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<document id="81F22AE8F35663F7A67744ED9DF63399" ID-CLB-Dataset="45651" ID-DOI="10.11646/zootaxa.3802.4.5" ID-GBIF-Dataset="71e8a84e-1a9b-4849-9a09-ec2023643364" ID-ISSN="1175-5326" ID-Zenodo-Dep="249605" ID-ZooBank="4095CA09-8EA4-4941-8286-32E95F0206AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427480340" checkinUser="plazi" docAuthor="Carmona, Leila, Pola, Marta, Gosliner, Terrence M. &amp; Cervera, Juan Lucas" docDate="2014" docId="ED36FA51A020FF9FFF1BFA87E06DF8A3" docLanguage="en" docName="zt03802p514.pdf" docOrigin="Zootaxa 3802 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baeolidia" docType="treatment" docVersion="9" lastPageNumber="506" masterDocId="110F8229A03CFF82FF8CFFB7E275FFB7" masterDocTitle="Review of Baeolidia, the largest genus of Aeolidiidae (Mollusca: Nudibranchia), with the description of five new species" masterLastPageNumber="514" masterPageNumber="477" pageNumber="505" updateTime="1698330923557" updateUser="ExternalLinkService">
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<mods:title id="4620F40CBBEB710C2C28062203AF997A">Review of Baeolidia, the largest genus of Aeolidiidae (Mollusca: Nudibranchia), with the description of five new species</mods:title>
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<mods:namePart id="ED00D35B39253E9DCB44BB4AD411584D">Carmona, Leila</mods:namePart>
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<mods:namePart id="D0C63957A5BCCDD8DEB99990D9621FC6">Pola, Marta</mods:namePart>
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<mods:namePart id="6E9CA8A686B76DEB20BD8F44B7BDCED7">Gosliner, Terrence M.</mods:namePart>
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<mods:namePart id="4669D1AD82AD371D31BAB57BC11DFECE">Cervera, Juan Lucas</mods:namePart>
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<mods:title id="C072B12391EC48BAFD7B85C3BE23FD59">Zootaxa</mods:title>
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<mods:date id="FDA7C984BD7F0E10D767963A1B6B1324">2014</mods:date>
<mods:detail id="7A9B7DA0093BE5E3AC477D5196F6EBF2" type="volume">
<mods:number id="E7EC5FA58BAB9FEC4571D0CFE3E8EF86">3802</mods:number>
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<mods:classification id="AAA07E40394E479EEB84A41B8D3E19A0">journal article</mods:classification>
<mods:identifier id="C4D3E02BAAA48EC63286A0C069536999" type="CLB-Dataset">45651</mods:identifier>
<mods:identifier id="CD75CC1ECFE8A179BC3ADD019A0568EB" type="DOI">10.11646/zootaxa.3802.4.5</mods:identifier>
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<treatment id="ED36FA51A020FF9FFF1BFA87E06DF8A3" ID-DOI="http://doi.org/10.5281/zenodo.6134239" ID-GBIF-Taxon="119429018" ID-Zenodo-Dep="6134239" LSID="urn:lsid:plazi:treatment:ED36FA51A020FF9FFF1BFA87E06DF8A3" httpUri="http://treatment.plazi.org/id/ED36FA51A020FF9FFF1BFA87E06DF8A3" lastPageId="29" lastPageNumber="506" pageId="28" pageNumber="505">
<subSubSection id="2D8518CCA020FF9FFF1BFA87E06DF8A3" lastPageId="29" lastPageNumber="506" pageId="28" pageNumber="505" type="nomenclature">
<paragraph id="65204B47A020FF9EFF1BFA87E0CAFAFE" blockId="28.[151,1436,1328,2038]" box="[151,703,1328,1353]" pageId="28" pageNumber="505">
<heading id="3E68FC2BA020FF9EFF1BFA87E0CAFAFE" bold="true" box="[151,703,1328,1353]" fontSize="10" level="2" pageId="28" pageNumber="505" reason="2">
<emphasis id="57EB9755A020FF9EFF1BFA87E0CAFAFE" bold="true" box="[151,703,1328,1353]" pageId="28" pageNumber="505">
<taxonomicName id="A29F30C4A020FF9EFF1BFA87E373FAFE" ID-CoL="7NQCC" box="[151,262,1328,1353]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFF1BFA87E373FAFE" bold="true" box="[151,262,1328,1353]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
vs
<emphasis id="57EB9755A020FF9EFEA0FA87E381FAFE" bold="true" box="[300,500,1328,1353]" italics="true" pageId="28" pageNumber="505">
<taxonomicName id="A29F30C4A020FF9EFEA0FA87E3FEFAFE" box="[300,395,1328,1353]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Spurilla</taxonomicName>
,
<taxonomicName id="A29F30C4A020FF9EFE1BFA87E381FAFE" box="[407,500,1328,1353]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Berghia</taxonomicName>
</emphasis>
and
