179 lines
20 KiB
XML
179 lines
20 KiB
XML
<document id="52906D5766BBBBDCC8CEEC36AAC4D13C" ID-CLB-Dataset="5809" ID-DOI="10.2478/aemnp-2018-0028" ID-GBIF-Dataset="54b60ce2-158f-4ce8-a9b0-f42b60242162" ID-ISSN="1804-6487" ID-Zenodo-Dep="4504772" ID-ZooBank="71176108-A738-46FF-89EF-4658841391F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612473328331" checkinUser="felipe" docAuthor="Chigray, Svetlana, Nabozhenko, Maxim, Keskin, Bekir & Abdurakhmanov, Gayirbeg" docDate="2018" docId="960EFA1AFF9789197305FF44FAEBFD66" docLanguage="en" docName="ActaEntMusNatPra.58.2.331-346.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 58 (2)" docSource="http://dx.doi.org/10.2478/aemnp-2018-0028" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Calyptopsis kaszabi Chigray & Nabozhenko & Keskin & Abdurakhmanov 2018, sp. nov." docType="treatment" docVersion="9" lastPageNumber="336" masterDocId="6A378262FF92891C722BFFF9FFA7FFF3" masterDocTitle="Taxonomic review of the genus Calyptopsis of Turkey and adjacent areas (Coleoptera: Tenebrionidae: Tentyriini)" masterLastPageNumber="346" masterPageNumber="331" pageNumber="336" updateTime="1698927389047" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="8669905EEEBE4DEF536D76215F5723B5">Taxonomic review of the genus Calyptopsis of Turkey and adjacent areas (Coleoptera: Tenebrionidae: Tentyriini)</mods:title>
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<mods:name id="7F27D1E8E4BBA646C4E55CE2FE7538FB" type="personal">
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<mods:roleTerm id="F975837E26857707A52D93056A88D454">Author</mods:roleTerm>
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<mods:namePart id="9A0E22947C0FE01E8B1FEFD17D713D6D">Chigray, Svetlana</mods:namePart>
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<mods:affiliation id="2411DABCD1434A5883617457FF925966">St Petersburg State University, Lieutenant Schmidt emb., 11 /</mods:affiliation>
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</mods:role>
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<mods:namePart id="2F9FE04FB263AEDA461982F66CA54323">Nabozhenko, Maxim</mods:namePart>
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<mods:affiliation id="5F48727B2C9AA5CD10A3E0ADCF38A507">St. Petersburg 199034 Russia; e-mail: s. voloboeva 95 @ mail. ru & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia</mods:affiliation>
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</mods:name>
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<mods:name id="85A527780C7C8BD4C6D5631A20DC3CCD" type="personal">
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<mods:roleTerm id="2D160D92D934FB3D67CF055A7E9969E4">Author</mods:roleTerm>
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<mods:namePart id="F0E338C93FC147BCDD336F8BD804364F">Keskin, Bekir</mods:namePart>
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<mods:affiliation id="BEA4F36D13CFE16C821D85FE45CBB99B">Ege University, Bornova-Izmir 35100 Turkey; e-mail: bekir. keskin. phd @ gmail. com</mods:affiliation>
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<mods:namePart id="8F3F7E7E26A15AD97BF128B890264400">Abdurakhmanov, Gayirbeg</mods:namePart>
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<mods:affiliation id="8A1BFE86342087C38066132FF576F203">St. Petersburg 199034 Russia; e-mail: s. voloboeva 95 @ mail. ru & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia</mods:affiliation>
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<mods:title id="9B0243785BCC79F5A16F0994A7049692">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="AA8F28FEEDA8E3CE71FE77B084FDBFEE">2018</mods:date>
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<mods:title id="565A4EA7577D6D882AC62290B859E60A">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="BC46F535D2A709ECA56A852821C5A0BB">2018-08-29</mods:number>
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<mods:url id="37315EBC4FD0DC6CC3182C6F6E4CA430">http://dx.doi.org/10.2478/aemnp-2018-0028</mods:url>
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<treatment id="960EFA1AFF9789197305FF44FAEBFD66" ID-DOI="http://doi.org/10.5281/zenodo.5061721" ID-GBIF-Taxon="183237286" ID-Zenodo-Dep="5061721" LSID="urn:lsid:plazi:treatment:960EFA1AFF9789197305FF44FAEBFD66" httpUri="http://treatment.plazi.org/id/960EFA1AFF9789197305FF44FAEBFD66" lastPageNumber="336" pageId="5" pageNumber="336">
