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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1102.84953" ID-Pensoft-Pub="1313-2970-1102-1" ID-Pensoft-UUID="9856BA5AE24B5C68B128B57DC644E3F0" ID-ZooBank="2398E5CC94134AE5976BDEF9F138A814" ModsDocID="1313-2970-1102-1" checkinTime="1653045140087" checkinUser="pensoft" docAuthor="Liao, Hauchuan, Terayama, Mamoru &amp; Eguchi, Katsuyuki" docDate="2022" docId="960BB0F4DC9A57DEA6A85910EC3582B1" docLanguage="en" docName="ZooKeys 1102: 1-42" docOrigin="ZooKeys 1102" docPubDate="2022-05-19" docSource="http://dx.doi.org/10.3897/zookeys.1102.84953" docTitle="Pristepyris zhejiangensis" docType="treatment" docVersion="1" id="9856BA5AE24B5C68B128B57DC644E3F0" lastPageNumber="1" masterDocId="9856BA5AE24B5C68B128B57DC644E3F0" masterDocTitle="Revision of Taiwanese and Ryukyuan species of Pristepyris Kieffer, 1905, with description of a new species (Hymenoptera, Bethylidae)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="1" updateTime="1653045140087" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Taiwanese and Ryukyuan species of Pristepyris Kieffer, 1905, with description of a new species (Hymenoptera, Bethylidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liao, Hauchuan</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-Osawa 1 - 1, Hachioji, Tokyo 192 - 0397, Japan</mods:affiliation>
<mods:nameIdentifier type="email">hachiliao0808@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Terayama, Mamoru</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-Osawa 1 - 1, Hachioji, Tokyo 192 - 0397, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eguchi, Katsuyuki</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-Osawa 1 - 1, Hachioji, Tokyo 192 - 0397, Japan &amp; Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, Sakamoto 1 - 12 - 4, Nagasaki, Nagasaki 852 - 8523, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1102</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>42</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1102.84953</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1102.84953</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1102-1</mods:identifier>
<mods:identifier type="ZooBank">2398E5CC94134AE5976BDEF9F138A814</mods:identifier>
<mods:identifier type="Pensoft-UUID">9856BA5AE24B5C68B128B57DC644E3F0</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1" httpUri="http://treatment.plazi.org/id/960BB0F4DC9A57DEA6A85910EC3582B1" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="960BB0F4-DC9A-57DE-A6A8-5910EC3582B1" authority="(Terayama, Xu &amp; He, 2002)" baseAuthorityName="Terayama, Xu &amp; He" baseAuthorityYear="2002" class="Insecta" family="Bethylidae" genus="Pristepyris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pristepyris zhejiangensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zhejiangensis">Pristepyris zhejiangensis (Terayama, Xu &amp; He, 2002)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pristepyris zhejiangensis, male A-C, F, G TNT 180504 _ 01 D, E JM 190717 _ 33 A head in full-face view B antenna (left) C mandible D mesosoma in lateral view; arrow indicating an angulate corner present on cervical pronotal area E mesosoma in dorsal view; arrow indicating transverse pronotal carina present F forewing G hindwing. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1102.84953.figure11" httpUri="https://binary.pensoft.net/fig/686039" pageId="0" pageNumber="1">Figs 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pristepyris zhejiangensis, male genitalia, TNT 180504 _ 01 A hypopygium B, C genitalia in ventral view D apical lobe in outer-lateral view. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1102.84953.figure12" httpUri="https://binary.pensoft.net/fig/686040" pageId="0" pageNumber="1">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pristepyris zhejiangensis, female, TN 170427 _ 01 A head in full-face view B antenna (left) C mandible D mesosoma in lateral view E mesosoma in dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1102.84953.figure13" httpUri="https://binary.pensoft.net/fig/686041" pageId="0" pageNumber="1">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Terayama, Xu &amp; He" authorityYear="2002" class="Insecta" family="Bethylidae" genus="Acrepyris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acrepyris zhejiangensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zhejiangensis">Acrepyris zhejiangensis</taxonomicName>
<bibRefCitation author="Terayama, M" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="0" pageNumber="1" pagination="81 - 86" refId="B35" refString="Terayama, M, Xu, ZF, He, JH, 2002. Three new species of the genus Acrepyris Kieffer, 1905 (Hymenoptera, Bethylidae) from China. Japanese Journal of Systematic Entomology 8: 81 - 86" title="Three new species of the genus Acrepyris Kieffer, 1905 (Hymenoptera, Bethylidae) from China." volume="8" year="2002">Terayama et al. 2002</bibRefCitation>
: 83, figs 9-16. Holotype, type loc.: Deqing, Zhejiang, China.
