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<document id="17386E24DBCDCCADD440AB76E39705CD" ID-CLB-Dataset="107633" ID-DOI="10.11646/zootaxa.5168.5.9" ID-GBIF-Dataset="c31732d0-a394-4a8b-b64b-00ce6c4834da" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911215" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658901058927" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="D975DF2AB95AFFC3928AFB28E6456C73" docLanguage="en" docName="zootaxa.5168.5.9.pdf" docOrigin="Zootaxa 5168 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caloboreaphilus schuelkei Shavrin 2022, sp. n." docType="treatment" docVersion="7" lastPageNumber="600" masterDocId="254CA752B95AFFC0921DFFECE30D6D68" masterDocTitle="A new species of the genus Caloboreaphilus Zerche, 1990 (Coleoptera: Staphylinidae: Coryphiini) from Gansu, China" masterLastPageNumber="600" masterPageNumber="597" pageNumber="597" updateTime="1699362124535" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="33BD1020066A62E128CAC8DFA115AD7C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="121AA7250807A30D58BF6643849F17D7">
<mods:title id="1961EC907F952EB33E372F3C1306D57F">A new species of the genus Caloboreaphilus Zerche, 1990 (Coleoptera: Staphylinidae: Coryphiini) from Gansu, China</mods:title>
</mods:titleInfo>
<mods:name id="6AC776C22BE172D84C09750620B4A0A9" type="personal">
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</mods:role>
<mods:namePart id="FD8F56F55A12CD78D2E2E0508CBD5C29">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="FB091024FD064E9DFA0C99BB1FD98A09">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="002A69FA85DE32184F484E9D4C0BBAB7">
<mods:date id="55A57815719F8FAD6D34DCA41DDF7C42">2022</mods:date>
<mods:detail id="40F8E2E9367F54DBDF50789E82B7A8A7" type="pubDate">
<mods:number id="E0EA3ED9B9C1602228BA6C9190E1E12F">2022-07-26</mods:number>
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<mods:number id="309F02847D3483B2722EE734F9D71C06">5168</mods:number>
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<mods:number id="363A5E8A12A20115234AAC56167F1BDB">5</mods:number>
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<mods:classification id="C6314378D62C343BCBD4DABBD3B61FC0">journal article</mods:classification>
<mods:identifier id="CBB8DDB794E5D552B6A7A836D6AF85E8" type="CLB-Dataset">107633</mods:identifier>
<mods:identifier id="C37CDA7386431F14B94CFA8D4D38EC38" type="DOI">10.11646/zootaxa.5168.5.9</mods:identifier>
<mods:identifier id="F39984F0233DDFEB7FF6D7E15F21E3D4" type="GBIF-Dataset">c31732d0-a394-4a8b-b64b-00ce6c4834da</mods:identifier>
<mods:identifier id="757433D02B1A4FB9C988F85953135E53" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E96B970F67A2631F65607DFCB14ADED2" type="Zenodo-Dep">6911215</mods:identifier>
</mods:mods>
<treatment id="D975DF2AB95AFFC3928AFB28E6456C73" ID-DOI="http://doi.org/10.5281/zenodo.6924003" ID-GBIF-Taxon="197865033" ID-Zenodo-Dep="6924003" LSID="urn:lsid:plazi:treatment:D975DF2AB95AFFC3928AFB28E6456C73" httpUri="http://treatment.plazi.org/id/D975DF2AB95AFFC3928AFB28E6456C73" lastPageId="3" lastPageNumber="600" pageId="0" pageNumber="597">
<subSubSection id="19C63DB7B95AFFC0928AFB28E12E69B7" box="[151,547,1220,1247]" pageId="0" pageNumber="597" type="nomenclature">
<paragraph id="51636E3CB95AFFC0928AFB28E12E69B7" blockId="0.[151,1437,1220,2039]" box="[151,547,1220,1247]" pageId="0" pageNumber="597">
<heading id="0A2BD950B95AFFC0928AFB28E12E69B7" bold="true" box="[151,547,1220,1247]" fontSize="11" level="1" pageId="0" pageNumber="597" reason="1">
