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<mods:title id="71604F5890BF5D367C067B71CC71557B">Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar</mods:title>
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<mods:affiliation id="C8C1E986D1DEE7C5BF5BF35AFB12CB33">Mécanismes adaptatifs et évolution (MECADEV), Muséum national dHistoire naturelle, CNRS, 57 rue Cuvier, CP 50, 75005 Paris, France.</mods:affiliation>
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<mods:namePart id="7905B3B6C1DB265A18A7A53289F0881B">Cesne, Maxime Le</mods:namePart>
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<paragraph id="60678614D107FFB59252B30AFBD5F992" blockId="23.[524,1064,1632,1693]" box="[524,1064,1632,1659]" pageId="23" pageNumber="189">
<heading id="3B2F3178D107FFB59252B30AFBD5F992" box="[524,1064,1632,1659]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="189" reason="2">
<taxonomicName id="A7D8FD97D107FFB59252B30AFC63F993" authority="(Fallou, 1890)" authorityName="Bucher &amp; Bouteille &amp; Cesne" authorityYear="2024" baseAuthorityName="Fallou" baseAuthorityYear="1890" box="[524,926,1632,1659]" class="Insecta" family="Cercopidae" genus="Soulierana" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="189" phylum="Arthropoda" rank="species" species="rugosa" status="comb. nov.">
<emphasis id="52AC5A06D107FFB59252B30AFD12F993" bold="true" box="[524,751,1632,1659]" italics="true" pageId="23" pageNumber="189">Soulierana rugosa</emphasis>
(
<bibRefCitation id="0449FBE5D107FFB592A1B30AFC6BF993" author="Fallou G." box="[767,918,1632,1658]" firstAuthor="Fallou" journalOrPublisher="Revue d'Entomologie. Societe Francaise d'Entomologie" pageId="23" pageNumber="189" pagination="351 - 354" part="9" refId="ref15133" refString="Fallou G. 1890. Diagnoses d'Homopteres nouveaux. Revue d'Entomologie. Societe Francaise d'Entomologie 9: 351 - 354." title="Diagnoses d'Homopteres nouveaux" type="journal article" year="1890">Fallou, 1890</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="499FE77DD107FFB593F8B30AFBD5F992" box="[934,1064,1632,1659]" pageId="23" pageNumber="189" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="28C2D59FD107FFB592AEB3E9FCBEF974" box="[752,835,1667,1693]" pageId="23" pageNumber="189" type="description">
<paragraph id="60678614D107FFB592AEB3E9FCBEF974" blockId="23.[524,1064,1632,1693]" box="[752,835,1667,1693]" pageId="23" pageNumber="189">
<figureCitation id="F8E39A91D107FFB592AEB3E9FCBEF974" box="[752,835,1667,1693]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[243,1327,272,1836]" captionTargetId="figure-81@24.[230,1358,262,1857]" captionTargetPageId="24" captionText="Fig. 10. Soulierana rugosa (Fallou, 1890), ♂ (MNHN(EH) 24695). A. Dorsal view. B. Lateral view. C. Frontal view.D. Labels.E. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate. F. Aedeagus. G. Left paramere, outside view on the left and top view on the rigth. H. Distribution map. Scale bars: AC = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11400848" httpUri="https://zenodo.org/record/11400848/files/figure.png" pageId="23" pageNumber="189">Fig. 10</figureCitation>
</paragraph>
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<paragraph id="60678614D107FFB590E3B3AFFD70F936" blockId="23.[189,1082,1732,1795]" box="[189,653,1732,1759]" pageId="23" pageNumber="189">
<treatmentCitationGroup id="40C8A13AD107FFB590E3B3AFFD70F936" box="[189,653,1732,1759]" pageId="23" pageNumber="189">
<treatmentCitation id="E179A005D107FFB590E3B3AFFD74F936" author="Fallou G." box="[189,649,1732,1759]" page="352" pageId="23" pageNumber="189" year="1890">
<taxonomicName id="A7D8FD97D107FFB590E3B3AFFD74F936" ID-CoL="6RNFL" authority="Fallou, 1890: 352" authorityName="Fallou" authorityPageNumber="352" authorityYear="1890" box="[189,649,1732,1759]" class="Insecta" family="Cercopidae" genus="Monecphora" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="189" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="52AC5A06D107FFB590E3B3AFFE53F937" box="[189,430,1733,1759]" italics="true" pageId="23" pageNumber="189">Monecphora rugosa</emphasis>