<taxonomicName baseAuthorityName="Terayama, Xu &amp; He" baseAuthorityYear="2002" class="Insecta" family="Bethylidae" genus="Pristepyris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pristepyris zhejiangensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zhejiangensis">Pristepyris zhejiangensis</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4489.1.1" author="Azevedo, CO" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 294" refId="B8" refString="Azevedo, CO, Alencar, IDCC, Ramos, MS, Barbosa, DN, Colombo, WD, Vargas, RJM, Lim, J, 2018. Global guide of the flat wasps (Hymenoptera, Bethylidae). Zootaxa 4889 (1): 1 - 294, DOI: https://doi.org/10.11646/zootaxa.4489.1.1" title="Global guide of the flat wasps (Hymenoptera, Bethylidae)." url="https://doi.org/10.11646/zootaxa.4489.1.1" volume="4889" year="2018">Azevedo et al. 2018</bibRefCitation>
: 104 (genus transfer).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="male diagnosis">
<paragraph pageId="0" pageNumber="1">Male diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
TL ≈ 6.1-9.3 mm. HL/HW
<normalizedToken originalValue="×">x</normalizedToken>
100 = 88-103. Frons and vertex with deep foveolae (ca. 0.05-0.06 mm in diameter), of which intervals are smooth and shining and narrower than diameter of foveolate. Anterior clypeal margin nearly straight medially. Mandible with five apical teeth. Transverse pronotal carina present. Cervical pronotal area in lateral view round. LP/WP = 0.97-1.04. Metapostnotal median carina incomplete posteriorly. Tergum II with longitudinal sulcus and ridge, sternum II with longitudinal median carina. Apical margin of hypopygium straight medially. Apical lobe of aedeagus in lateral view elongate and spatulate, with broadened and rounded apex, in ventral view somewhat winding.
</paragraph>
<caption doi="10.3897/zookeys.1102.84953.figure11" httpUri="https://binary.pensoft.net/fig/686039" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
<taxonomicName baseAuthorityName="Terayama, Xu &amp; He" baseAuthorityYear="2002" class="Insecta" family="Bethylidae" genus="Pristepyris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pristepyris zhejiangensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pristepyris zhejiangensis</emphasis>
</taxonomicName>
, male
<emphasis bold="true" pageId="0" pageNumber="1">A-C, F, G</emphasis>
TNT180504_01
<emphasis bold="true" pageId="0" pageNumber="1">D, E</emphasis>
JM190717_33
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna (left)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
mandible
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
mesosoma in lateral view; arrow indicating an angulate corner present on cervical pronotal area
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
mesosoma in dorsal view; arrow indicating transverse pronotal carina present
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
forewing
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
hindwing. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="female diagnosis">
<paragraph pageId="0" pageNumber="1">Female diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
TL ≈ 6.5 mm. HL/
<normalizedToken originalValue="HW×">HWx</normalizedToken>
100 = 118-126. Frons and vertex with deep foveolae (ca. 0.03-0.04 mm in diameter), of which intervals are imbricate; intervals in vertex wider than diameter of foveolae; intervals in lateral and submedian part of frons as narrow as or narrower than diameter of foveolae; the area along mesal line without foveolae. Median portion of clypeus roundly and relatively strongly produced anteriad; apical clypeal margin deeply incurved medially. Compound eye less developed. Mandible with four teeth. Transverse pronotal carina absent. Dorsal face of pronotum, mesoscutellum, mesopleuron and dorsal and lateral faces of metapectal-propodeal complex imbricate. Mesosoma excluding dorsal and lateral faces of metapectal-propodeal complex with dense foveolae. Tarsal claws with thin and curved tooth. Tergum II with weak longitudinal ridge, without longitudinal sulcus. Sternum II without longitudinal median carina.