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<subSubSection id="56BD1887FF9789197305FF44FDC6FF26" box="[302,609,189,213]" pageId="5" pageNumber="336" type="nomenclature">
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<paragraph id="1E184B0CFF9789197305FF44FDC6FF26" blockId="5.[302,609,189,213]" box="[302,609,189,213]" pageId="5" pageNumber="336">
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<heading id="4550FC60FF9789197305FF44FDC6FF26" box="[302,609,189,213]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="336" reason="3">
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<emphasis id="2CD3971EFF9789197305FF44FDC6FF26" bold="true" box="[302,609,189,213]" pageId="5" pageNumber="336">
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<taxonomicName id="D9A7308FFF9789197305FF44FDA3FF26" authority="Chigray & Nabozhenko & Keskin & Abdurakhmanov, 2018" authorityName="Chigray & Nabozhenko & Keskin & Abdurakhmanov" authorityYear="2018" box="[302,516,189,213]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="kaszabi" status="sp. nov.">
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<emphasis id="2CD3971EFF9789197305FF44FDA3FF26" bold="true" box="[302,516,189,213]" italics="true" pageId="5" pageNumber="336">Calyptopsis kaszabi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="37E02A65FF9789197020FF47FDC6FF26" box="[523,609,190,213]" pageId="5" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="56BD1887FF97891973ADFF25FDAEFF1C" box="[390,521,220,239]" pageId="5" pageNumber="336" type="description">
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<paragraph id="1E184B0CFF97891973ADFF25FDAEFF1C" blockId="5.[390,521,220,239]" box="[390,521,220,239]" pageId="5" pageNumber="336">
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(
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<figureCitation id="869C5789FF97891973A7FF25FE6AFF1C" box="[396,461,220,239]" captionStart="Fig" captionStartId="2.[151,182,1719,1738]" captionTargetBox="[161,1358,199,1681]" captionTargetId="figure-13@2.[151,1361,189,1689]" captionTargetPageId="2" captionText="Fig. 1. Calyptopsis spp., habitus. A – C. capnisiformis capnisiformis Reitter, 1903, ♁; B – C. capnisiformis volcanica subsp. nov., ♁; C – C. caucasica (Kraatz, 1865), ♁; D – C. emarginata Reitter, 1889, ♁; E – C. emarginata, ♀; F – C. kaszabi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4504774" httpUri="https://zenodo.org/record/4504774/files/figure.png" pageId="5" pageNumber="336">Figs 1F</figureCitation>
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,
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<figureCitation id="869C5789FF97891973FDFF25FE78FF1C" box="[470,479,220,239]" captionStart="Fig" captionStartId="11.[151,182,2027,2046]" captionTargetBox="[237,1344,461,2000]" captionTargetId="figure-16@11.[237,1352,458,2009]" captionTargetPageId="11" captionText="Fig. 8. Calyptopsis spp., details of structure, ♁. A – C. solieri Reiche & Saulcy, 1857, mesoventrite; B – C. escherichi Reitter, 1900, protibia; C – C. solieri, protibia; D–I – C. kaszabi sp. nov.: D – aedeagus; E – penis; F – gastral spicula; G – inner sternite VIII; H – protibia; I – mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.4504788" httpUri="https://zenodo.org/record/4504788/files/figure.png" pageId="5" pageNumber="336">8</figureCitation>
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D–I)
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</paragraph>
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</subSubSection>
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<subSubSection id="56BD1887FF97891972BCFEFFFE17FEBD" pageId="5" pageNumber="336" type="materials_examined">
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<paragraph id="1E184B0CFF97891972BCFEFFFD65FEEA" blockId="5.[151,759,262,334]" box="[151,706,262,281]" pageId="5" pageNumber="336">
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<emphasis id="2CD3971EFF97891972BCFEFFFEB7FEEA" bold="true" box="[151,272,262,281]" pageId="5" pageNumber="336">