<bibRefCitation id="0449FBE5D107FFB591E8B3AFFD74F936" author="Fallou G." box="[438,649,1732,1759]" firstAuthor="Fallou" journalOrPublisher="Revue d'Entomologie. Societe Francaise d'Entomologie" page="352" pageId="23" pageNumber="189" pagination="351 - 354" part="9" refId="ref15133" refString="Fallou G. 1890. Diagnoses d'Homopteres nouveaux. Revue d'Entomologie. Societe Francaise d'Entomologie 9: 351 - 354." title="Diagnoses d'Homopteres nouveaux" type="journal article" year="1890">Fallou, 1890: 352</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="60678614D107FFB590E3B383FBC7F8EB" blockId="23.[189,1082,1732,1795]" box="[189,1082,1768,1795]" pageId="23" pageNumber="189">
<treatmentCitationGroup id="40C8A13AD107FFB590E3B383FBC7F8EB" box="[189,1082,1768,1795]" pageId="23" pageNumber="189">
<treatmentCitation id="E179A005D107FFB590E3B383FD78F8EA" author="Lallemand V." box="[189,645,1768,1795]" page="285" pageId="23" pageNumber="189" year="1920">
<taxonomicName id="A7D8FD97D107FFB590E3B383FD78F8EA" ID-CoL="3VHDD" authority="Lallemand, 1920: 285" authorityName="Lallemand" authorityPageNumber="285" authorityYear="1920" box="[189,645,1768,1795]" class="Insecta" family="Cercopidae" genus="Literna" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="189" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="52AC5A06D107FFB590E3B383FE81F8EB" box="[189,380,1768,1794]" italics="true" pageId="23" pageNumber="189">Literna jeanneli</emphasis>
<bibRefCitation id="0449FBE5D107FFB591DDB382FD78F8EA" author="Lallemand V." box="[387,645,1768,1795]" firstAuthor="Lallemand" journalOrPublisher="Annales de la Societe entomologique de France" page="285" pageId="23" pageNumber="189" pagination="280 - 288" part="88" refId="ref15265" refString="Lallemand V. 1920. Cercopides nouveaux de Madagascar. Annales de la Societe entomologique de France 88: 280 - 288." title="Cercopides nouveaux de Madagascar" type="journal article" year="1920">Lallemand, 1920: 285</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(synonymy by
<treatmentCitation id="E179A005D107FFB59363B382FBD0F8EB" author="Lallemand V." box="[829,1069,1768,1794]" page="67" pageId="23" pageNumber="189" year="1949">
<bibRefCitation id="0449FBE5D107FFB59363B382FBD0F8EB" author="Lallemand V." box="[829,1069,1768,1794]" firstAuthor="Lallemand" journalOrPublisher="Memoires de l'Institut royal des Sciences naturelles de Belgique" page="67" pageId="23" pageNumber="189" pagination="1 - 193" part="2" refId="ref15288" refString="Lallemand V. 1949. Revision des Cercopinae (Hemiptera Homoptera) Premiere partie. Memoires de l'Institut royal des Sciences naturelles de Belgique 2 (32): 1 - 193." title="Revision des Cercopinae (Hemiptera Homoptera) Premiere partie" type="journal article" year="1949">Lallemand 1949: 67</bibRefCitation>
</treatmentCitation>
).
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</paragraph>
<paragraph id="60678614D107FFB590E3B240FDA1F8AD" blockId="23.[189,604,1833,1860]" box="[189,604,1833,1860]" pageId="23" pageNumber="189">
<treatmentCitationGroup id="40C8A13AD107FFB590E3B240FDA1F8AD" box="[189,604,1833,1860]" pageId="23" pageNumber="189">
<emphasis id="52AC5A06D107FFB590E3B240FE78F8AA" box="[189,389,1834,1859]" italics="true" pageId="23" pageNumber="189">
<taxonomicName id="A7D8FD97D107FFB590E3B240FE93F8AA" ID-CoL="3VHDV" baseAuthorityName="Fallou" baseAuthorityYear="1890" box="[189,366,1834,1859]" class="Insecta" family="Cercopidae" genus="Literna" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="189" phylum="Arthropoda" rank="species" species="rugosa">Literna rugosa</taxonomicName>
</emphasis>
<treatmentCitation id="E179A005D107FFB591D2B243FDA5F8AD" author="Jacobi A." box="[396,600,1833,1860]" page="541" pageId="23" pageNumber="189" year="1917">
<bibRefCitation id="0449FBE5D107FFB591D2B243FDA5F8AD" author="Jacobi A." box="[396,600,1833,1860]" firstAuthor="Jacobi" journalOrPublisher="Voeltzkow's Reise in Ostafrika" page="541" pageId="23" pageNumber="189" pagination="519 - 552" part="3" refId="ref15154" refString="Jacobi A. 1917. Die Zikadenfauna Madagascars und der Comoren. Voeltzkow's Reise in Ostafrika 3: 519 - 552." title="Die Zikadenfauna Madagascars und der Comoren" type="journal article" year="1917">Jacobi 1917: 541</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="28C2D59FD107FFBB90E3B200FDC1FD2D" lastPageId="25" lastPageNumber="191" pageId="23" pageNumber="189" type="materials_examined">