<taxonomicName id="A29F30C4A020FF9EFDBCFA87E0CAFAFE" box="[560,703,1328,1353]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFDBCFA87E0CAFAFE" bold="true" box="[560,703,1328,1353]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="65204B47A020FF9EFF4BFAE3E741FA4A" blockId="28.[151,1436,1328,2038]" pageId="28" pageNumber="505">
The aeolidid genera
<emphasis id="57EB9755A020FF9EFE39FAE3E00CFADA" box="[437,633,1364,1389]" italics="true" pageId="28" pageNumber="505">
<taxonomicName id="A29F30C4A020FF9EFE39FAE3E067FADA" box="[437,530,1364,1389]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Spurilla</taxonomicName>
,
<taxonomicName id="A29F30C4A020FF9EFDACFAE3E00CFADA" box="[544,633,1364,1389]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Berghia</taxonomicName>
</emphasis>
and
<taxonomicName id="A29F30C4A020FF9EFD3AFAE3E156FADA" box="[694,803,1364,1389]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFD3AFAE3E156FADA" box="[694,803,1364,1389]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
were considered to be closely related. Indeed,
<bibRefCitation id="010E36B6A020FF9EFAC4FAE3E294FA26" author="Odhner" pageId="28" pageNumber="505" refString="Odhner, N. H. (1939) Opisthobranchiate Mollusca from the western and northern coasts of Norway. Det Kgl Norske Vidensk Selsk Skrift, 1, 1 - 93." type="journal article" year="1939">Odhner (1939)</bibRefCitation>
placed
<taxonomicName id="A29F30C4A020FF9EFECCFACFE3EFFA26" box="[320,410,1400,1425]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFECCFACFE3EFFA26" box="[320,410,1400,1425]" italics="true" pageId="28" pageNumber="505">Spurilla</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A020FF9EFE5BFACFE045FA26" box="[471,560,1400,1425]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFE5BFACFE045FA26" box="[471,560,1400,1425]" italics="true" pageId="28" pageNumber="505">Berghia</emphasis>
</taxonomicName>
in a new family called
<taxonomicName id="A29F30C4A020FF9EFCC0FACFE1BFFA26" box="[844,970,1400,1425]" class="Gastropoda" family="Aeolidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="family">Spurillidae</taxonomicName>
, including
<taxonomicName id="A29F30C4A020FF9EFBC1FACFE6CFFA26" box="[1101,1210,1400,1425]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFBC1FACFE6CFFA26" box="[1101,1210,1400,1425]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
ten years later and overlooking
<taxonomicName id="A29F30C4A020FF9EFEA8FA2BE3D8FA02" box="[292,429,1436,1461]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFEA8FA2BE3D8FA02" box="[292,429,1436,1461]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
(Odhner in
<bibRefCitation id="010E36B6A020FF9EFDB8FA29E0C3FA02" author="Franc" box="[564,694,1436,1461]" pageId="28" pageNumber="505" refString="Franc, A. (1968) Sous-classe des opisthobranches. In: G r asse, P. P. (E d.), Traite de Zoologie. Masson, Paris, pp. 608 - 893." type="book chapter" year="1968">Franc 1968</bibRefCitation>
). In the manner of Odhner (in
<bibRefCitation id="010E36B6A020FF9EFB8BFA29E6FCFA02" author="Franc" box="[1031,1161,1436,1461]" pageId="28" pageNumber="505" refString="Franc, A. (1968) Sous-classe des opisthobranches. In: G r asse, P. P. (E d.), Traite de Zoologie. Masson, Paris, pp. 608 - 893." type="book chapter" year="1968">Franc 1968</bibRefCitation>
), many authors regarded
<taxonomicName id="A29F30C4A020FF9EFF1BFA77E355FA6E" box="[151,288,1472,1497]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFF1BFA77E355FA6E" box="[151,288,1472,1497]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="A29F30C4A020FF9EFDB8FA77E0D4FA6E" box="[564,673,1472,1497]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFDB8FA77E0D4FA6E" box="[564,673,1472,1497]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
. Recently, the molecular phylogeny of
<taxonomicName id="A29F30C4A020FF9EFBECFA77E2ADFA4A" authority="Carmona et al. 2013" authorityName="Carmona et al." authorityYear="2013" class="Gastropoda" family="Aeolidiidae" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="family">
Aeolidiidae (
<bibRefCitation id="010E36B6A020FF9EFB7EFA76E2A4FA4A" author="Carmona" pageId="28" pageNumber="505" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2013) A tale that morphology fails to tell: a molecular phylogeny of Aeolidiidae (Aeolidida, Nudibranchia, Gastropoda). PloS One, 8, 5. http: // dx. doi. org / 10.1371 / journal. pone. 0063000" type="journal article" year="2013">