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<typeStatus id="C11CF5AEFF97891972BCFEFFFF63FEEA" box="[151,196,262,281]" pageId="5" pageNumber="336">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="66B00B9CFF978919733EFEFFFEF4FEEA" box="[277,339,262,281]" name="Turkey" pageId="5" pageNumber="336">Turkey</collectingCountry>
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,
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<collectingRegion id="DC6385EEFF9789197370FEFFFE54FEEA" box="[347,499,262,281]" country="Turkey" name="Bayburt" pageId="5" pageNumber="336">Bayburt Province</collectingRegion>
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,
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<collectingRegion id="DC6385EEFF97891973D0FEFFFDE7FEEA" box="[507,576,262,281]" country="Turkey" name="Bayburt" pageId="5" pageNumber="336">Bayburt</collectingRegion>
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town,
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<quantity id="D95FE6E9FF9789197050FEFFFD19FEEA" box="[635,702,262,281]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="336" unit="m" value="1200.0">1200 m</quantity>
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.
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</paragraph>
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<paragraph id="1E184B0CFF97891972BCFED9FE17FEBD" blockId="5.[151,759,262,334]" pageId="5" pageNumber="336">
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||
<emphasis id="2CD3971EFF97891972BCFED9FEBEFEC0" bold="true" box="[151,281,288,307]" pageId="5" pageNumber="336">Type material.</emphasis>
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||
<materialsCitation id="AECF4151FF9789197336FED9FE17FEBD" ID-GBIF-Occurrence="3313296304" collectionCode="HNHM" collectorName="Osella" country="Turkey" location="Gumushane Bayburt" pageId="5" pageNumber="336" specimenCount="1" stateProvince="Gumushane Bayburt" typeStatus="holotype">
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<typeStatus id="C11CF5AEFF9789197336FED9FEC8FEC2" box="[285,367,288,307]" pageId="5" pageNumber="336" type="holotype">HOLOTYPE</typeStatus>
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: ♁, ‘
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<collectingCountry id="66B00B9CFF97891973BEFED9FE7DFEC0" box="[405,474,288,307]" name="Turkey" pageId="5" pageNumber="336">Turchia</collectingCountry>
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- vil.
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<collectingRegion id="DC6385EEFF9789197023FED9FD12FEC0" box="[520,693,288,307]" isEnumeration="true" pageId="5" pageNumber="336">
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<collectingRegion id="DC6385EEFF9789197023FED9FDCAFEC0" box="[520,621,288,307]" country="Turkey" name="Gumushane" pageId="5" pageNumber="336">Gümüshane</collectingRegion>
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<collectingRegion id="DC6385EEFF978919705AFED9FD12FEC0" box="[625,693,288,307]" country="Turkey" name="Bayburt" pageId="5" pageNumber="336">Bayburt</collectingRegion>
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</collectingRegion>
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m 1200 / 1.vii.75 /
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<collectorName id="B3522EDAFF97891972DFFEC2FE8CFEBD" box="[244,299,315,334]" pageId="5" pageNumber="336">Osella</collectorName>
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leg.’ (
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<collectionCode id="78B6D3C9FF978919734BFEC2FE01FEBD" box="[352,422,315,334]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="336">HNHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="56BD1887FF97891972BCFE9FFB7FFEE6" pageId="5" pageNumber="336" type="description">
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<paragraph id="1E184B0CFF97891972BCFE9FFD2AFC6E" blockId="5.[151,761,357,2077]" pageId="5" pageNumber="336">
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<emphasis id="2CD3971EFF97891972BCFE9FFECDFE8E" bold="true" box="[151,362,357,381]" pageId="5" pageNumber="336">
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Description.