<paragraph id="60678614D107FFB590E3B200FE57F86D" blockId="23.[189,426,1898,1924]" box="[189,426,1898,1924]" pageId="23" pageNumber="189">
<heading id="3B2F3178D107FFB590E3B200FE57F86D" bold="true" box="[189,426,1898,1924]" fontSize="11" level="3" pageId="23" pageNumber="189" reason="3">
<emphasis id="52AC5A06D107FFB590E3B200FE57F86D" bold="true" box="[189,426,1898,1924]" pageId="23" pageNumber="189">Material examined</emphasis>
</heading>
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<paragraph id="60678614D107FFB590B4B2F3FC73F83E" blockId="23.[189,910,1945,2007]" pageId="23" pageNumber="189">
<materialsCitation id="D0B08C49D107FFB590B4B2F3FC73F83E" ID-GBIF-Occurrence="4873642345" collectionCode="MNHN" country="Madagascar" location="MADAGASCAR" pageId="23" pageNumber="189" specimenCode="MNHN EH 24695" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<emphasis id="52AC5A06D107FFB590B4B2F3FE9BF85A" bold="true" box="[234,358,1945,1971]" pageId="23" pageNumber="189">
<typeStatus id="BF6338B6D107FFB590B4B2F3FE9BF85A" box="[234,358,1945,1971]" pageId="23" pageNumber="189" type="lectotype">Lectotype</typeStatus>
</emphasis>
<collectingCountry id="18CFC684D107FFB590E3B2D7FE73F83E" box="[189,398,1981,2007]" name="Madagascar" pageId="23" pageNumber="189">MADAGASCAR</collectingCountry>
<specimenCount id="76DE4D9DD107FFB591F8B2D7FE40F83E" box="[422,445,1981,2007]" count="1" pageId="23" pageNumber="189" type="female"></specimenCount>
; collection Fallou;
<collectionCode id="06C91ED1D107FFB592FFB2D7FD03F83E" box="[673,766,1981,2007]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="189">MNHN</collectionCode>
<specimenCode id="307E2E6FD107FFB592A0B2D7FC77F83E" box="[766,906,1980,2007]" pageId="23" pageNumber="189">(EH) 24695</specimenCode>
.
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</paragraph>
<caption id="34A7D69CD108FFBA90E3B20EFCDBF800" ID-DOI="http://doi.org/10.5281/zenodo.11400848" ID-Zenodo-Dep="11400848" httpUri="https://zenodo.org/record/11400848/files/figure.png" pageId="24" pageNumber="190" startId="24.[189,232,1892,1918]" targetBox="[243,1327,272,1836]" targetPageId="24">
<paragraph id="60678614D108FFBA90E3B20EFCDBF800" blockId="24.[187,1399,1892,2025]" pageId="24" pageNumber="190">
<emphasis id="52AC5A06D108FFBA90E3B20EFEE1F896" bold="true" box="[189,284,1892,1919]" pageId="24" pageNumber="190">Fig. 10.</emphasis>
<taxonomicName id="A7D8FD97D108FFBA9178B20FFD41F896" authority="(Fallou, 1890)" authorityName="Bucher &amp; Bouteille &amp; Cesne" authorityYear="2024" baseAuthorityName="Fallou" baseAuthorityYear="1890" box="[294,700,1893,1919]" class="Insecta" family="Cercopidae" genus="Soulierana" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="190" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="52AC5A06D108FFBA9178B20FFDFEF896" box="[294,515,1893,1919]" italics="true" pageId="24" pageNumber="190">Soulierana rugosa</emphasis>
(
<bibRefCitation id="0449FBE5D108FFBA9246B20FFD49F896" author="Fallou G." box="[536,692,1893,1919]" firstAuthor="Fallou" journalOrPublisher="Revue d'Entomologie. Societe Francaise d'Entomologie" pageId="24" pageNumber="190" pagination="351 - 354" part="9" refId="ref15133" refString="Fallou G. 1890. Diagnoses d'Homopteres nouveaux. Revue d'Entomologie. Societe Francaise d'Entomologie 9: 351 - 354." title="Diagnoses d'Homopteres nouveaux" type="journal article" year="1890">Fallou, 1890</bibRefCitation>
)
</taxonomicName>
, ♂ (MNHN(EH) 24695).
<emphasis id="52AC5A06D108FFBA93ACB20EFBFAF897" bold="true" box="[1010,1031,1892,1918]" pageId="24" pageNumber="190">A</emphasis>
. Dorsal view.
<emphasis id="52AC5A06D108FFBA94E7B20FFB36F896" bold="true" box="[1209,1227,1893,1919]" pageId="24" pageNumber="190">B</emphasis>
. Lateral view.
<emphasis id="52AC5A06D108FFBA90E1B2E2FF2FF84B" bold="true" box="[191,210,1928,1954]" pageId="24" pageNumber="190">C</emphasis>
. Frontal view.