</paragraph>
<caption doi="10.3897/zookeys.1102.84953.figure12" httpUri="https://binary.pensoft.net/fig/686040" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<taxonomicName baseAuthorityName="Terayama, Xu &amp; He" baseAuthorityYear="2002" class="Insecta" family="Bethylidae" genus="Pristepyris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pristepyris zhejiangensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pristepyris zhejiangensis</emphasis>
</taxonomicName>
, male genitalia, TNT180504_01
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
hypopygium
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
genitalia in ventral view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
apical lobe in outer-lateral view. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Male redescription.</paragraph>
<paragraph pageId="0" pageNumber="1">
Full description was given by
<bibRefCitation author="Terayama, M" journalOrPublisher="Japanese Journal of Systematic Entomology" pageId="0" pageNumber="1" pagination="81 - 86" refId="B35" refString="Terayama, M, Xu, ZF, He, JH, 2002. Three new species of the genus Acrepyris Kieffer, 1905 (Hymenoptera, Bethylidae) from China. Japanese Journal of Systematic Entomology 8: 81 - 86" title="Three new species of the genus Acrepyris Kieffer, 1905 (Hymenoptera, Bethylidae) from China." volume="8" year="2002">Terayama et al. (2002)</bibRefCitation>
. Additional information as below.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
HL/
<normalizedToken originalValue="HW×">HWx</normalizedToken>
100 = 88-103 (88 in holotype). Frons and vertex with deep foveolae (ca. 0.05-0.06 mm in diameter), of which intervals are smooth and shining and narrower than diameter of foveolate. Occipital carina present. Median portion of clypeus roundly produced anteriad; median clypeal carina moderately distinct, almost reaching anterior margin; anterior clypeal margin nearly straight medially. Compound eye large and convex, with sparse thin erect setae. Mandible with five teeth.
</paragraph>
<caption doi="10.3897/zookeys.1102.84953.figure13" httpUri="https://binary.pensoft.net/fig/686041" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<taxonomicName baseAuthorityName="Terayama, Xu &amp; He" baseAuthorityYear="2002" class="Insecta" family="Bethylidae" genus="Pristepyris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pristepyris zhejiangensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis italics="true" pageId="0" pageNumber="1">Pristepyris zhejiangensis</emphasis>
</taxonomicName>
, female, TN170427_01
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
antenna (left)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
mandible
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
mesosoma in lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
mesosoma in dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
Dorsal area of pronotum smooth and shining, with deep foveolae, with distinct transverse pronotal carinae (arrow in Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pristepyris mieae, female, holotype A head in full-face view B mandible C mesosoma in dorsal view D mesosoma in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1102.84953.figure8" httpUri="https://binary.pensoft.net/fig/686036" pageId="0" pageNumber="1">8E</figureCitation>
). Cervical pronotal area in lateral view round. Mesopleuron elongate; anterior, upper and lower fovea distinct; acropleural area smooth and shining. Mesopleural pit absent. Mesodiscrimen concave, with weak median carina. Metasternum with metafurcal pit. Lateral face of metapectal-propodeal complex smooth and shining anteriorly, irregularly rugose on posterior half of central area. Metapectal-propodeal complex in dorsal view with LP/WP = 0.97-1.04, with lateral margins subparallel and slightly convex; metapostnotal median carina distinct, but incomplete posteriorly; submedian rugae irregularly running; sublateral margin distinct, incomplete posteriorly; posterior transverse margin distinct; dorsomedian and dorsolateral faces weakly rugoso-scabrous; median portion of propodeal declivity transversely rugoso-scabrous. Forewing with r-m2 flexion line (arrows in Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Pristepyris mieae, female, holotype A head in full-face view B mandible C mesosoma in dorsal view D mesosoma in lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1102.84953.figure8" httpUri="https://binary.pensoft.net/fig/686036" pageId="0" pageNumber="1">8F</figureCitation>
), without R2 and 2M12 flexion line. Hindwing with five distal hamuli. Tarsal claws bifid, with thin and curved apical teeth.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Metasoma</emphasis>
.