Carmona
<emphasis id="57EB9755A020FF9EFAEFFA75E7E0FA6E" box="[1379,1429,1472,1497]" italics="true" pageId="28" pageNumber="505">et al</emphasis>
. 2013
</bibRefCitation>
)
</taxonomicName>
rejected the validity of
<taxonomicName id="A29F30C4A020FF9EFE6FFA53E08BFA4A" authority="Odhner" authorityName="Odhner" box="[483,766,1508,1533]" class="Gastropoda" family="Aeolidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="family">Spurillidae sensu Odhner</taxonomicName>
and the synonymy of
<taxonomicName id="A29F30C4A020FF9EFC7BFA53E6F5FA4A" box="[1015,1152,1508,1533]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFC7BFA53E6F5FA4A" box="[1015,1152,1508,1533]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
with
<taxonomicName id="A29F30C4A020FF9EFB4CFA53E758FA4A" box="[1216,1325,1508,1533]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFB4CFA53E758FA4A" box="[1216,1325,1508,1533]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="65204B47A020FF9EFF1BF9BEE609F996" blockId="28.[151,1436,1328,2038]" box="[151,1148,1544,1569]" pageId="28" pageNumber="505">However, the morphological differences among these genera have not been well defined.</paragraph>
<paragraph id="65204B47A020FF9FFF4BF99AE1F7FED2" blockId="28.[151,1436,1328,2038]" lastBlockId="29.[151,1436,151,501]" lastPageId="29" lastPageNumber="506" pageId="28" pageNumber="505">
Despite difficulties in finding synapomorphies for some genera within
<taxonomicName id="A29F30C4A020FF9EFBB6F99AE6B5F9F1" box="[1082,1216,1581,1606]" class="Gastropoda" family="Aeolidiidae" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="family">Aeolidiidae</taxonomicName>
, it is possible to differentiate
<emphasis id="57EB9755A020FF9EFEA0F9E7E001F9DE" box="[300,628,1616,1641]" italics="true" pageId="28" pageNumber="505">
<taxonomicName id="A29F30C4A020FF9EFEA0F9E7E3E9F9DE" box="[300,412,1616,1641]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Baeolidia</taxonomicName>
,
<taxonomicName id="A29F30C4A020FF9EFE21F9E7E07FF9DE" box="[429,522,1616,1641]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Spurilla</taxonomicName>
,
<taxonomicName id="A29F30C4A020FF9EFD97F9E7E001F9DE" box="[539,628,1616,1641]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Berghia</taxonomicName>
</emphasis>
and
<taxonomicName id="A29F30C4A020FF9EFD3AF9E7E14AF9DE" box="[694,831,1616,1641]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFD3AF9E7E14AF9DE" box="[694,831,1616,1641]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
based on some features. Regarding body size and morphology, we could divide these genera in two groups:
<taxonomicName id="A29F30C4A020FF9EFCB4F9C2E1E7F939" box="[824,914,1653,1678]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFCB4F9C2E1E7F939" box="[824,914,1653,1678]" italics="true" pageId="28" pageNumber="505">Spurilla</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A020FF9EFC42F9C2E64EF939" box="[974,1083,1653,1678]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFC42F9C2E64EF939" box="[974,1083,1653,1678]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
, and
<taxonomicName id="A29F30C4A020FF9EFBF2F9C2E6A2F939" box="[1150,1239,1653,1678]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFBF2F9C2E6A2F939" box="[1150,1239,1653,1678]" italics="true" pageId="28" pageNumber="505">Berghia</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A020FF9EFA9FF9C2E7E9F939" box="[1299,1436,1653,1678]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFA9FF9C2E7E9F939" box="[1299,1436,1653,1678]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
(
<figureCitation id="FDA457C2A020FF9EFF13F92FE283F906" box="[159,246,1688,1713]" captionStart="FIGURE 15" captionStartId="27.[151,250,1869,1891]" captionTargetBox="[289,1324,452,1834]" captionTargetId="figure@27.[231,1355,423,1851]" captionTargetPageId="27" captionText="FIGURE 15. Schematic representation of differences of body shape among the genera Spurilla (A), drawing of Ascensión Cespedosa; Berghia (B), drawing of Ascensión Cespedosa; Limenandra (C), taken from Haefelfinger &amp; Stamm (1958); and Baeolidia (D), taken from Rudman (1982)." httpUri="https://zenodo.org/record/249619/files/figure.png" pageId="28" pageNumber="505">Fig. 15</figureCitation>