<emphasis id="52AC5A06D108FFBA9125B2E2FE6DF84B" bold="true" box="[379,400,1928,1954]" pageId="24" pageNumber="190">D</emphasis>
. Labels.
<emphasis id="52AC5A06D108FFBA91A6B2E2FDF1F84B" bold="true" box="[504,524,1928,1954]" pageId="24" pageNumber="190">E</emphasis>
. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate.
<emphasis id="52AC5A06D108FFBA91B4B2C1FE06F82C" bold="true" box="[490,507,1963,1989]" pageId="24" pageNumber="190">F</emphasis>
. Aedeagus.
<emphasis id="52AC5A06D108FFBA92D3B2C1FD59F82C" bold="true" box="[653,676,1963,1989]" pageId="24" pageNumber="190">G</emphasis>
. Left paramere, outside view on the left and top view on the rigth.
<emphasis id="52AC5A06D108FFBA915DB2A4FEE7F801" bold="true" box="[259,282,1998,2024]" pageId="24" pageNumber="190">H</emphasis>
. Distribution map. Scale bars: AC = 1 mm.
</paragraph>
</caption>
<paragraph id="60678614D109FFBB90B4B463FC59FEAF" blockId="25.[189,932,265,326]" pageId="25" pageNumber="191">
<materialsCitation id="D0B08C49D109FFBB90B4B463FC5DFEAF" ID-GBIF-Occurrence="4873642361" collectionCode="MNHN" country="Madagascar" location="MADAGASCAR" pageId="25" pageNumber="191" specimenCode="MNHN EH 24694" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<emphasis id="52AC5A06D109FFBB90B4B463FE6AFECA" bold="true" box="[234,407,265,291]" pageId="25" pageNumber="191">
<typeStatus id="BF6338B6D109FFBB90B4B463FE6AFECA" box="[234,407,265,291]" pageId="25" pageNumber="191" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
<collectingCountry id="18CFC684D109FFBB90E3B446FE73FEAF" box="[189,398,300,326]" name="Madagascar" pageId="25" pageNumber="191">MADAGASCAR</collectingCountry>
<specimenCount id="76DE4D9DD109FFBB91F8B446FE2EFEAF" box="[422,467,300,326]" count="1" pageId="25" pageNumber="191" type="female">1 ♀</specimenCount>
; collection Fallou;
<collectionCode id="06C91ED1D109FFBB92E9B446FCE9FEAF" box="[695,788,300,326]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="191">MNHN</collectionCode>
<specimenCode id="307E2E6FD109FFBB934AB446FC5DFEAF" box="[788,928,300,326]" pageId="25" pageNumber="191">(EH) 24694</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="60678614D109FFBB90B4B407FE5BFE6E" blockId="25.[189,1399,365,708]" box="[234,422,365,391]" pageId="25" pageNumber="191">
<emphasis id="52AC5A06D109FFBB90B4B407FE5BFE6E" bold="true" box="[234,422,365,391]" pageId="25" pageNumber="191">Other material</emphasis>
</paragraph>
<paragraph id="60678614D109FFBB90E3B4FAFDC1FD2D" blockId="25.[189,1399,365,708]" pageId="25" pageNumber="191">
<materialsCitation id="D0B08C49D109FFBB90E3B4FAFB68FE27" ID-GBIF-Occurrence="4873642320" collectedFrom="foret humide" collectingDate="2011-11-09" collectionCode="MNHN" collectorName="A. Soulier-Perkins" country="Madagascar" county="Province" elevation="944" latitude="-18.981184" location="Toamasina Andasibe" longLatPrecision="1" longitude="48.420033" pageId="25" pageNumber="191" specimenCode="MNHN EH 24118" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina Andasibe">
<collectingCountry id="18CFC684D109FFBB90E3B4FAFE76FE43" box="[189,395,400,426]" name="Madagascar" pageId="25" pageNumber="191">MADAGASCAR</collectingCountry>
<specimenCount id="76DE4D9DD109FFBB91F3B4FAFE22FE42" box="[429,479,400,427]" count="1" pageId="25" pageNumber="191" type="male">1 ♂</specimenCount>
; Province de
<collectingRegion id="A21C48F6D109FFBB92D4B4FAFC7DFE43" box="[650,896,400,426]" pageId="25" pageNumber="191">Toamasina Andasibe</collectingRegion>
,
<collectedFrom id="D3C4C5E9D109FFBB93CEB4FAFBD6FE43" box="[912,1067,400,426]" pageId="25" pageNumber="191">forêt humide</collectedFrom>
, s/
<emphasis id="52AC5A06D109FFBB9410B4FAFB3CFE43" box="[1102,1217,400,426]" italics="true" pageId="25" pageNumber="191">Hisbiscus</emphasis>
;
<geoCoordinate id="05ECE0D3D109FFBB9488B4E5FA8CFE43" box="[1238,1393,399,427]" degrees="18" direction="south" minutes="58.871" orientation="latitude" pageId="25" pageNumber="191" precision="1" value="-18.981184">18°58.871 S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D109FFBB90E3B4D8FEA5FE27" box="[189,344,434,462]" degrees="48" direction="east" minutes="25.202" orientation="longitude" pageId="25" pageNumber="191" precision="1" value="48.420033">48°25.202 E</geoCoordinate>