<emphasis id="63A8B22EB95AFFC0928AFB28E12E69B7" bold="true" box="[151,547,1220,1247]" pageId="0" pageNumber="597">
<taxonomicName id="96DC15BFB95AFFC0928AFB28E2D469B7" ID-CoL="9GDKS" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,473,1220,1247]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="63A8B22EB95AFFC0928AFB28E2D469B7" bold="true" box="[151,473,1220,1247]" italics="true" pageId="0" pageNumber="597">Caloboreaphilus schuelkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="789B0F55B95AFFC093FDFB29E12E69B7" box="[480,547,1221,1247]" pageId="0" pageNumber="597" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="19C63DB7B95AFFC0928AFB06E20B686B" box="[151,262,1258,1283]" pageId="0" pageNumber="597" type="description">
<paragraph id="51636E3CB95AFFC0928AFB06E20B686B" blockId="0.[151,1437,1220,2039]" box="[151,262,1258,1283]" pageId="0" pageNumber="597">
(
<figureCitation id="C9E772B9B95AFFC09282FB06E3F1686B" box="[159,252,1258,1283]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURE 8" captionStartId-0="1.[151,264,1851,1875]" captionStartId-1="2.[151,250,1990,2014]" captionTargetBox-0="[157,1428,749,1824]" captionTargetBox-1="[151,1435,1244,1967]" captionTargetId-0="figure-296@1.[151,1436,740,1828]" captionTargetId-1="figure-474@2.[151,1435,1244,1967]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." captionText-1="FIGURE 8. Distribution of Caloboreaphilus in China: C. schuelkei (round), C. concisus (square), C. hammondi (traingle)." figureDoi-0="http://doi.org/10.5281/zenodo.6911217" figureDoi-1="http://doi.org/10.5281/zenodo.6911219" httpUri-0="https://zenodo.org/record/6911217/files/figure.png" httpUri-1="https://zenodo.org/record/6911219/files/figure.png" pageId="0" pageNumber="597">Figs 1–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="19C63DB7B95AFFC0928AFAE3E0ED6896" pageId="0" pageNumber="597" type="materials_examined">
<paragraph id="51636E3CB95AFFC0928AFAE3E12268DE" blockId="0.[151,1437,1220,2039]" pageId="0" pageNumber="597">
<emphasis id="63A8B22EB95AFFC0928AFAE3E2B6684E" bold="true" box="[151,443,1294,1319]" inLineHeading="true" pageId="0" pageNumber="597" reason="1">Type material examined.</emphasis>
<materialsCitation id="E1B46461B95AFFC093DAFAE2E12268DE" ID-GBIF-Occurrence="3866796301" pageId="0" pageNumber="597" typeStatus="holotype">
<typeStatus id="8E67D09EB95AFFC093DAFAE2E127684E" box="[455,554,1294,1318]" pageId="0" pageNumber="597" type="holotype">Holotype</typeStatus>
<specimenCount id="47DAA5B5B95AFFC09028FAE3E146684E" box="[565,587,1295,1318]" pageId="0" pageNumber="597" type="female">♀</specimenCount>
: ‘
<emphasis id="63A8B22EB95AFFC09078FAE2E1CA684E" bold="true" box="[613,711,1294,1318]" pageId="0" pageNumber="597">
<collectingCountry id="29CB2EACB95AFFC09078FAE2E1CF684E" box="[613,706,1294,1318]" name="China" pageId="0" pageNumber="597">CHINA</collectingCountry>
:
</emphasis>
<collectingRegion id="9318A0DEB95AFFC090CEFAE3E08E684E" box="[723,899,1294,1319]" country="China" name="Gansu" pageId="0" pageNumber="597">Gansu province</collectingRegion>
, | DaGcaNGLHamo (= | Langmusi env., 34°04.6- 05.1’N 102°37.7- | 38.1’E,
<quantity id="9624C3D9B95AFFC093A6FADEE1466822" box="[443,587,1330,1354]" metricMagnitude="3" metricUnit="m" metricValue="3.554" metricValueMax="3.6439999999999997" metricValueMin="3.464" pageId="0" pageNumber="597" unit="m" value="3554.0" valueMax="3644.0" valueMin="3464.0">3464-3644 m</quantity>
| (GPS), [Ch 5]’ &lt;printed&gt;, ’
<date id="256248FCB95AFFC09161FADEE0F06822" box="[892,1021,1330,1354]" pageId="0" pageNumber="597" value="2005-06-25">
25.