). The body of the members of the first group is quite broad although species of
<taxonomicName id="A29F30C4A020FF9EFB2BF92FE761F906" box="[1191,1300,1688,1713]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFB2BF92FE761F906" box="[1191,1300,1688,1713]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
are usually smaller and wider than species of
<taxonomicName id="A29F30C4A020FF9EFD9CF90AE01FF961" box="[528,618,1725,1750]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFD9CF90AE01FF961" box="[528,618,1725,1750]" italics="true" pageId="28" pageNumber="505">Spurilla</emphasis>
</taxonomicName>
. On the other hand,
<taxonomicName id="A29F30C4A020FF9EFCC0F90AE1D0F961" box="[844,933,1725,1750]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFCC0F90AE1D0F961" box="[844,933,1725,1750]" italics="true" pageId="28" pageNumber="505">Berghia</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A020FF9EFC56F90AE616F961" box="[986,1123,1725,1750]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFC56F90AE616F961" box="[986,1123,1725,1750]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
species are both slender but
<taxonomicName id="A29F30C4A020FF9EFF1BF957E355F94E" box="[151,288,1760,1785]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFF1BF957E355F94E" box="[151,288,1760,1785]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
species are more cylindrical. The rhinophorial ornamentation is also very useful to separate these genera. Although
<taxonomicName id="A29F30C4A020FF9EFEEDF8B2E3CFF8A9" box="[353,442,1797,1822]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFEEDF8B2E3CFF8A9" box="[353,442,1797,1822]" italics="true" pageId="28" pageNumber="505">Berghia</emphasis>
</taxonomicName>
,
<taxonomicName id="A29F30C4A020FF9EFE45F8B2E043F8A9" box="[457,566,1797,1822]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFE45F8B2E043F8A9" box="[457,566,1797,1822]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A020FF9EFDE1F8B2E083F8A9" box="[621,758,1797,1822]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFDE1F8B2E083F8A9" box="[621,758,1797,1822]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
were supposed to have papillate rhinophores, we could find consistent differences in the morphology of the papillae. While the rhinophorial papillae of
<taxonomicName id="A29F30C4A020FF9EFB30F89FE760F8F6" box="[1212,1301,1832,1857]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFB30F89FE760F8F6" box="[1212,1301,1832,1857]" italics="true" pageId="28" pageNumber="505">Berghia</emphasis>
</taxonomicName>
are usually asymmetrical, being more or less rounded on posterior side, and elongate and perpendicular to rhinophores on their lateral sides (see figure 2 of
<bibRefCitation id="010E36B6A020FF9EFE59F8C6E0B8F83E" author="Carmona" box="[469,717,1904,1929]" pageId="28" pageNumber="505" refString="Carmona, L., Pola, M, Gosliner, T. M., Cervera, J. L. (2014 b) The Atlantic-Mediterranean genus Berghia (Mollusca, Nudibranchia, Aeolidiidae). Taxonomic review and phylogenetic analysis. Journal of Molluscan Studies. [in press]" type="book" year="2014" yearSuffix="b">
Carmona
<emphasis id="57EB9755A020FF9EFDC8F8C5E001F83E" box="[580,628,1904,1929]" italics="true" pageId="28" pageNumber="505">et al</emphasis>
. 2014b
</bibRefCitation>
), in
<taxonomicName id="A29F30C4A020FF9EFD71F8C7E1F3F83E" box="[765,902,1904,1929]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFD71F8C7E1F3F83E" box="[765,902,1904,1929]" italics="true" pageId="28" pageNumber="505">Limenandra</emphasis>
</taxonomicName>
they are bifurcated (see figure 4 of
<bibRefCitation id="010E36B6A020FF9EFA96F8C6E37EF81A" author="Carmona" pageId="28" pageNumber="505" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2014 c) The end of a long controversy: systematics of the genus Limenandra (Mollusca: Nudibranchia: Aeolidiidae). Helgoland Marine Research, 68, 37 - 48. http: // dx. doi. org / 10.1007 / s 10152 - 013 - 0367 - y" type="journal article" year="2014" yearSuffix="c">
Carmona
<emphasis id="57EB9755A020FF9EFA04F8C5E2D8F819" italics="true" pageId="28" pageNumber="505">et al</emphasis>
., 2014c
</bibRefCitation>
). In
<taxonomicName id="A29F30C4A020FF9EFEB0F822E3DCF819" box="[316,425,1941,1966]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFEB0F822E3DCF819" box="[316,425,1941,1966]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
, we find more variability in the morphology of these papillae and in the ornamentation of the rhinophores in general (
<figureCitation id="FDA457C2A020FF9EFE43F80FE063F866" box="[463,534,1976,2001]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[151,1436,373,1863]" captionTargetId="figure@3.[151,1436,373,1865]" captionTargetPageId="3" captionText="FIGURE 2. Photographs of the living animals. A, B, Baeolidia moebii: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05 / 63447; (C) Baeolidia macleayi, specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E) Baeolidia japonica: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F) Baeolidia ransoni, specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208." httpUri="https://zenodo.org/record/249607/files/figure.png" pageId="28" pageNumber="505">Fig. 2</figureCitation>
). However, due to the particular rhinophores together with other morphological features,
<taxonomicName id="A29F30C4A020FF9EFF73F86AE319F841" box="[255,364,2013,2038]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFF73F86AE319F841" box="[255,364,2013,2038]" italics="true" pageId="28" pageNumber="505">Baeolidia</emphasis>
</taxonomicName>
cannot be confounded with
<emphasis id="57EB9755A020FF9EFD38F86AE1DBF841" box="[692,942,2013,2038]" italics="true" pageId="28" pageNumber="505">
<taxonomicName id="A29F30C4A020FF9EFD38F86AE164F841" box="[692,785,2013,2038]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Spurilla</taxonomicName>
,
<taxonomicName id="A29F30C4A020FF9EFC92F86AE1DFF841" box="[798,938,2013,2038]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">Limenandra</taxonomicName>
,
</emphasis>
and
<taxonomicName id="A29F30C4A020FF9EFC65F86AE637F841" box="[1001,1090,2013,2038]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="28" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A020FF9EFC65F86AE637F841" box="[1001,1090,2013,2038]" italics="true" pageId="28" pageNumber="505">Berghia</emphasis>
</taxonomicName>
. Although the arrangement of the cerata is not appropriate to separate genera in
<taxonomicName id="A29F30C4A021FF9FFD4AFF20E63EFF07" authority="Carmona et al. 2013" authorityName="Carmona et al." authorityYear="2013" box="[710,1099,151,176]" class="Gastropoda" family="Aeolidiidae" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="family">
Aeolidiidae (
<bibRefCitation id="010E36B6A021FF9FFCD4FF2FE636FF07" author="Carmona" box="[856,1091,151,176]" pageId="29" pageNumber="506" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2013) A tale that morphology fails to tell: a molecular phylogeny of Aeolidiidae (Aeolidida, Nudibranchia, Gastropoda). PloS One, 8, 5. http: // dx. doi. org / 10.1371 / journal. pone. 0063000" type="journal article" year="2013">
Carmona
<emphasis id="57EB9755A021FF9FFC4BFF2EE675FF07" box="[967,1024,151,176]" italics="true" pageId="29" pageNumber="506">et al.</emphasis>
2013
</bibRefCitation>
)
</taxonomicName>
, the morphology of the cerata may give some information. Our research suggests that all the
<taxonomicName id="A29F30C4A021FF9FFCC2FF0BE1D2FF62" box="[846,935,188,213]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFCC2FF0BE1D2FF62" box="[846,935,188,213]" italics="true" pageId="29" pageNumber="506">Berghia</emphasis>
</taxonomicName>
species have cylindrical cerata, with a round apex and uniform diameter throughout most of their length (
<bibRefCitation id="010E36B6A021FF9FFCB7FF57E640FF4F" author="Carmona" box="[827,1077,223,248]" pageId="29" pageNumber="506" refString="Carmona, L., Pola, M, Gosliner, T. M., Cervera, J. L. (2014 b) The Atlantic-Mediterranean genus Berghia (Mollusca, Nudibranchia, Aeolidiidae). Taxonomic review and phylogenetic analysis. Journal of Molluscan Studies. [in press]" type="book" year="2014" yearSuffix="b">
Carmona
<emphasis id="57EB9755A021FF9FFC27FF56E1A8FF4F" box="[939,989,223,248]" italics="true" pageId="29" pageNumber="506">et al</emphasis>
. 2014b
</bibRefCitation>
); the cerata of
<taxonomicName id="A29F30C4A021FF9FFB50FF68E710FF4F" box="[1244,1381,223,248]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFB50FF68E710FF4F" box="[1244,1381,223,248]" italics="true" pageId="29" pageNumber="506">Limenandra</emphasis>
</taxonomicName>
may have papillae over their surface (
<bibRefCitation id="010E36B6A021FF9FFD8BFEB3E08AFEAB" author="Carmona" box="[519,767,260,285]" pageId="29" pageNumber="506" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2014 c) The end of a long controversy: systematics of the genus Limenandra (Mollusca: Nudibranchia: Aeolidiidae). Helgoland Marine Research, 68, 37 - 48. http: // dx. doi. org / 10.1007 / s 10152 - 013 - 0367 - y" type="journal article" year="2014" yearSuffix="c">
Carmona
<emphasis id="57EB9755A021FF9FFDFAFEB2E0DAFEAA" box="[630,687,260,285]" italics="true" pageId="29" pageNumber="506">et al.</emphasis>
2014c
</bibRefCitation>
);
<taxonomicName id="A29F30C4A021FF9FFC9FFEB3E11BFEAA" box="[787,878,260,285]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFC9FFEB3E11BFEAA" box="[787,878,260,285]" italics="true" pageId="29" pageNumber="506">Spurilla</emphasis>
</taxonomicName>
has cerata that are recurved inwards (
<bibRefCitation id="010E36B6A021FF9FFA94FEB3E37DFEF7" author="Carmona" pageId="29" pageNumber="506" refString="Carmona, L., Lei, B. R., Pola, M., Gosliner, T. M., Valdes, A. &amp; Cervera, J. L. (2014 a) Untangling the Spurilla neapolitana (Delle Chiaje, 1841) species complex: a review of the genus Spurilla Bergh, 1864 (Mollusca: Nudibranchia: Aeolidiidae). Zoological Journal of the Linnean Society, 17, 132 - 154. http: // dx. doi. org / 10.1111 / zoj. 12098" type="journal article" year="2014" yearSuffix="a">
Carmona
<emphasis id="57EB9755A021FF9FFA04FEB2E2D8FEF7" italics="true" pageId="29" pageNumber="506">et al</emphasis>
. 2014a
</bibRefCitation>
), which are also present in some species of
<taxonomicName id="A29F30C4A021FF9FFC9AFE90E1F6FEF7" box="[790,899,295,320]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFC9AFE90E1F6FEF7" box="[790,899,295,320]" italics="true" pageId="29" pageNumber="506">Baeolidia</emphasis>
</taxonomicName>
. However, the latter usually have leaf-shaped cerata, being broad at their base and narrowing towards the apices.
</paragraph>
<paragraph id="65204B47A021FF9FFF4BFED8E0FBFE42" blockId="29.[151,1436,151,501]" pageId="29" pageNumber="506">
Regarding the radular morphology (
<figureCitation id="FDA457C2A021FF9FFDE2FED8E0B3FE3F" box="[622,710,367,392]" captionStart="FIGURE 16" captionStartId="28.[151,250,1201,1223]" captionTargetBox="[155,1432,193,1177]" captionTargetId="figure@28.[155,1432,193,1179]" captionTargetPageId="28" captionText="FIGURE 16. Comparison of the radular morphology among the genera Spurilla (A), Berghia (B), Limenandra (C) and Baeolidia (D)." httpUri="https://zenodo.org/record/249620/files/figure.png" pageId="29" pageNumber="506">Fig. 16</figureCitation>
), only
<taxonomicName id="A29F30C4A021FF9FFC92FED8E1D2FE3F" box="[798,935,367,392]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFC92FED8E1D2FE3F" box="[798,935,367,392]" italics="true" pageId="29" pageNumber="506">Limenandra</emphasis>
</taxonomicName>
is easily recognizable from the remaining genera (
<figureCitation id="FDA457C2A021FF9FFF7FFE23E332FE1A" box="[243,327,404,429]" captionStart="FIGURE 16" captionStartId="28.[151,250,1201,1223]" captionTargetBox="[155,1432,193,1177]" captionTargetId="figure@28.[155,1432,193,1179]" captionTargetPageId="28" captionText="FIGURE 16. Comparison of the radular morphology among the genera Spurilla (A), Berghia (B), Limenandra (C) and Baeolidia (D)." httpUri="https://zenodo.org/record/249620/files/figure.png" pageId="29" pageNumber="506">Fig. 16</figureCitation>
C). The radular teeth of
<taxonomicName id="A29F30C4A021FF9FFDD2FE23E0CDFE1A" box="[606,696,404,429]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFDD2FE23E0CDFE1A" box="[606,696,404,429]" italics="true" pageId="29" pageNumber="506">Spurilla</emphasis>
</taxonomicName>
,
<taxonomicName id="A29F30C4A021FF9FFD45FE23E157FE1A" box="[713,802,404,429]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFD45FE23E157FE1A" box="[713,802,404,429]" italics="true" pageId="29" pageNumber="506">Berghia</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A021FF9FFCD0FE23E1BCFE1A" box="[860,969,404,429]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFCD0FE23E1BCFE1A" box="[860,969,404,429]" italics="true" pageId="29" pageNumber="506">Baeolidia</emphasis>