;
<elevation id="EBF56127D109FFBB913BB4DEFE12FE24" box="[357,495,435,462]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">
<quantity id="A7202BF1D109FFBB913BB4DEFE52FE27" box="[357,431,436,462]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">944 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D109FFBB91A3B4DEFD72FE27" box="[509,655,436,462]" pageId="25" pageNumber="191" value="2011-11-09">
<collectingDate id="0422593CD109FFBB91A3B4DEFD72FE27" box="[509,655,436,462]" pageId="25" pageNumber="191" value="2011-11-09">9 Nov. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D109FFBB92C4B4DEFC8CFE24" box="[666,881,435,462]" pageId="25" pageNumber="191">A. Soulier-Perkins</collectorName>
leg.;
<collectionCode id="06C91ED1D109FFBB93EDB4DEFBF3FE27" box="[947,1038,436,462]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="191">MNHN</collectionCode>
(
<specimenCode id="307E2E6FD109FFBB9448B4DEFB68FE27" box="[1046,1173,436,462]" pageId="25" pageNumber="191">EH) 24118</specimenCode>
</materialsCitation>
<materialsCitation id="D0B08C49D109FFBB94EEB4DEFD3CFDFD" ID-GBIF-Occurrence="4873642378" collectedFrom="foret humide" collectingDate="2011-11-09" collectionCode="MNHN" collectorName="A. Soulier-Perkins" country="Madagascar" county="Province" elevation="944" latitude="-18.981184" location="Toamasina Andasibe" longLatPrecision="1" longitude="48.420033" pageId="25" pageNumber="191" specimenCode="MNHN EH 24634" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina Andasibe">
<specimenCount id="76DE4D9DD109FFBB94EEB4DEFB23FE27" box="[1200,1246,436,462]" count="1" pageId="25" pageNumber="191" type="male">1 ♂</specimenCount>
; Province de
<collectingRegion id="A21C48F6D109FFBB90E3B4BDFE4DFE18" box="[189,432,471,497]" pageId="25" pageNumber="191">Toamasina Andasibe</collectingRegion>
,
<collectedFrom id="D3C4C5E9D109FFBB91E3B4BDFDA8FE18" box="[445,597,471,497]" pageId="25" pageNumber="191">forêt humide</collectedFrom>
, s/Hisbiscus;
<geoCoordinate id="05ECE0D3D109FFBB92ABB4BFFC73FE18" box="[757,910,469,497]" degrees="18" direction="south" minutes="58.871" orientation="latitude" pageId="25" pageNumber="191" precision="1" value="-18.981184">18°58.871 S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D109FFBB93C2B4BFFBCAFE18" box="[924,1079,469,497]" degrees="48" direction="east" minutes="25.202" orientation="longitude" pageId="25" pageNumber="191" precision="1" value="48.420033">48°25.202 E</geoCoordinate>
;
<elevation id="EBF56127D109FFBB941BB4BDFB2DFE18" box="[1093,1232,471,497]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">
<quantity id="A7202BF1D109FFBB941BB4BDFB6DFE18" box="[1093,1168,471,497]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">944 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D109FFBB9481B4BDFA8FFE18" box="[1247,1394,471,497]" pageId="25" pageNumber="191" value="2011-11-09">
<collectingDate id="0422593CD109FFBB9481B4BDFA8FFE18" box="[1247,1394,471,497]" pageId="25" pageNumber="191" value="2011-11-09">9 Nov. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D109FFBB90E3B490FE68FDFD" box="[189,405,506,532]" pageId="25" pageNumber="191">A. Soulier-Perkins</collectorName>
leg.;
<collectionCode id="06C91ED1D109FFBB9182B490FDCAFDFD" box="[476,567,506,532]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="191">MNHN</collectionCode>
<specimenCode id="307E2E6FD109FFBB9266B490FD3CFDFD" box="[568,705,506,532]" pageId="25" pageNumber="191">(EH) 24634</specimenCode>
</materialsCitation>
<materialsCitation id="D0B08C49D109FFBB92BEB490FEF8FDB3" ID-GBIF-Occurrence="4873642339" collectedFrom="foret humide" collectingDate="2011-11-09" collectionCode="MNHN" collectorName="A. Soulier-Perkins" country="Madagascar" county="Province" elevation="944" latitude="-18.981184" location="Toamasina Andasibe" longLatPrecision="1" longitude="48.420033" pageId="25" pageNumber="191" specimenCode="MNHN EH 24698" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina Andasibe">
<specimenCount id="76DE4D9DD109FFBB92BEB490FCECFDFD" box="[736,785,506,532]" count="1" pageId="25" pageNumber="191" type="male">1 ♂</specimenCount>
; Province de
