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<document id="2051225B0532029ED5484BC54C9E779E" ID-CLB-Dataset="46361" ID-DOI="10.11646/zootaxa.3768.2.6" ID-GBIF-Dataset="807bb695-5592-4089-9142-010a98add339" ID-ISSN="1175-5326" ID-Zenodo-Dep="225197" ID-ZooBank="12362EE3-1A25-4DC6-8413-4E1264826680" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460388721214" checkinUser="plazi" docAuthor="Sugaya, Kazuki &amp; Yamasako, Junsuke" docDate="2014" docId="BA388799F94FFFEB4C98210D1791FE09" docLanguage="en" docName="zt03768p195.pdf" docOrigin="Zootaxa 3768 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ishikawatrechus ishiharai Ueno 1994" docType="treatment" docVersion="9" lastPageNumber="195" masterDocId="4601FFE1F94AFFED4C0F275D1627FFC6" masterDocTitle="A new species of the genus Ishikawatrechus (Coleoptera, Trechinae) from Japan" masterLastPageNumber="195" masterPageNumber="189" pageNumber="194" updateTime="1698297955780" updateUser="ExternalLinkService">
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<mods:title id="581D288DF7E4FA39FA648706EC1EB996">A new species of the genus Ishikawatrechus (Coleoptera, Trechinae) from Japan</mods:title>
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<mods:namePart id="FF0494767A0566DC1FC3B766D7FB2DAA">Sugaya, Kazuki</mods:namePart>
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<mods:namePart id="BF7EF4DF08AF0A1611DD49C576F00D5D">Yamasako, Junsuke</mods:namePart>
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<paragraph id="322E368FF94FFFE84C98210D1441F9AD" blockId="5.[151,614,1616,1677]" box="[151,614,1616,1643]" pageId="5" pageNumber="194">
<heading id="696681E3F94FFFE84C98210D1441F9AD" bold="true" box="[151,614,1616,1643]" fontSize="11" level="1" pageId="5" pageNumber="194" reason="1">
<taxonomicName id="F5914D0CF94FFFE84C98210D1441F9AD" ID-CoL="3Q73S" authority="Ueno, 1994" authorityName="Ueno" authorityYear="1994" box="[151,614,1616,1643]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis id="00E5EA9DF94FFFE84C98210D1441F9AD" bold="true" box="[151,614,1616,1643]" pageId="5" pageNumber="194">
<emphasis id="00E5EA9DF94FFFE84C98210D17F4F9AC" bold="true" box="[151,467,1616,1642]" italics="true" pageId="5" pageNumber="194">Ishikawatrechus ishiharai</emphasis>
<bibRefCitation id="56004B7EF94FFFE84DD4210C1441F9AD" author="Ueno" box="[475,614,1617,1643]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1994) Ishikawatrechus ishiharai, a new anophthalmic trechine beetle from the Ishizuchi Mountains, Southwest Japan (Coleoptera, Trechinae). Transactions of the Shikoku Entomological Society, 20, 339 - 344." type="journal article" year="1994">Uéno, 1994</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="322E368FF94FFFE84C9821291783F94B" blockId="5.[151,614,1616,1677]" box="[151,420,1652,1677]" pageId="5" pageNumber="194">
(
<figureCitation id="AAAA2A0AF94FFFE84C902129176DF94A" box="[159,330,1652,1677]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 10. Genital organs of Ishikawatrechus spp. 3 4, 7 8, I. bidilatatus sp. nov.; 5 6, 9 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="5" pageNumber="194">Figs 56, 910</figureCitation>
,
<figureCitation id="AAAA2A0AF94FFFE84D5A212917BBF94B" box="[341,412,1652,1677]" captionStart="FIGURES 11 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 19. Internal sac of Ishikawatrechus spp. 11 15, I. bidilatatus sp. nov.; 16 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">1619</figureCitation>
)
</paragraph>
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<paragraph id="322E368FF94FFFE84C9821E11494F914" blockId="5.[151,691,1724,1746]" box="[151,691,1724,1746]" pageId="5" pageNumber="194">
<treatmentCitationGroup id="128111A1F94FFFE84C9821E11494F914" box="[151,691,1724,1746]" pageId="5" pageNumber="194">
<taxonomicName id="F5914D0CF94FFFE84C9821E11477F914" ID-CoL="3Q73S" authority="Ueno, 1994: 339" authorityName="Ueno" authorityPageNumber="339" authorityYear="1994" box="[151,592,1724,1746]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="ishiharai">
<emphasis id="00E5EA9DF94FFFE84C9821E117BAF914" box="[151,413,1724,1746]" italics="true" pageId="5" pageNumber="194">Ishikawatrechus ishiharai</emphasis>
<treatmentCitation id="B330109EF94FFFE84DAB21E11477F914" author="Ueno" box="[420,592,1724,1746]" page="339" pageId="5" pageNumber="194" year="1994">
<bibRefCitation id="56004B7EF94FFFE84DAB21E1143AF914" author="Ueno" box="[420,541,1724,1746]" pageId="5" pageNumber="194" refString="Ueno, S. - I. (1994) Ishikawatrechus ishiharai, a new anophthalmic trechine beetle from the Ishizuchi Mountains, Southwest Japan (Coleoptera, Trechinae). Transactions of the Shikoku Entomological Society, 20, 339 - 344." type="journal article" year="1994">Uéno, 1994</bibRefCitation>
: 339
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</taxonomicName>
; 2008: 14
</treatmentCitationGroup>
</paragraph>
<paragraph id="322E368FF94FFFE84C9821A3133FF899" blockId="5.[151,1436,1790,2031]" pageId="5" pageNumber="194">
<emphasis id="00E5EA9DF94FFFE84C9821A317B0F8D1" bold="true" box="[151,407,1790,1815]" pageId="5" pageNumber="194">Specimens examined.</emphasis>
<specimenCount id="2497FD06F94FFFE84D9221A217E5F8D1" box="[413,450,1791,1815]" pageId="5" pageNumber="194" type="generic">1♂</specimenCount>
, Iwaguro-yama, alt. ca.
<quantity id="F5699B6AF94FFFE84EDD21A31516F8D0" box="[722,817,1790,1815]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="194" unit="m" value="1550.0">1,550 m</quantity>
, Wakayama, Kumakogen-chô, Kamiukena-gun, Ehime Pref.,
<date id="462F104FF94FFFE84CD4207E174CF8FD" box="[219,363,1827,1851]" pageId="5" pageNumber="194" value="2011-09-24">24. IX. 2011</date>
, K. Sugaya leg.; 1♀, same locality but
<date id="462F104FF94FFFE84F2C207E1595F8FD" box="[803,946,1827,1851]" pageId="5" pageNumber="194" value="2011-11-21">21. XI. 2011</date>
, H. Hara leg.;
<specimenCount id="2497FD06F94FFFE84859207E125BF8FD" box="[1110,1148,1827,1851]" pageId="5" pageNumber="194" type="generic">2♂</specimenCount>
, 1♀, Odama-dani, alt. ca.
<quantity id="F5699B6AF94FFFE84C98201B16D2F898" box="[151,245,1862,1887]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="194" unit="m" value="1500.0">1,500 m</quantity>
, Nishinokawa-chô, Saijô-shi, Ehime Pref.,
<date id="462F104FF94FFFE84ED6201A154DF899" box="[729,874,1863,1887]" pageId="5" pageNumber="194" value="2012-09-12">12. IX. 2012</date>
, K. Sugaya leg. (our own collections).
</paragraph>
<paragraph id="322E368FF94FFFEB4CC82037173FFE86" blockId="5.[151,1436,1790,2031]" lastBlockId="6.[151,1436,151,464]" lastPageId="6" lastPageNumber="195" pageId="5" pageNumber="194">
<emphasis id="00E5EA9DF94FFFE84CC8203717C7F845" bold="true" box="[199,480,1898,1923]" pageId="5" pageNumber="194">Additional description.</emphasis>
Male genitalia (n = 3,
<figureCitation id="AAAA2A0AF94FFFE84EF120371540F845" box="[766,871,1898,1923]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 10. Genital organs of Ishikawatrechus spp. 3 4, 7 8, I. bidilatatus sp. nov.; 5 6, 9 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="5" pageNumber="194">Figs 56</figureCitation>
,
<figureCitation id="AAAA2A0AF94FFFE84F77203715E6F845" box="[888,961,1898,1923]" captionStart="FIGURES 11 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 19. Internal sac of Ishikawatrechus spp. 11 15, I. bidilatatus sp. nov.; 16 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">1619</figureCitation>
). Internal sac in fully inflated condition through eversion (n = 3,
<figureCitation id="AAAA2A0AF94FFFE84DB620D3141EF861" box="[441,569,1934,1959]" captionStart="FIGURES 11 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 19. Internal sac of Ishikawatrechus spp. 11 15, I. bidilatatus sp. nov.; 16 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="5" pageNumber="194">Figs 1619</figureCitation>
) about twice as long as aedeagus, elongated pyriform, asymmetric, twisted leftwards behind middle; basal half swollen into elongated tube-shaped, without any projections and copulatory pieces, covered with spicules excepted for small crescent area on basal 1/3 of left lateral side; apical half well swollen into an elongated bulb, with spicules on the longitudinal area which is extending from basal half to periphery of gonopore through ventral side; gonopore (GP,
<figureCitation id="AAAA2A0AF94CFFEB4F3F27E115FFFF12" box="[816,984,188,213]" captionStart="FIGURES 11 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 19. Internal sac of Ishikawatrechus spp. 11 15, I. bidilatatus sp. nov.; 16 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="6" pageNumber="195">Figs 1617, 19</figureCitation>
) opening on ventral side. Spicules (
<figureCitation id="AAAA2A0AF94CFFEB496527E116FAFF3E" captionStart="FIGURES 11 19" captionStartId="4.[151,264,1743,1766]" captionTargetBox="[159,1419,200,1714]" captionTargetId="figure@4.[151,1436,193,1722]" captionTargetPageId="4" captionText="FIGURES 11 19. Internal sac of Ishikawatrechus spp. 11 15, I. bidilatatus sp. nov.; 16 19, I. ishiharai. 11, 16, Right lateral view; 12, 17, Left lateral view; 13, 18, Dorsal view; 14, 19, Ventral view; 15, Copulatory piece of I. bidilatatus. Abbreviations; CP, copulatory piece; GP, gonopore." httpUri="https://zenodo.org/record/225200/files/figure.png" pageId="6" pageNumber="195">Figs 1619</figureCitation>
) on basal half usually fine, scaly and weakly sclerotized, but those on the area obliquely and curvedly extending from dorsal left side to dorsum distinctly spinous and well sclerotized; those on apical half rudimentary sclerotized.
</paragraph>
<paragraph id="322E368FF94CFFEB4CC826111791FE09" blockId="6.[151,1436,151,464]" pageId="6" pageNumber="195">
Female genitalia (n = 2,
<figureCitation id="AAAA2A0AF94CFFEB4DD72611146CFEA2" box="[472,587,332,357]" captionStart="FIGURES 3 10" captionStartId="3.[151,264,1894,1917]" captionTargetBox="[161,1430,200,1870]" captionTargetId="figure@3.[151,1436,193,1874]" captionTargetPageId="3" captionText="FIGURES 3 10. Genital organs of Ishikawatrechus spp. 3 4, 7 8, I. bidilatatus sp. nov.; 5 6, 9 10, I. ishiharai. 3, 5, Aedeagus in left lateral view; 4, 6, Apex of aedeagus in dorsal view; 7, 9, Left stylus in ventral view; 8, 10, Ditto in dorsal view. Abbreviations; GC 1, basal gonocoxite 1; GC 2, apical gonocoxite 2." httpUri="https://zenodo.org/record/225199/files/figure.png" pageId="6" pageNumber="195">Figs 910</figureCitation>
). GC1 near quadrate, with 4 short setae on ventral side of apical margin and 2 short setae on dorsal side of inner margin near apex; GC2 slender, slightly curved outwardly, pointed at apex, well emarginated at middle of outer margin, with 2 ensiform setae on dorsal side near base and 2 nematiform setae on ventral side near apex.
</paragraph>
</subSubSection>
</treatment>
</document>