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<emphasis id="2CD3971EFF9789197307FE9CFECDFE8E" bold="true" box="[300,362,357,381]" italics="true" pageId="5" pageNumber="336">Male.</emphasis>
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</emphasis>
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Body black, shiny, robust. Apex of frontoclypeus (frontoclypeal tooth) obtuse. Anterior margin of head not separated from the other surface. Lateral margins of head weakly sinuated between genae and frontoclypeus. Lateral margins of genae regularly rounded, not sinuated. Supraocular keels extended almost from level of basal margin of eye. Ratio of width of head at eyes to distance between eyes 1.3. Antennae short, with apical antennomere reaching basal third of pronotum. Ratio of length (width) of antennomeres II–XI: 0.4(0.5): 0.7(0.5): 1.0(0.6): 0.6(0.5): 0.6(0.5): 0.6(0.5): 0.5(0.5): 0.5(0.5): 0.5(0.5): 0.4(0.4). Punctation of head moderately coarse and dense, puncture diameter 1.5× as wide as distance among punctures. Vertex with deep transverse depression, shiny, with coarse and dense rasp-shaped punctation (puncture diameter 1.5–2.0× as wide as distance among punctures). Mentum weakly shiny, with smoothed punctation of weakly rasp-shaped punctures and wrinkles.
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</paragraph>
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<paragraph id="1E184B0CFF978919729EFC5FFD8AFA2E" blockId="5.[151,761,357,2077]" pageId="5" pageNumber="336">Pronotum transverse (1.3× as wide as long), widest in middle, 1.6× as wide as head. Ratio of pronotal width at anterior margin, in middle and at base 4.5: 6.5: 5.5. Anterior margin of pronotum straight, base weakly bisinuate, weakly rounded in middle. Lateral margins of pronotum moderately rounded, straight near base. Anterior angles of pronotum straight, rounded, posterior angles weakly obtuse, narrowly rounded at apex. All margins of pronotum beaded, only anterior margin with interrupted bead in middle. Disc of pronotum moderately regularly convex. Punctation of disc regularly moderately coarse and dense (puncture diameter from subequal to 1.5–2.0× as wide as distance among punctures). Prothoracic hypomera with inner two thirds of surface punctate by rasp-shaped punctures. Prosternum with coarse and dense punctation. Prosternal process moderately convex, not strongly projected, not beaded, with longitudinal impression in middle, with fine and moderately dense punctation.</paragraph>
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<paragraph id="1E184B0CFF978919729EFA1FFD37F8AE" blockId="5.[151,761,357,2077]" pageId="5" pageNumber="336">Elytra oval (1.3× as long as wide), strongly convex, distinctly depressed along suture in anterior quarter, 2.0 × longer and 1.27× wider than pronotum, 2.0 × wider than head. Base of elytra 1.06× wider than base of pronotum. Elytra without microwrinkles, with distinct fine and sparse punctation (puncture diameter 3.0× as wide as distance among punctures). Epipleura wrinkled. Mesoventrite with multiple short regularly rounded bracket-shaped foveae connected by apices. Metaventrite with fine, moderately dense punctation along meso- and metacoxae, remaining surface of metaventrite with smooth sparse punctation. Metepisterna with smoothed coarse punctation.</paragraph>
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<paragraph id="1E184B0CFF978919729EF89FFEF4F82E" blockId="5.[151,761,357,2077]" pageId="5" pageNumber="336">First and second abdominal ventrite completely beaded, third ventrite beaded on less than half its surface, the other ventrites not beaded. All ventrites with fine, moderately dense punctation.</paragraph>
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<paragraph id="1E184B0CFF978919729EF81FFC52FF06" blockId="5.[151,761,357,2077]" lastBlockId="5.[828,1437,190,661]" pageId="5" pageNumber="336">Legs short. Protibia not fossorial, 3.24× as long as wide. All tibiae straight. Ratio of femur/tibia/tarsus: 4.5: 3.0: 2.0 in fore legs, 4.5: 3.5: 3.0 in middle legs, and 5.0: 4.5: 3.0 in hind legs.</paragraph>