<collectionCode id="37CDF6F9B95AFFC09181FADEE0B16822" box="[924,956,1330,1354]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="0" pageNumber="597">VI</collectionCode>
.2005
</date>
, J. Hájek, | D. Král &amp; J. Růžička leg.; |
<emphasis id="63A8B22EB95AFFC0928AFABAE3D86806" bold="true" box="[151,213,1366,1390]" pageId="0" pageNumber="597">sifted</emphasis>
detritus and wet | moss, wet
<emphasis id="63A8B22EB95AFFC09017FABAE18D6806" bold="true" box="[522,640,1366,1390]" pageId="0" pageNumber="597">coniferous</emphasis>
|
<emphasis id="63A8B22EB95AFFC0908EFABAE1D96806" bold="true" box="[659,724,1366,1390]" pageId="0" pageNumber="597">forest</emphasis>
(
<taxonomicName id="96DC15BFB95AFFC090FEFABBE02D6806" box="[739,800,1367,1390]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="0" pageNumber="597" phylum="Tracheophyta" rank="genus">
<emphasis id="63A8B22EB95AFFC090FEFABBE02D6806" box="[739,800,1367,1390]" italics="true" pageId="0" pageNumber="597">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="96DC15BFB95AFFC09130FABAE0AC6806" authority=", Rho" authorityName="Rho" box="[813,929,1366,1390]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="0" pageNumber="597" phylum="Tracheophyta" rank="genus">
<emphasis id="63A8B22EB95AFFC09130FABAE0656806" box="[813,872,1366,1390]" italics="true" pageId="0" pageNumber="597">Abies</emphasis>
,
<emphasis id="63A8B22EB95AFFC0916BFABAE0AC6806" box="[886,929,1366,1390]" italics="true" pageId="0" pageNumber="597">Rho</emphasis>
</taxonomicName>
- |
<emphasis id="63A8B22EB95AFFC091A1FABAE7226806" box="[956,1071,1366,1390]" italics="true" pageId="0" pageNumber="597">dodendron</emphasis>
) on N slope’ &lt;printed&gt;, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ &lt;printed&gt;, ‘HOLOTYPE |
<emphasis id="63A8B22EB95AFFC091DAFA96E78968FA" box="[967,1156,1402,1426]" italics="true" pageId="0" pageNumber="597">Caloboreraphilus</emphasis>
|
<taxonomicName id="96DC15BFB95AFFC0968AFA96E7F768FA" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[1175,1274,1402,1426]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="63A8B22EB95AFFC0968AFA96E7F768FA" box="[1175,1274,1402,1426]" italics="true" pageId="0" pageNumber="597">schuelkei</emphasis>
</taxonomicName>
<emphasis id="63A8B22EB95AFFC0971CFA97E63B68FB" bold="true" box="[1281,1334,1403,1427]" pageId="0" pageNumber="597">
<taxonomicNameLabel id="789B0F55B95AFFC0971CFA97E63B68FB" box="[1281,1334,1403,1427]" pageId="0" pageNumber="597" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="37CDF6F9B95AFFC092ADFA73E3C868DF" box="[176,197,1439,1463]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="597" type="Museum">V</collectionCode>
. des. 2022’ &lt;red, printed&gt; (cSch).
</materialsCitation>
</paragraph>
<paragraph id="51636E3CB95AFFC092DAFA2FE0ED6896" blockId="0.[151,1437,1220,2039]" pageId="0" pageNumber="597">
<materialsCitation id="E1B46461B95AFFC092DAFA2FE7A668B2" ID-GBIF-Occurrence="3866796303" box="[199,1195,1474,1499]" pageId="0" pageNumber="597" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="8E67D09EB95AFFC092DAFA2FE23E68B3" box="[199,307,1475,1499]" pageId="0" pageNumber="597" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="47DAA5B5B95AFFC0935CFA2FE28D68B2" box="[321,384,1475,1499]" pageId="0" pageNumber="597" type="female">2 ♀♀</specimenCount>
(dissected): same data as the holotype, with additional red printed label
</materialsCitation>
: ‘
<materialsCitation id="E1B46461B95AFFC096A1FA2FE0D06896" ID-GBIF-Occurrence="3866796302" collectionCode="V" collectorName="Shavrin A." pageId="0" pageNumber="597" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="8E67D09EB95AFFC096A1FA2FE64B68B3" box="[1212,1350,1475,1499]" pageId="0" pageNumber="597" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="96DC15BFB95AFFC09742FA2EE2186896" authorityName="Zerche" authorityYear="1990" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="genus">
<emphasis id="63A8B22EB95AFFC09742FA2EE2186896" italics="true" pageId="0" pageNumber="597">Caloboreaphilus</emphasis>
</taxonomicName>
|