<collectingRegion id="A21C48F6D109FFBB93E9B490FB51FDFD" box="[951,1196,506,532]" pageId="25" pageNumber="191">Toamasina Andasibe</collectingRegion>
,
<collectedFrom id="D3C4C5E9D109FFBB94E5B490FAA9FDFD" box="[1211,1364,506,532]" pageId="25" pageNumber="191">forêt humide</collectedFrom>
, s/ Hisbiscus;
<geoCoordinate id="05ECE0D3D109FFBB9162B776FE29FDDE" box="[316,468,540,568]" degrees="18" direction="south" minutes="58.871" orientation="latitude" pageId="25" pageNumber="191" precision="1" value="-18.981184">18°58.871 S</geoCoordinate>
, E
<geoCoordinate id="05ECE0D3D109FFBB91ADB776FD70FDD1" box="[499,653,540,568]" degrees="48" direction="east" minutes="25.202" orientation="longitude" pageId="25" pageNumber="191" precision="1" value="48.420033">48°25.202 E</geoCoordinate>
;
<elevation id="EBF56127D109FFBB92C4B777FCDEFDDE" box="[666,803,541,568]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">
<quantity id="A7202BF1D109FFBB92C4B777FD19FDD1" box="[666,740,541,568]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">944 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D109FFBB936EB777FC3FFDDE" box="[816,962,541,568]" pageId="25" pageNumber="191" value="2011-11-09">
<collectingDate id="0422593CD109FFBB936EB777FC3FFDDE" box="[816,962,541,568]" pageId="25" pageNumber="191" value="2011-11-09">9 Nov. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D109FFBB9393B777FB5EFDDE" box="[973,1187,541,567]" pageId="25" pageNumber="191">A. Soulier-Perkins</collectorName>
leg.;
<collectionCode id="06C91ED1D109FFBB94BAB777FABDFDDE" box="[1252,1344,541,567]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="191">MNHN</collectionCode>
<specimenCode id="307E2E6FD109FFBB951EB777FEF8FDB3" pageId="25" pageNumber="191">(EH) 24698</specimenCode>
</materialsCitation>
<materialsCitation id="D0B08C49D109FFBB917FB72AFBF9FD94" ID-GBIF-Occurrence="4873642370" collectedFrom="foret humide" collectingDate="2011-11-09" collectionCode="MNHN" collectorName="A. Soulier-Perkins" country="Madagascar" county="Province" elevation="944" latitude="-18.981184" location="Toamasina Andasibe" longLatPrecision="1" longitude="48.420033" pageId="25" pageNumber="191" specimenCode="MNHN EH 24699" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina Andasibe">
<specimenCount id="76DE4D9DD109FFBB917FB72AFEAFFDB2" box="[289,338,576,603]" count="1" pageId="25" pageNumber="191" type="male">1 ♂</specimenCount>
; Province de
<collectingRegion id="A21C48F6D109FFBB91AAB72AFD15FDB3" box="[500,744,576,602]" pageId="25" pageNumber="191">Toamasina Andasibe</collectingRegion>
,
<collectedFrom id="D3C4C5E9D109FFBB92ABB72AFC70FDB3" box="[757,909,576,602]" pageId="25" pageNumber="191">forêt humide</collectedFrom>
, s/Hisbiscus;
<geoCoordinate id="05ECE0D3D109FFBB9471B755FB35FDB3" box="[1071,1224,575,603]" degrees="18" direction="south" minutes="58.871" orientation="latitude" pageId="25" pageNumber="191" precision="1" value="-18.981184">18°58.871 S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D109FFBB9488B755FA8FFDB2" box="[1238,1394,575,603]" degrees="48" direction="east" minutes="25.202" orientation="longitude" pageId="25" pageNumber="191" precision="1" value="48.420033">48°25.202 E</geoCoordinate>
;
<elevation id="EBF56127D109FFBB90E3B70EFEB6FD94" box="[189,331,611,638]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">
<quantity id="A7202BF1D109FFBB90E3B70EFEF7FD97" box="[189,266,612,638]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">944 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D109FFBB9105B70EFE0FFD97" box="[347,498,612,638]" pageId="25" pageNumber="191" value="2011-11-09">
<collectingDate id="0422593CD109FFBB9105B70EFE0FFD97" box="[347,498,612,638]" pageId="25" pageNumber="191" value="2011-11-09">9 Nov. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D109FFBB91A1B70EFD25FD94" box="[511,728,611,638]" pageId="25" pageNumber="191">A. Soulier-Perkins</collectorName>
leg.;
<collectionCode id="06C91ED1D109FFBB9341B70EFC87FD97" box="[799,890,612,638]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="191">MNHN</collectionCode>
<specimenCode id="307E2E6FD109FFBB9324B70EFBF9FD94" box="[890,1028,611,638]" pageId="25" pageNumber="191">(EH) 24699</specimenCode>
</materialsCitation>
<materialsCitation id="D0B08C49D109FFBB947DB70EFDC5FD2D" ID-GBIF-Occurrence="4873642306" collectedFrom="foret humide" collectingDate="2011-11-09" collectionCode="MNHN" collectorName="A. Soulier-Perkins" country="Madagascar" county="Province" elevation="944" latitude="-18.981184" location="Toamasina Andasibe" longLatPrecision="1" longitude="48.420033" pageId="25" pageNumber="191" specimenCode="MNHN EH 24703" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina Andasibe">
<specimenCount id="76DE4D9DD109FFBB947DB70EFBA9FD97" box="[1059,1108,612,638]" count="1" pageId="25" pageNumber="191" type="male">1 ♂</specimenCount>
; Province de
<collectingRegion id="A21C48F6D109FFBB94A4B709FED0FD48" pageId="25" pageNumber="191">Toamasina Andasibe</collectingRegion>
,
<collectedFrom id="D3C4C5E9D109FFBB9167B7EDFE2CFD48" box="[313,465,647,673]" pageId="25" pageNumber="191">forêt humide</collectedFrom>
, s/Hisbiscus;
<geoCoordinate id="05ECE0D3D109FFBB922FB7EFFCF4FD48" box="[625,777,645,673]" degrees="18" direction="south" minutes="58.871" orientation="latitude" pageId="25" pageNumber="191" precision="1" value="-18.981184">18°58.871 S</geoCoordinate>
,
<geoCoordinate id="05ECE0D3D109FFBB9349B7EFFC4FFD48" box="[791,946,645,673]" degrees="48" direction="east" minutes="25.202" orientation="longitude" pageId="25" pageNumber="191" precision="1" value="48.420033">48°25.202 E</geoCoordinate>
;
<elevation id="EBF56127D109FFBB93E1B7EDFBB4FD48" box="[959,1097,647,673]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">
<quantity id="A7202BF1D109FFBB93E1B7EDFBF7FD48" box="[959,1034,647,673]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="25" pageNumber="191" unit="m" value="944.0">944 m</quantity>
a.s.l.
</elevation>
;
<date id="1466A0D4D109FFBB9406B7EDFB17FD48" box="[1112,1258,647,673]" pageId="25" pageNumber="191" value="2011-11-09">
<collectingDate id="0422593CD109FFBB9406B7EDFB17FD48" box="[1112,1258,647,673]" pageId="25" pageNumber="191" value="2011-11-09">9 Nov. 2011</collectingDate>
</date>
;
<collectorName id="CD2DE3C2D109FFBB94A8B7EDFEEEFD2D" pageId="25" pageNumber="191">A. Soulier-Perkins</collectorName>
leg.;
<collectionCode id="06C91ED1D109FFBB910DB7C0FE53FD2D" box="[339,430,682,708]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="191">MNHN</collectionCode>
<specimenCode id="307E2E6FD109FFBB91F0B7C0FDC5FD2D" box="[430,568,682,708]" pageId="25" pageNumber="191">(EH) 24703</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="28C2D59FD109FFBB90E3B781FD02F962" pageId="25" pageNumber="191" type="description">
<paragraph id="60678614D109FFBB90E3B781FEB3FCEC" blockId="25.[189,334,747,773]" box="[189,334,747,773]" pageId="25" pageNumber="191">
<heading id="3B2F3178D109FFBB90E3B781FEB3FCEC" bold="true" box="[189,334,747,773]" fontSize="11" level="3" pageId="25" pageNumber="191" reason="3">
<emphasis id="52AC5A06D109FFBB90E3B781FEB3FCEC" bold="true" box="[189,334,747,773]" pageId="25" pageNumber="191">Description</emphasis>
</heading>
</paragraph>
<paragraph id="60678614D109FFBB90E3B670FD45FCDD" blockId="25.[189,696,794,820]" box="[189,696,794,820]" pageId="25" pageNumber="191">TOTAL LENGTH. Between 9.0 and 10.0 mm.</paragraph>
<paragraph id="60678614D109FFBB90E3B636FDDFFBEA" blockId="25.[187,1399,859,1027]" pageId="25" pageNumber="191">HEAD. Median part of vertex strongly domed forming an obtuse angle with postclypeus. Frons concave in its middle. Postclypeus not exceeding anterior margin of head. Furrow on postclypeus wide, rounded at its upper extremity. Lateral bulges parallel, narrower than width of furrow and keeping same width over their entire length. In lateral view, postclypeus convex, forming an obtuse angle slightly pronounced. Rostrum reaching mesocoxae.</paragraph>
<paragraph id="60678614D109FFBB90E3B140FB90FBAD" blockId="25.[189,1133,1065,1092]" box="[189,1133,1065,1092]" pageId="25" pageNumber="191">THORAX. Scutellum thick and main dimple deep; tip of scutellum slightly raised.</paragraph>
<paragraph id="60678614D109FFBB90E3B101FB1AFB41" blockId="25.[189,1399,1131,1193]" pageId="25" pageNumber="191">TEGMINA. Tegmina 2.5 times as long as wide with many tubers all over surface; anal vein on clavus strongly wave-shaped. Apical network dense with more than 20 concave cells per tegmina.</paragraph>
<paragraph id="60678614D109FFBB90E3B1BAFF12F9ED" blockId="25.[189,1400,1232,1540]" pageId="25" pageNumber="191">MALE GENITALIA. Under anal lobe, posterior margin of pygofer forming convex arc; margin becoming concave on ventral part. Subgenital plates shorter than length of pygofer; in lateral view, dorsal margin forming lobe on anterior part. Apical extremity of subgenital plates folded inwards with flattened internal part covered with setae. Sterno-lateral plates extend from pygofer in tapered shape; slightly dorsally oriented with convex apex. Dorsal margin of parameres bearing thin dorsal extension and posterior area excavated in its middle forming two separate bulbs. Aedeagus developed anteriorly in one-pointed protrusion at end of convex arc. Aedeagus bifid on apical part; posterior extension with serrated apex on last quarter. Anterior dorsal extension with a rounded apex representing one quarter length of posterior one.</paragraph>
<paragraph id="60678614D109FFBB90E3B341FD02F962" blockId="25.[189,1398,1579,1675]" pageId="25" pageNumber="191">COLORATION. Head red with end of rostrum and pedicels black. Pronotum and tegmina red; apical network sometimes with black colouration. In ventral view, thorax and abdomen fully red. Red legs with tibias fore and mid-leg black; all tarsi black.</paragraph>
</subSubSection>
<subSubSection id="28C2D59FD109FFBB90E3B3D9FD06F827" pageId="25" pageNumber="191" type="discussion">
<paragraph id="60678614D109FFBB90E3B3D9FE4BF924" blockId="25.[189,438,1715,1741]" box="[189,438,1715,1741]" pageId="25" pageNumber="191">
<heading id="3B2F3178D109FFBB90E3B3D9FE4BF924" bold="true" box="[189,438,1715,1741]" fontSize="11" level="3" pageId="25" pageNumber="191" reason="3">
<emphasis id="52AC5A06D109FFBB90E3B3D9FE4BF924" bold="true" box="[189,438,1715,1741]" pageId="25" pageNumber="191">Nomenclatural note</emphasis>
</heading>
</paragraph>
<paragraph id="60678614D109FFBB90E3B38BFD06F827" blockId="25.[187,1400,1761,1999]" pageId="25" pageNumber="191">
The type specimens designated by Fallou in the monograph of
<taxonomicName id="A7D8FD97D109FFBB93F8B38BFB65F912" authorityName="Fallou" authorityYear="1890" box="[934,1176,1761,1787]" class="Insecta" family="Cercopidae" genus="Monecphora" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="191" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="52AC5A06D109FFBB93F8B38BFB65F912" box="[934,1176,1761,1787]" italics="true" pageId="25" pageNumber="191">Monecphora rugosa</emphasis>
</taxonomicName>
were conserved in his personal collection located in the MNHN. The number of type specimens was not mentioned in his original publication. Nevertheless, in Fallous collection, the pinholes at the bottom of the collection boxes correspond to the number of specimens referenced. Only
<specimenCount id="76DE4D9DD109FFBB9390B221FBA8F88C" box="[974,1109,1867,1893]" count="2" pageId="25" pageNumber="191" type="female">two female</specimenCount>
specimens were placed under the pinned label of
<taxonomicName id="A7D8FD97D109FFBB91AAB204FD1BF861" authorityName="Fallou" authorityYear="1890" box="[500,742,1902,1928]" class="Insecta" family="Cercopidae" genus="Monecphora" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="191" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="52AC5A06D109FFBB91AAB204FD1BF861" box="[500,742,1902,1928]" italics="true" pageId="25" pageNumber="191">Monecphora rugosa</emphasis>
</taxonomicName>
, with no supernumerary pinholes, suggesting that only these
<specimenCount id="76DE4D9DD109FFBB915FB2F8FE50F842" box="[257,429,1937,1964]" count="2" pageId="25" pageNumber="191" type="generic">two specimens</specimenCount>
were considered to be the types. We choose to designate the better-shaped specimen as
<typeStatus id="BF6338B6D109FFBB9085B2DEFEBAF827" box="[219,327,1972,1998]" pageId="25" pageNumber="191" type="lectotype">lectotype</typeStatus>
and the second one as
<typeStatus id="BF6338B6D109FFBB920BB2DEFD0AF827" box="[597,759,1972,1998]" pageId="25" pageNumber="191" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>