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<paragraph id="1E184B0CFF9789197171FF07FB7FFEE6" blockId="5.[828,1437,190,661]" box="[858,1240,254,277]" pageId="5" pageNumber="336">
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Body length
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<quantity id="D95FE6E9FF97891971CFFF07FB96FEE6" box="[996,1073,254,277]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="336" unit="mm" value="10.0">10 mm</quantity>
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, width 4.0 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="56BD1887FF9789197117FEE7FB05FD86" pageId="5" pageNumber="336" type="etymology">
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||
<paragraph id="1E184B0CFF9789197117FEE7FABEFEA6" blockId="5.[828,1437,190,661]" pageId="5" pageNumber="336">
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<emphasis id="2CD3971EFF9789197117FEE7FC1BFEC6" bold="true" box="[828,956,286,309]" pageId="5" pageNumber="336">Etymology.</emphasis>
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The species is named in honor of the famous specialist on darkling beetles Zoltan Kaszab.
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</paragraph>
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<paragraph id="1E184B0CFF9789197117FEA7FB05FD86" blockId="5.[828,1437,190,661]" pageId="5" pageNumber="336">
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<emphasis id="2CD3971EFF9789197117FEA7FBE0FE86" bold="true" box="[828,1095,350,373]" pageId="5" pageNumber="336">Comparative diagnosis.</emphasis>
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The new species is most similar to
|
||
<taxonomicName id="D9A7308FFF9789197170FE87FC46FE66" box="[859,993,382,405]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF9789197170FE87FC46FE66" box="[859,993,382,405]" italics="true" pageId="5" pageNumber="336">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
, which also has short roundebracket- shaped foveae on mesoventrite and elytra depressed in basal part along suture, but clearly differs in shiny body (
|
||
<taxonomicName id="D9A7308FFF978919716EFE27FC6DFE06" box="[837,970,478,501]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF978919716EFE27FC6DFE06" box="[837,970,478,501]" italics="true" pageId="5" pageNumber="336">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
has dull body), regular, coarse and dense punctation of pronotum (
|
||
<taxonomicName id="D9A7308FFF978919766BFE07FB18FDE6" box="[1088,1215,510,533]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF978919766BFE07FB18FDE6" box="[1088,1215,510,533]" italics="true" pageId="5" pageNumber="336">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
has coarse and dense punctation only on sides of pronotum), moderately dense distinct elytral punctation (
|
||
<taxonomicName id="D9A7308FFF978919764EFDC7FB4DFDA6" box="[1125,1258,574,597]" class="Insecta" family="Tenebrionidae" genus="Calyptopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="2CD3971EFF978919764EFDC7FB4DFDA6" box="[1125,1258,574,597]" italics="true" pageId="5" pageNumber="336">C. pulchella</emphasis>
|
||
</taxonomicName>
|
||
has very sparse poorly visible elytral punctation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="56BD1887FF9789197117FD87FAEBFD66" box="[828,1356,638,661]" pageId="5" pageNumber="336" type="distribution">
|
||
<paragraph id="1E184B0CFF9789197117FD87FAEBFD66" blockId="5.[828,1437,190,661]" box="[828,1356,638,661]" pageId="5" pageNumber="336">
|
||
<emphasis id="2CD3971EFF9789197117FD87FC69FD66" bold="true" box="[828,974,638,661]" pageId="5" pageNumber="336">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="C11CF5AEFF97891976EAFD87FB48FD66" box="[1217,1263,638,661]" pageId="5" pageNumber="336">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |