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<document id="F7AC0E84CB3C9407901A7AD0A0E2D1DB" ID-DOI="10.11646/zootaxa.5486.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13209709" ID-ZooBank="E696DDCA-FEAE-4EC6-9FF4-0CC3E5F55063" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1722787464916" checkinUser="plazi" docAuthor="Luu, Vinh Quang, Hoang, Tuoi Thi, Ha, Hong Bich, Grismer, Jesse L., Murdoch, Matthew, Sitthivong, Saly, Phimpasone, Vilay &amp; Grismer, L. Lee" docDate="2024" docId="867887EBFFB2FFB4FF13FF4FF0C4FEBC" docLanguage="en" docName="zootaxa.5486.1.3.pdf" docOrigin="Zootaxa 5486 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5486.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemiphyllodactylus vanhoensis Luu, Hoang, Ha, Grismer, Murdoch, Sitthivong, Phimpasone &amp; Grismer, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="104" masterDocId="7A41FF93FFADFF96FF84FFD8F353FFD9" masterDocTitle="Integrative taxonomy reveals two new species of karst-dwelling Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from the border region of Laos and Vietnam" masterLastPageNumber="108" masterPageNumber="71" pageNumber="102" updateTime="1722950403596" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4C362AF1F2F26452EA7F1A374B4D23D4">Integrative taxonomy reveals two new species of karst-dwelling Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from the border region of Laos and Vietnam</mods:title>
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<mods:namePart id="2D30D303DEF20A2B1624D2E442CD9C5D">Luu, Vinh Quang</mods:namePart>
<mods:nameIdentifier id="F3BCB5737611D0A60C1E23E2CBE31D69" type="ORCID">0000-0002-0634-1338</mods:nameIdentifier>
<mods:affiliation id="76DE6CAB99FD1806AC74E2D933EFB89A">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="639FBFAC52905DBC991EC8DBA7C6A7F2" type="email">vinhlq@vnuf.edu.vn</mods:nameIdentifier>
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<mods:namePart id="44371A9E7A71078EB997B262A558F57F">Hoang, Tuoi Thi</mods:namePart>
<mods:nameIdentifier id="D06F4E2520CC5F708EAF80513C733E81" type="ORCID">0000-0002-3564-0601</mods:nameIdentifier>
<mods:affiliation id="DB758A5A3C3BBFE35B5448B306978BFD">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="197972A441C5E5A88FE3AD66F3DF6E4C" type="email">tuoiht@vnuf.edu.vn</mods:nameIdentifier>
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<mods:namePart id="11F8D591304CE9EB376478756784E74A">Ha, Hong Bich</mods:namePart>
<mods:nameIdentifier id="6C4E2BCA050A4AC8112B027CAF7F2A94" type="ORCID">0000-0003-4777-5463</mods:nameIdentifier>
<mods:affiliation id="D2A5DC87BDB2AD7AE7DE862C7844ACB3">College of Forestry Biotechnology, Vietnam National University of Forestry, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="354B3A674B56A0DFA96334E84B6CD882" type="email">honghb@vnuf.edu.vn</mods:nameIdentifier>
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<mods:namePart id="EFB388B085BDE280C23F2CEC8CC48E8E">Grismer, Jesse L.</mods:namePart>
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<mods:namePart id="0E1BF3053767B120DEBE3C0A250D8BDF">Murdoch, Matthew</mods:namePart>
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<mods:nameIdentifier id="6BC5814ECD7B15B40A2AB0B09A47E741" type="email">hewmurdoch@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C38E40D54851822CA60AB1E62E360D74">Sitthivong, Saly</mods:namePart>
<mods:nameIdentifier id="2289D8369D32D385DED507DEC5DF3726" type="ORCID">0000-0002-9993-2136</mods:nameIdentifier>
<mods:affiliation id="9E083F5371DE9F3F93497E58948B3F91">Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR</mods:affiliation>