<taxonomicName id="96DC15BFB95AFFC09335FA0AE2866896" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[296,395,1510,1534]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="597" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="63A8B22EB95AFFC09335FA0AE2866896" box="[296,395,1510,1534]" italics="true" pageId="0" pageNumber="597">schuelkei</emphasis>
</taxonomicName>
<emphasis id="63A8B22EB95AFFC0938FFA0BE2CA6897" bold="true" box="[402,455,1511,1535]" pageId="0" pageNumber="597">
<taxonomicNameLabel id="789B0F55B95AFFC0938FFA0BE2CA6897" box="[402,455,1511,1535]" pageId="0" pageNumber="597" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FC290BEAB95AFFC093C7FA0AE1466897" box="[474,587,1510,1535]" pageId="0" pageNumber="597">Shavrin A.</collectorName>
<collectionCode id="37CDF6F9B95AFFC09056FA0BE16D6897" box="[587,608,1511,1535]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="0" pageNumber="597" type="Museum">V</collectionCode>
. des. 2022’ (
<specimenCount id="47DAA5B5B95AFFC090F5FA0BE0296896" box="[744,804,1511,1535]" pageId="0" pageNumber="597" type="female">1 ♀♀</specimenCount>
: cSh;
<specimenCount id="47DAA5B5B95AFFC0917BFA0BE09D6896" box="[870,912,1511,1535]" pageId="0" pageNumber="597" type="female">1 ♀</specimenCount>
: cSch)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19C63DB7B95AFFC292DAF9E6E00369AB" lastPageId="2" lastPageNumber="599" pageId="0" pageNumber="597" type="description">
<paragraph id="51636E3CB95AFFC092DAF9E6E03D6B92" blockId="0.[151,1437,1220,2039]" pageId="0" pageNumber="597">
<emphasis id="63A8B22EB95AFFC092DAF9E6E25F6B4A" box="[199,338,1546,1570]" inLineHeading="true" pageId="0" pageNumber="597" reason="1">
<emphasis id="63A8B22EB95AFFC092DAF9E6E2466B4A" bold="true" box="[199,331,1546,1570]" pageId="0" pageNumber="597">Description</emphasis>
.
</emphasis>
Measurements (n=3): maximum width of head including eyes: 0.87–0.96; length of head (from base of labrum to neck constriction along head midline): 0.62–0.67; length of antenna (
<typeStatus id="8E67D09EB95AFFC09611F9C2E77D6B2E" box="[1036,1136,1582,1606]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
): 1.42; ocular length (longitudinal): 0.17; length of pronotum: 0.60–0.74; maximum width of pronotum: 0.87–0.92; sutural length of elytra (length of elytra from the apex of scutellum to the posterior margin of sutural angle): 1.10–1.12; maximum width of elytra: 1.35–1.41; maximum width of abdomen: 1.47–1.64; length of metatibia (
<typeStatus id="8E67D09EB95AFFC09187F976E0F36BDA" box="[922,1022,1690,1714]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
): 0.87; length of metatarsus (
<typeStatus id="8E67D09EB95AFFC09730F976E69C6BDA" box="[1325,1425,1690,1714]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
): 0.59 (combined length of metatarsomeres 1–4: 0.42; length of metatarsomere 5: 0.17); total body length (from anterior margin of clypeus to apex of abdomen): 3.69 (
<typeStatus id="8E67D09EB95AFFC09098F90EE1EA6B92" box="[645,743,1762,1786]" pageId="0" pageNumber="597" type="holotype">holotype</typeStatus>
)–4.75.
</paragraph>
<paragraph id="51636E3CB95AFFC092DAF8EAE1476ABA" blockId="0.[151,1437,1220,2039]" pageId="0" pageNumber="597">
Habitus as in
<figureCitation id="C9E772B9B95AFFC0937CF8EAE2AF6A77" box="[353,418,1798,1823]" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="0" pageNumber="597">Fig. 1</figureCitation>
. Forebody reddish-brown (
<specimenCount id="47DAA5B5B95AFFC090D0F8EBE0566A77" box="[717,859,1799,1823]" pageId="0" pageNumber="597" type="generic" typeStatus="paratype">one paratype</specimenCount>
slightly darker); abdomen dark-brown, with yellowbrown paratergites (
<specimenCount id="47DAA5B5B95AFFC09373F8C7E2F16A2B" box="[366,508,1835,1859]" pageId="0" pageNumber="597" type="generic" typeStatus="paratype">one paratype</specimenCount>
) and intersegmental membranes between tergites III–VII; mouthparts, legs and antennomeres yellow to yellow-brown. Body shiny, without microsculpture, except supra-antennal prominences with traces of diagonal meshes, neck with dense isodiametric sculpture, apical portion of scutellum with fine transverse meshes and abdomen with dense, indistinct or distinct, isodiametric microreticulation. Body covered by moderately dense semierect setae, longer on anterior portion of head.