</emphasis>
Tergum II with longitudinal sulcus and ridge; sternum II with longitudinal median carina. Hypopygium with spiculum much longer than S9ala (spiculum broken in Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pristepyris ryukyuensis, male, holotype A head in full-face view B mandible C mesosoma in lateral view; arrow indicating an angulate corner present on cervical pronotal area D mesosoma in dorsal view; arrow indicating transverse pronotal carina present. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1102.84953.figure9" httpUri="https://binary.pensoft.net/fig/686037" pageId="0" pageNumber="1">9A</figureCitation>
); apical margin straight medially; ventral face of apicomedian part with relatively dense setae. Gonostipes glabrous, unfused to harpe. Harpe in ventral view widely elongated, slightly curved inward, with blunt apex, entirely covered with setae which increase in length toward apex; median basal portion with concavity which accommodates digitus and cuspis. Cuspis lobate and extending laterad, curled, with short, thick, conical setae at apex; subbasal part facing digitus with short and thin setea. Digitus extending laterad, curled; lateral face with short, thick, conical setae near apex. Apical lobe of aedeagus in lateral view elongate and spatulate, with broadened and rounded apex, in ventral view somewhat winding.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Female description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female of this species was newly-recognised by molecular phylogenetic analyses in the present study.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Body mostly dark brown; mandible, antenna, anterior flange of pronotum and legs brown or light brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Head capsule with posterior margin slightly incurved, with posterolateral corner round; HL/HW
<normalizedToken originalValue="×">x</normalizedToken>
100 = 118-126. Occipital carina present. Frons and vertex with deep foveolae (ca. 0.03-0.04 mm in diameter), of which intervals are imbricate; intervals in vertex wider than diameter of foveolae; intervals in lateral and submedian part of frons as narrow as or narrower than diameter of foveolae; the area along mesal line without foveolae. Median portion of clypeus roundly and relatively strongly produced anteriad, imbricate; median longitudinal carina not reaching anterior clypeal margin; anterior clypeal margin deeply incurved medially. Compound eye less developed. Mandible with four teeth; basalmost tooth relatively shorter than other ones. Antennomere I (excluding basal condylar bulb) 2.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width; antennomere I:II:III = 5:1:1 in length; antennomere II 0.9
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, narrowed and bent in basal part; antennomere III-XII each 0.76-0.85
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, elongate-cylindrical; antennomere XIII (terminal) 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, with round apex. Tarsal claws with thin and curved tooth.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
Pronotum with anterior flange extending anteriad beyond anterior margin of propleuron; transverse pronotal carina absent; cervical pronotal area in lateral view round, with a steep anterior face; dorsal area subtrapezoidal, with almost straight posterior margin, with deep foveolae of which intervals are wider than diameter of foveolae and weakly imbricate; LPD/WPD = 1.00-1.07. Mesoscutum overlain by posteromedian portion of pronotum. Mesoscutellum trapezoidal, 0.63-0.64
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, weakly imbricate, with sparse and deep foveolae. Mesopleuron largely imbricate excluding smooth anterodorsal part, with sparse and deep foveolae; anterior, upper and lower fovea absent; mesopleural pit absent. Mesodiscrimen with weak median carina. Metasternum with metafurcal pit. Lateral face of metapectal-propodeal complex imbricates entirely. Metapectal-propodeal complex in dorsal view weakly constricted behind propodeal spiracles and then widened again posteriad, without any distinct carinae which subdivide dorsal face; LP/WP = 2.28-2.42; dorsomedian face weakly imbricate; median portion of propodeal declivity weakly and transversely rugoso-scabrous, with sparse foveolae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Metasoma</emphasis>
.
</emphasis>
Tergum II with weak longitudinal ridge, without longitudinal sulcus; sternum II without longitudinal median carina.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="JM190717" collectingDate="2019-07-23" collectorName="HauChuan Liao" country="Japan" elevation="93" latitude="24.815832" location="Makiyama Park" longLatPrecision="21" longitude="125.21667" municipality="Irabu" specimenCount="15" specimenCount-male="15">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Japan">Japan</collectingCountry>
:
<collectingMunicipality>Irabu</collectingMunicipality>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="1">jima.</emphasis>
<specimenCount type="male">15 males</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/JM190717">JM190717</accessionNumber>
_31-45);
<location LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1:5655F0B14BB11065B37EDB658F118786" country="Japan" latitude="24.815832" longLatPrecision="21" longitude="125.21667" municipality="Irabu" name="Makiyama Park">Makiyama Park</location>
,
<geoCoordinate degrees="24" direction="north" minutes="48" orientation="latitude" precision="15" seconds="57" value="24.815832">24°48'57&quot;N</geoCoordinate>
,
<geoCoordinate degrees="125" direction="east" minutes="13" orientation="longitude" precision="15" seconds="00" value="125.21667">125°13'00&quot;E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.3" unit="m" value="93.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.3" unit="m" value="93.0">93 m</elevation>
</quantity>
alt.;
<collectorName>HauChuan Liao</collectorName>
leg. (sweeping);
<collectingDate value="2019-07-23">23/VII/2019</collectingDate>
</materialsCitation>
.
<materialsCitation accessionNumber="JM190717" collectingDate="2019-07-23" collectorName="HauChuan Liao" country="Japan" elevation="93" latitude="24.815832" location="Makiyama Park" longLatPrecision="21" longitude="125.21667" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/JM190717">JM190717</accessionNumber>
_28);
<location LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1:24152A96C946F14103BEA5666F59175F" country="Japan" latitude="24.815832" longLatPrecision="21" longitude="125.21667" name="Makiyama Park">Makiyama Park</location>
,
<geoCoordinate degrees="24" direction="north" minutes="48" orientation="latitude" precision="15" seconds="57" value="24.815832">24°48'57&quot;N</geoCoordinate>
,
<geoCoordinate degrees="125" direction="east" minutes="13" orientation="longitude" precision="15" seconds="00" value="125.21667">125°13'00&quot;E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.3" unit="m" value="93.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.3" unit="m" value="93.0">93 m</elevation>
</quantity>
alt.;
<collectorName>HauChuan Liao</collectorName>
leg. (sweeping);
<collectingDate value="2019-07-23">23/VII/2019</collectingDate>
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="HauChuan Liao" country="Japan" elevation="33" latitude="24.396667" location="Tropical Biosphere Research Center" longLatPrecision="21" longitude="123.803055" municipality="Iriomote" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingMunicipality>Iriomote</collectingMunicipality>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="1">jima</emphasis>
<specimenCount type="male">1 male</specimenCount>
(JIR190717_47);
<location LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1:CCA7979111838F02F96A63E69EE53F2D" country="Japan" latitude="24.396667" longLatPrecision="21" longitude="123.803055" municipality="Iriomote" name="Tropical Biosphere Research Center">Tropical Biosphere Research Center</location>
,
<geoCoordinate degrees="24" direction="north" minutes="23" orientation="latitude" precision="15" seconds="48" value="24.396667">24°23'48&quot;N</geoCoordinate>
,
<geoCoordinate degrees="123" direction="east" minutes="48" orientation="longitude" precision="15" seconds="11" value="123.803055">123°48'11&quot;E</geoCoordinate>
,
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.3" unit="m" value="33.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.3" unit="m" value="33.0">33 m</elevation>
</quantity>
alt.
<collectorName>HauChuan Liao</collectorName>
leg. (sweeping)
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Hauchuan Liao" country="Taiwan" elevation="548" latitude="24.8525" location="Mt. Dadao Wurai" longLatPrecision="21" longitude="121.5575" specimenCount="2" specimenCount-male="2" stateProvince="New Taipei City">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
: N.
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
.
</emphasis>
<specimenCount type="male">2 males</specimenCount>
(TNT171019_04, TNT180504_01);
<location LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1:962F4078EF78DD490E914B60589BD5D0" country="Taiwan" latitude="24.8525" longLatPrecision="21" longitude="121.5575" name="Mt. Dadao Wurai" stateProvince="New Taipei City">Mt. Dadao Wurai</location>
,
<collectingRegion country="Taiwan" name="New Taipei">New Taipei City</collectingRegion>
,
<geoCoordinate degrees="24" direction="north" minutes="51" orientation="latitude" precision="15" seconds="09" value="24.8525">24°51'09&quot;N</geoCoordinate>
,
<geoCoordinate degrees="121" direction="east" minutes="33" orientation="longitude" precision="15" seconds="27" value="121.5575">121°33'27&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" unit="m" value="548.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" unit="m" value="548.0">548 m</elevation>
</quantity>
alt.;
<collectorName>Hauchuan Liao</collectorName>
leg. (sweeping);
<collectingDate value="2017-10-26">26/X/2017</collectingDate>
,
<collectingDate value="2018-05-04">4/V/2018</collectingDate>
</materialsCitation>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<taxonomicName genus="C." lsidName="C." pageId="0" pageNumber="1" rank="genus">C.</taxonomicName>
<materialsCitation accessionNumber="TN190315" collectingDate="2019-03-20" collectorName="HauChuan Liao" country="Taiwan" elevation="789" latitude="23.868334" location="Sungpolun Trail" longLatPrecision="21" longitude="120.92889" specimenCount="1" specimenCount-male="1" stateProvince="Nantou County">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
.
<specimenCount type="male">1 male</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TN190315">TN190315</accessionNumber>
_24);
<location LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1:90E8131D789D3DF4A794455C7EF475E2" country="Taiwan" latitude="23.868334" longLatPrecision="21" longitude="120.92889" name="Sungpolun Trail" stateProvince="Nantou County">Sungpolun Trail</location>
,
<collectingRegion country="Taiwan" name="Nantou">Nantou County</collectingRegion>
,
<geoCoordinate degrees="23" direction="north" minutes="52" orientation="latitude" precision="15" seconds="06" value="23.868334">23°52'06&quot;N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="55" orientation="longitude" precision="15" seconds="44" value="120.92889">120°55'44&quot;E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.89" unit="m" value="789.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.89" unit="m" value="789.0">789 m</elevation>
</quantity>
alt.;
<collectorName>HauChuan Liao</collectorName>
leg. (sweeping);
<collectingDate value="2019-03-20">20/III/2019</collectingDate>
</materialsCitation>
</emphasis>
.
<materialsCitation accessionNumber="TN170427" collectingDate="2017-04-27" collectorName="Po-Cheng Hsu" country="Taiwan" location="Huisun Experimental Forest" specimenCount="1" specimenCount-female="1" stateProvince="Nantou County">
<specimenCount type="female">1 female</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TN170427">TN170427</accessionNumber>
_01);
<location LSID="urn:lsid:plazi:treatment:960BB0F4DC9A57DEA6A85910EC3582B1:2E9F2969F6A7F81D23A36007D02C9E7B" country="Taiwan" name="Huisun Experimental Forest" stateProvince="Nantou County">Huisun Experimental Forest</location>
,
<collectingRegion country="Taiwan" name="Nantou">Nantou County</collectingRegion>
.
<collectorName>Po-Cheng Hsu</collectorName>
leg.;
<collectingDate value="2017-04-27">27/IV/2017</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="taxonomic remarks">
<paragraph pageId="0" pageNumber="1">Taxonomic remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is most similar in general appearance to
<taxonomicName genus="P." lsidName="P. ryukyuensis" pageId="0" pageNumber="1" rank="species" species="ryukyuensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. ryukyuensis</emphasis>
</taxonomicName>
among the named species known from East and Southeast Asia (for details, see under Taxonomic remarks of &quot;
<taxonomicName genus="P." lsidName="P. ryukyuensis" pageId="0" pageNumber="1" rank="species" species="ryukyuensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. ryukyuensis</emphasis>
</taxonomicName>
&quot;).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">Eastern China (Zhejiang), southern Ryukyu, northern and central Taiwan (new to Taiwan); evergreen broadleaf forests.</paragraph>
</subSubSection>
</treatment>
</document>
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