</taxonomicName>
are very similar, corresponding with the radular teeth represented in the
<figureCitation id="FDA457C2A021FF9FFE7BFE00E013FE67" box="[503,614,439,464]" captionStart="FIGURE 17" captionStartId="29.[151,250,1180,1202]" captionText="FIGURE 17. Most common radular teeth morphology in Aeolidiidae." pageId="29" pageNumber="506">Figure 17</figureCitation>
. Although there is quite a lot of intrageneric variability,
<figureCitation id="FDA457C2A021FF9FFB56FE00E733FE67" box="[1242,1350,439,464]" captionStart="FIGURE 16" captionStartId="28.[151,250,1201,1223]" captionTargetBox="[155,1432,193,1177]" captionTargetId="figure@28.[155,1432,193,1179]" captionTargetPageId="28" captionText="FIGURE 16. Comparison of the radular morphology among the genera Spurilla (A), Berghia (B), Limenandra (C) and Baeolidia (D)." httpUri="https://zenodo.org/record/249620/files/figure.png" pageId="29" pageNumber="506">Figure 16</figureCitation>
depicts the archetypical radular teeth for each genus.
</paragraph>
<caption id="31E01BCFA021FF9FFF1BFB2BE12BFB05" box="[151,862,1180,1202]" pageId="29" pageNumber="506">
<paragraph id="65204B47A021FF9FFF1BFB2BE12BFB05" blockId="29.[151,862,1180,1202]" box="[151,862,1180,1202]" pageId="29" pageNumber="506">
<emphasis id="57EB9755A021FF9FFF1BFB2BE36AFB06" bold="true" box="[151,287,1180,1202]" pageId="29" pageNumber="506">FIGURE 17.</emphasis>
Most common radular teeth morphology in
<taxonomicName id="A29F30C4A021FF9FFD6CFB2BE12FFB05" box="[736,858,1180,1202]" class="Gastropoda" family="Aeolidiidae" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="family">Aeolidiidae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="65204B47A021FF9FFF4BFAB4E103FAD4" blockId="29.[151,1436,1282,1451]" pageId="29" pageNumber="506">
In terms of biogeography, so far
<taxonomicName id="A29F30C4A021FF9FFDE7FAB5E0B1FAAC" box="[619,708,1282,1307]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFDE7FAB5E0B1FAAC" box="[619,708,1282,1307]" italics="true" pageId="29" pageNumber="506">Berghia</emphasis>
</taxonomicName>
is the only genus restricted to one ocean basin (Atlantic- Mediterranean) (
<figureCitation id="FDA457C2A021FF9FFEDFFA91E3D7FA88" box="[339,418,1318,1343]" captionStart="FIGURE 18" captionStartId="30.[151,250,1519,1541]" captionTargetBox="[151,1427,195,1497]" captionTargetId="figure@30.[151,1436,195,1498]" captionTargetPageId="30" captionText="FIGURE 18. Map of the range of the different species of Spurilla, Berghia, Limenandra, and Baeolidia. A. Berghia and Limenandra. B. Spurilla and Baeolidia. Abbreviation: Isolated records of Baeolidia cryporos (star) and Spurilla braziliana (square)." httpUri="https://zenodo.org/record/249621/files/figure.png" pageId="29" pageNumber="506">Fig. 18</figureCitation>
A) although the occurrence of otherwise Indo-Pacific
<taxonomicName id="A29F30C4A021FF9FFB8CFA91E618FA88" box="[1024,1133,1318,1343]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFB8CFA91E618FA88" box="[1024,1133,1318,1343]" italics="true" pageId="29" pageNumber="506">Baeolidia</emphasis>
</taxonomicName>
in the Atlantic Ocean with
<taxonomicName id="A29F30C4A021FF9FFF1BFAFCE352FAD5" box="[151,295,1355,1378]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="species" species="cryoporos">
<emphasis id="57EB9755A021FF9FFF1BFAFCE352FAD5" box="[151,295,1355,1378]" italics="true" pageId="29" pageNumber="506">B. cryoporos</emphasis>
</taxonomicName>
would need further comparatives studies (
<figureCitation id="FDA457C2A021FF9FFC8FFAFDE120FAD4" box="[771,853,1354,1379]" captionStart="FIGURE 18" captionStartId="30.[151,250,1519,1541]" captionTargetBox="[151,1427,195,1497]" captionTargetId="figure@30.[151,1436,195,1498]" captionTargetPageId="30" captionText="FIGURE 18. Map of the range of the different species of Spurilla, Berghia, Limenandra, and Baeolidia. A. Berghia and Limenandra. B. Spurilla and Baeolidia. Abbreviation: Isolated records of Baeolidia cryporos (star) and Spurilla braziliana (square)." httpUri="https://zenodo.org/record/249621/files/figure.png" pageId="29" pageNumber="506">Fig. 18</figureCitation>
B).
</paragraph>
<paragraph id="65204B47A021FF9FFF4BFAD8E3BDFA1C" blockId="29.[151,1436,1282,1451]" pageId="29" pageNumber="506">In order to facilitate the identification of these four genera, here we present a small key based on external characters and distribution.</paragraph>
<paragraph id="65204B47A021FF9FFF1BFA6EE06DF8A3" blockId="29.[151,1436,1497,1812]" pageId="29" pageNumber="506">
1 Rhinophores perfoliate...........................................................................
<taxonomicName id="A29F30C4A021FF9FFAC2FA6EE7EEFA58" box="[1358,1435,1497,1519]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFAC2FA6EE7EEFA58" box="[1358,1435,1497,1519]" italics="true" pageId="29" pageNumber="506">Spurilla</emphasis>
</taxonomicName>
- Rhinophores ribbed.....................................................................
<taxonomicName id="A29F30C4A021FF9FFB6EFA41E7E9F9BB" box="[1250,1436,1526,1548]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="species" species="macleayi">
<emphasis id="57EB9755A021FF9FFB6EFA41E7E9F9BB" box="[1250,1436,1526,1548]" italics="true" pageId="29" pageNumber="506">Baeolidia macleayi</emphasis>
</taxonomicName>
- Rhinophores smooth.....................................................................
<taxonomicName id="A29F30C4A021FF9FFB63F9A3E7EEF99D" box="[1263,1435,1556,1578]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="species" species="ransoni">
<emphasis id="57EB9755A021FF9FFB63F9A3E7EEF99D" box="[1263,1435,1556,1578]" italics="true" pageId="29" pageNumber="506">Baeolidia ransoni</emphasis>
</taxonomicName>
- Rhinophores papillate................................................................................. 2 2 Notum with colouration pattern of concentric circles.................................................
<taxonomicName id="A29F30C4A021FF9FFAAAF9F9E7EEF9D3" box="[1318,1435,1614,1636]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFAAAF9F9E7EEF9D3" box="[1318,1435,1614,1636]" italics="true" pageId="29" pageNumber="506">Limenandra</emphasis>
</taxonomicName>
- Notum without colouration pattern of concentric circles...................................................... 3 3 Cerata cylindrical.................................................................................... 4 - Cerata non-cylindrical................................................................................. 5 4 Indo-Pacific distribution.....................................................................
<taxonomicName id="A29F30C4A021FF9FFA83F973E7E9F96D" box="[1295,1436,1732,1754]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="species" species="dela">
<emphasis id="57EB9755A021FF9FFA83F973E7E9F96D" box="[1295,1436,1732,1754]" italics="true" pageId="29" pageNumber="506">Baeolidia dela</emphasis>
</taxonomicName>
- Atlantic-Mediterranean distribution.................................................................
<taxonomicName id="A29F30C4A021FF9FFAC3F956E7EEF940" box="[1359,1435,1761,1783]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFAC3F956E7EEF940" box="[1359,1435,1761,1783]" italics="true" pageId="29" pageNumber="506">Berghia</emphasis>
</taxonomicName>
5 Remaining species of
<taxonomicName id="A29F30C4A021FF9FFE39F949E067F8A3" box="[437,530,1790,1812]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="29" pageNumber="506" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A021FF9FFE39F949E067F8A3" box="[437,530,1790,1812]" italics="true" pageId="29" pageNumber="506">Baeolidia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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