</paragraph>
<paragraph id="51636E3CB95AFFC192DAF832E61F6FA3" blockId="0.[151,1437,1220,2039]" lastBlockId="1.[151,1437,151,716]" lastPageId="1" lastPageNumber="598" pageId="0" pageNumber="597">
Head very large and convex, 1.4 times as broad as long, about as broad as pronotum, with evenly elevated middle portion and slightly convex supra-antennal prominences; basal portions of clypeus with deep suboval impressions; mediobasal margin of head moderately deeply semicircularly concave. Punctation dense and moderately deep, sparser in middle portion, denser in latero-apical portions between clypeal impressions and margins of eyes. Temples about twice as long as longitudinal length of eyes, very wide and convex. Ocelli small, situated significantly behind level of posterior margins of eyes, distance between ocelli more than twice as long as distance between ocellus and posterior margin of eye. Eyes small, strongly convex. Neck without visible punctation. Labrum strongly transverse, with three short latero-apical, six apical setae, two lateral of which extremely long, and four very small sensile-like setae (
<figureCitation id="C9E772B9B95BFFC191E0FE83E7326CE0" box="[1021,1087,367,392]" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="1" pageNumber="598">Fig. 2</figureCitation>
). Mandible as in
<figureCitation id="C9E772B9B95BFFC196EAFE83E63A6CE0" box="[1271,1335,367,392]" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="1" pageNumber="598">Fig. 3</figureCitation>
. Mentum large, trapezoidal, with rounded laterobasal margins, strongly narrowed apicad, with deeply sinuate lateral margins; labium narrow, with elongate labial palpomeres, apical palpomere very narrow, distinctly shorter than preapical segment, glossa narrow and strongly protruded apicad (
<figureCitation id="C9E772B9B95BFFC1909CFE37E1CF6C9C" box="[641,706,475,500]" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="1" pageNumber="598">Fig. 4</figureCitation>
). Preapical maxillary palpomere very wide, slightly longer than preceding palpomere; galea moderately narrow and slightly longer than lacinia; lacinia slightly broader than galea in middle portion, with elongate spine-like apex, five moderately long preapical spines and long setation along medial margin (
<figureCitation id="C9E772B9B95BFFC1976CFDCFE3AB6F37" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
). Antenna exceeding basal margin of pronotum when reclined, antennomeres 4–10 covered by dense pubescence; basal antennomere 1.7 times as long as 2, antennomere 2 subovoid, 3 slightly longer than 2, 4 short, slightly longer than broad, 5–6 indistinctly broader and shorter than 4, 7–8 slightly broader than 6, 9–10 slightly longer and distinctly broader than 8, apical antennomere twice as long as 10, from middle gradually narrowed toward rounded or subacute apex.
</paragraph>
<caption id="05A33EB4B95BFFC1928AF8D7E0736AF3" ID-DOI="http://doi.org/10.5281/zenodo.6911217" ID-Zenodo-Dep="6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="1" pageNumber="598" startId="1.[151,264,1851,1875]" targetBox="[157,1428,749,1824]" targetPageId="1">
<paragraph id="51636E3CB95BFFC1928AF8D7E0736AF3" blockId="1.[151,1437,1851,1947]" pageId="1" pageNumber="598">
<emphasis id="63A8B22EB95BFFC1928AF8D7E2306A3B" bold="true" box="[151,317,1851,1875]" pageId="1" pageNumber="598">FIGURES 1–7.</emphasis>
<taxonomicName id="96DC15BFB95BFFC19358F8D7E1586A3B" authorityName="Shavrin" authorityYear="2022" box="[325,597,1851,1875]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="63A8B22EB95BFFC19358F8D7E1586A3B" box="[325,597,1851,1875]" italics="true" pageId="1" pageNumber="598">Caloboreaphilus schuelkei</emphasis>
</taxonomicName>
: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7).
</paragraph>
</caption>
<paragraph id="51636E3CB958FFC292DAFF74E6646CC3" blockId="2.[151,1437,151,1219]" pageId="2" pageNumber="599">
Pronotum 1.2–1.4 times as broad as long, significantly convex in middle, with very deep and wide medial impression, stretching from about apical margin toward middle, and with deep semicircular impression in mediobasal third, mediolateral portions with very wide and deep depressions; pronotum widest in middle, somewhat angulate, significantly narrowed posteriad toward widely rounded hind angles; latero-apical portions narrowed gradually toward neck; apical margin rounded, significantly narrower than somewhat straight or slightly rounded posterior margin; lateral edges with regular, very strong and rounded crenulation, larger on latero-apical and mediolateral and finer on latero-basal margins. Punctation denser, coarser and deeper than that on middle of head (
<typeStatus id="8E67D09EB958FFC29176FE83E0C76CEF" box="[875,970,367,391]" pageId="2" pageNumber="599" type="holotype">holotype</typeStatus>
with more sparser punctation on bottom of medioapical impression), distance between punctures in middle about as long as nearest puncture or slightly longer.
</paragraph>
<paragraph id="51636E3CB958FFC292DAFE5BE20B6C9B" blockId="2.[151,1437,151,1219]" pageId="2" pageNumber="599">Scutellum large, with rounded apex and distinct semicircular impression in middle, without or with several very fine punctures.</paragraph>
<paragraph id="51636E3CB958FFC292DAFE13E3FE6FCF" blockId="2.[151,1437,151,1219]" pageId="2" pageNumber="599">Elytra slightly convex, 1.5–1.8 times as long as pronotum, 1.2 times as long as broad, slightly or significantly broadened posteriad, reaching apical margin of abdominal tergite III or IV; apical margin straight; lateral portions narrowly flattened, with distinctly bordered margins; laterobasal margins with very fine acute crenulation. Punctation very dense, larger and deeper than that on pronotum, somewhat denser and coarser in basal portion near scutellum. Wings fully developed.</paragraph>
<paragraph id="51636E3CB958FFC292DAFD58E68F6FA3" blockId="2.[151,1437,151,1219]" box="[199,1410,691,716]" pageId="2" pageNumber="599">Metatarsus 1.4 times as long as metatibia; apical metatarsomere slightly shorter than three preceding tarwomeres.</paragraph>
<paragraph id="51636E3CB958FFC292DAFD3BE2316E5F" blockId="2.[151,1437,151,1219]" pageId="2" pageNumber="599">Abdomen distinctly broader than elytra, with fine and dense punctation, larger on tergite V, with transverse wingfolding spots in middle of tergites III and IV (slightly more transverse in III); apical margin of tergite VII with very narrow palisade fringe.</paragraph>
<paragraph id="51636E3CB958FFC292DAFCAFE2626E33" blockId="2.[151,1437,151,1219]" box="[199,367,835,859]" pageId="2" pageNumber="599">Male unknown.</paragraph>
<paragraph id="51636E3CB958FFC292DAFC8BE0346ECB" blockId="2.[151,1437,151,1219]" pageId="2" pageNumber="599">
Female. Apical margins of abdominal tergite VIII (
<figureCitation id="C9E772B9B958FFC290F0FC8BE0236E17" box="[749,814,871,895]" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="2" pageNumber="599">Fig. 6</figureCitation>
) and sternite VIII (
<figureCitation id="C9E772B9B958FFC291E2FC8BE74D6EE8" box="[1023,1088,871,896]" captionStart="FIGURES 1–7" captionStartId="1.[151,264,1851,1875]" captionTargetBox="[157,1428,749,1824]" captionTargetId="figure-296@1.[151,1436,740,1828]" captionTargetPageId="1" captionText="FIGURES 1–7. Caloboreaphilus schuelkei: 1—habitus (paratype), 2—labrum, dorsal view, 3—right mandible, dorsal view, 4—labium and mentum, ventral view, 5—left maxilla, ventral view, 6—abdominal tergite VIII, dorsal view, 7—abdominal sternite IX, ventral view. Scale bars: 1.0 mm (Fig. 1); 0.1 mm (Figs 2–7)." figureDoi="http://doi.org/10.5281/zenodo.6911217" httpUri="https://zenodo.org/record/6911217/files/figure.png" pageId="2" pageNumber="599">Fig. 7</figureCitation>
) rounded. Genital segment with long gonocoxites and elongate styli, each with long apical seta.
</paragraph>
<paragraph id="51636E3CB958FFC292DAFC43E00369AB" blockId="2.[151,1437,151,1219]" pageId="2" pageNumber="599">
<emphasis id="63A8B22EB958FFC292DAFC43E2AB6EA0" bold="true" box="[199,422,943,968]" inLineHeading="true" pageId="2" pageNumber="599" reason="1">Comparative notes.</emphasis>
<taxonomicName id="96DC15BFB958FFC293B0FC43E1C46EAF" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[429,713,943,967]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="63A8B22EB958FFC293B0FC43E1C46EAF" box="[429,713,943,967]" italics="true" pageId="2" pageNumber="599">Caloboreaphilus schuelkei</emphasis>
</taxonomicName>
<emphasis id="63A8B22EB958FFC290CDFC5CE0086EA0" bold="true" box="[720,773,944,968]" pageId="2" pageNumber="599">
<taxonomicNameLabel id="789B0F55B958FFC290CDFC5CE0086EA0" box="[720,773,944,968]" pageId="2" pageNumber="599" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
can be distinguished from the other congeners by the broader postocular portion of the head, the different shape of the mentum, the pronotum with more narrowed latero-apical margins, deeper medial impression, larger lateral crenulation, and larger and deeper punctation, and the shape of the elytra more broadened posteriad. Based on the head about as broad as the pronotum and angulate lateral margins of the pronotum, it is somewhat similar to
<taxonomicName id="96DC15BFB958FFC2938DFBACE12C693F" authorityName="Zerche" authorityYear="1990" box="[400,545,1087,1111]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="63A8B22EB958FFC2938DFBACE2AF693F" box="[400,418,1088,1111]" italics="true" pageId="2" pageNumber="599">C</emphasis>
.
<emphasis id="63A8B22EB958FFC293ADFBD3E12C693F" box="[432,545,1087,1111]" italics="true" pageId="2" pageNumber="599">hammondi</emphasis>
</taxonomicName>
, from which it can be distinguished by the significantly larger body. Regarding the body size and the presence of coarse crenulation on the lateral margins of the pronotum, the new species is also similar to
<taxonomicName id="96DC15BFB958FFC2928AFB64E21E69F7" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[151,275,1160,1183]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="concisus">
<emphasis id="63A8B22EB958FFC2928AFB64E3A469F7" box="[151,169,1160,1183]" italics="true" pageId="2" pageNumber="599">C</emphasis>
.
<emphasis id="63A8B22EB958FFC292AAFB64E21E69F7" box="[183,275,1160,1183]" italics="true" pageId="2" pageNumber="599">concisus</emphasis>
</taxonomicName>
, from which it can be distinguished by the broader forebody, more transverse pronotum being widest at the middle, and the shape of the elytra more broadened apicad.
</paragraph>
</subSubSection>
<caption id="05A33EB4B958FFC2928AF82AE67E6AB6" ID-DOI="http://doi.org/10.5281/zenodo.6911219" ID-Zenodo-Dep="6911219" box="[151,1395,1990,2014]" httpUri="https://zenodo.org/record/6911219/files/figure.png" pageId="2" pageNumber="599" startId="2.[151,250,1990,2014]" targetBox="[151,1435,1244,1967]" targetPageId="2">
<paragraph id="51636E3CB958FFC2928AF82AE67E6AB6" blockId="2.[151,1395,1990,2014]" box="[151,1395,1990,2014]" pageId="2" pageNumber="599">
<emphasis id="63A8B22EB958FFC2928AF82AE21E6AB6" bold="true" box="[151,275,1990,2014]" pageId="2" pageNumber="599">FIGURE 8.</emphasis>
Distribution of
<taxonomicName id="96DC15BFB958FFC293ABF82AE1526AB6" authorityName="Zerche" authorityYear="1990" box="[438,607,1990,2014]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="genus">
<emphasis id="63A8B22EB958FFC293ABF82AE1526AB6" box="[438,607,1990,2014]" italics="true" pageId="2" pageNumber="599">Caloboreaphilus</emphasis>
</taxonomicName>
in China:
<taxonomicName id="96DC15BFB958FFC290D4F82BE0486AB6" authorityName="Shavrin" authorityYear="2022" box="[713,837,1990,2014]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="schuelkei">
<emphasis id="63A8B22EB958FFC290D4F82BE1D76AB6" box="[713,730,1991,2014]" italics="true" pageId="2" pageNumber="599">C</emphasis>
.
<emphasis id="63A8B22EB958FFC290FAF82AE0486AB6" box="[743,837,1990,2014]" italics="true" pageId="2" pageNumber="599">schuelkei</emphasis>
</taxonomicName>
(round),
<taxonomicName id="96DC15BFB958FFC291B9F82BE7146AB6" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[932,1049,1991,2014]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="concisus">
<emphasis id="63A8B22EB958FFC291B9F82BE0B86AB6" box="[932,949,1991,2014]" italics="true" pageId="2" pageNumber="599">C</emphasis>
.
<emphasis id="63A8B22EB958FFC291DFF82BE7146AB6" box="[962,1049,1991,2014]" italics="true" pageId="2" pageNumber="599">concisus</emphasis>
</taxonomicName>
(square),
<taxonomicName id="96DC15BFB958FFC29662F82BE60A6AB6" authorityName="Zerche" authorityYear="1990" box="[1151,1287,1990,2014]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="63A8B22EB958FFC29662F82BE79D6AB6" box="[1151,1168,1991,2014]" italics="true" pageId="2" pageNumber="599">C</emphasis>
.
<emphasis id="63A8B22EB958FFC29681F82AE60A6AB6" box="[1180,1287,1990,2014]" italics="true" pageId="2" pageNumber="599">hammondi</emphasis>
</taxonomicName>
(traingle).
</paragraph>
</caption>
<subSubSection id="19C63DB7B959FFC392DAFF7BE1446D9F" pageId="3" pageNumber="600" type="distribution">
<paragraph id="51636E3CB959FFC392DAFF7BE6156DC7" blockId="3.[151,1436,151,284]" box="[199,1304,151,176]" pageId="3" pageNumber="600">
<emphasis id="63A8B22EB959FFC392DAFF7BE2576DC7" bold="true" box="[199,346,151,175]" inLineHeading="true" pageId="3" pageNumber="600" reason="1">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="8E67D09EB959FFC3919EFF74E0BC6DD8" box="[899,945,152,176]" pageId="3" pageNumber="600">type</typeStatus>
locality in
<collectingRegion id="9318A0DEB959FFC39637FF7BE77C6DC7" box="[1066,1137,151,175]" country="China" name="Gansu" pageId="3" pageNumber="600">Gansu</collectingRegion>
,
<collectingCountry id="29CB2EACB959FFC39661FF7BE7B16DC7" box="[1148,1212,151,175]" name="China" pageId="3" pageNumber="600">China</collectingCountry>
(
<figureCitation id="C9E772B9B959FFC396D7FF7BE6066DC7" box="[1226,1291,151,175]" captionStart="FIGURE 8" captionStartId="2.[151,250,1990,2014]" captionTargetBox="[151,1435,1244,1967]" captionTargetId="figure-474@2.[151,1435,1244,1967]" captionTargetPageId="2" captionText="FIGURE 8. Distribution of Caloboreaphilus in China: C. schuelkei (round), C. concisus (square), C. hammondi (traingle)." figureDoi="http://doi.org/10.5281/zenodo.6911219" httpUri="https://zenodo.org/record/6911219/files/figure.png" pageId="3" pageNumber="600">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="51636E3CB959FFC392DAFF57E1446D9F" blockId="3.[151,1436,151,284]" pageId="3" pageNumber="600">
<emphasis id="63A8B22EB959FFC392DAFF57E2486DBB" bold="true" box="[199,325,187,211]" inLineHeading="true" pageId="3" pageNumber="600" reason="1">Bionomics.</emphasis>
Specimens were collected at an elevation from
<quantity id="9624C3D9B959FFC39150FF57E0F46DBB" box="[845,1017,187,211]" metricMagnitude="3" metricUnit="m" metricValue="3.554" metricValueMax="3.6439999999999997" metricValueMin="3.464" pageId="3" pageNumber="600" unit="m" value="3554.0" valueMax="3644.0" valueMin="3464.0">3464 to 3644 m</quantity>
a.s.l. and were sifted from the detritus and wet mosses in wet coniferous forest.
</paragraph>
</subSubSection>
<subSubSection id="19C63DB7B959FFC392DAFEEFE6456C73" box="[199,1352,259,284]" pageId="3" pageNumber="600" type="etymology">
<paragraph id="51636E3CB959FFC392DAFEEFE6456C73" blockId="3.[151,1436,151,284]" box="[199,1352,259,284]" pageId="3" pageNumber="600">
<emphasis id="63A8B22EB959FFC392DAFEEFE2456C73" bold="true" box="[199,328,259,283]" inLineHeading="true" pageId="3" pageNumber="600" reason="1">Etymology.</emphasis>
Patronymic, the species is named to honour my colleague, Michael Schülke (
<collectingRegion id="9318A0DEB959FFC3969EFEEFE7C76C73" box="[1155,1226,259,283]" country="Germany" name="Berlin" pageId="3" pageNumber="600">Berlin</collectingRegion>
,
<collectingCountry id="29CB2EACB959FFC396C8FEEFE6306C73" box="[1237,1341,259,283]" name="Germany" pageId="3" pageNumber="600">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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