction it seems that the epigastric tooth and acute supra-orbital eaves evolved and got lost again independently several times. The variation in the latter character within 
<taxonomicName id="B8C1A3C27D08FFC43600FF4C8D326FDF" authorityName="Olthof &amp; Becking &amp; Fransen" authorityYear="2018" box="[770,1113,152,176]" class="Malacostraca" family="Palaemonidae" genus="Pseudocoutierea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="545" phylum="Arthropoda" rank="species" species="acutidorsata">
<emphasis id="4DB504537D08FFC43600FF4C8D326FDF" box="[770,1113,152,176]" italics="true" pageId="12" pageNumber="545">Pseudocoutierea acutidorsata</emphasis>
</taxonomicName>
is in agreement with this observation.
</paragraph>
</subSubSection>
</treatment>
</document>