<mods:nameIdentifier id="58F587A4626A845D442D9F5F32F27656" type="email">s.sitthivong@nuol.edu.la</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="6C3BCA9328CB373C1B253184A231C8F5">Phimpasone, Vilay</mods:namePart>
<mods:nameIdentifier id="AFD63AB8E0C8906857868FD49A5EFC7C" type="ORCID">0009-0001-9744-0728</mods:nameIdentifier>
<mods:affiliation id="E890209450E679D8C0DE7D3BD153C3B8">Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR</mods:affiliation>
<mods:nameIdentifier id="46BA33F3426D672DF452C237F2D3D428" type="email">v.phimpasone@nuol.edu.la</mods:nameIdentifier>
</mods:name>
<mods:name id="3E7F2DEB60824D3281A89B6BC4E6D633" type="personal">
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<mods:namePart id="96872E0B3ED7C4264F795D21B50620CB">Grismer, L. Lee</mods:namePart>
<mods:nameIdentifier id="7A866734E692D790E9AE6F65E0B1FDD1" type="ORCID">0000-0001-8422-3698</mods:nameIdentifier>
<mods:affiliation id="A272E1430FDD7AB7FA2481F0AD910D29">Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA &amp; Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
<mods:nameIdentifier id="6A933AF8A9AED03F16C9889647E1961A" type="email">lgrismer@lasierra.edu</mods:nameIdentifier>
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<mods:typeOfResource id="C64468C9ABB0B9604D4754E851827D78">text</mods:typeOfResource>
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<mods:title id="AA9850E8334D6DDEC6D98B71998BE4C7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B9860C8D69D2F631F2FC7573F654FE4D">
<mods:date id="FAD69FED033159D693057C57E3C6DA6C">2024</mods:date>
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<mods:number id="EB805824BD76DF6E27E3E18A41E00DD4">2024-07-24</mods:number>
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<mods:number id="D70628E0C38FF4E5D9026D969ED6EC67">5486</mods:number>
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<treatment id="867887EBFFB2FFB4FF13FF4FF0C4FEBC" LSID="urn:lsid:plazi:treatment:867887EBFFB2FFB4FF13FF4FF0C4FEBC" httpUri="http://treatment.plazi.org/id/867887EBFFB2FFB4FF13FF4FF0C4FEBC" lastPageId="34" lastPageNumber="104" pageId="31" pageNumber="102">
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<heading id="55268191FFB2FF89FF13FF4FF12FFF68" bold="true" box="[151,636,151,178]" fontSize="11" level="1" pageId="31" pageNumber="102" reason="1">
<emphasis id="3CA5EAEFFFB2FF89FF13FF4FF12FFF68" bold="true" box="[151,636,151,178]" pageId="31" pageNumber="102">
<taxonomicName id="C9D14D7EFFB2FF89FF13FF4FF144FF6B" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[151,535,151,178]" class="Repitilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="102" phylum="Chordata" rank="species" species="vanhoensis" status="sp. nov.">
<emphasis id="3CA5EAEFFFB2FF89FF13FF4FF144FF6B" bold="true" box="[151,535,151,178]" italics="true" pageId="31" pageNumber="102">Hemiphyllodactylus vanhoensis</emphasis>
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<taxonomicNameLabel id="27965794FFB2FF89FD9AFF4FF12FFF68" box="[542,636,151,177]" pageId="31" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="0E6E36FDFFB2FF89FF13FF04F29EFF2D" blockId="31.[151,461,220,244]" box="[151,461,220,244]" pageId="31" pageNumber="102">
Vanho Slender Gecko (
<figureCitation id="96EA2A78FFB2FF89FE06FF04F293FF2D" box="[386,448,220,244]" captionStart="FIGURE 9" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[284,1303,182,1918]" captionTargetId="figure-36@32.[284,1304,181,1919]" captionTargetPageId="32" captionText="FIGURE 9. Dorsal views of Hemiphyllodactylus vanhoensis sp. nov. from Bun Village, Tan Xuan Commune, Van Ho District, Son La Province, Vietnam. A: adult female holotype (VNUF R.2022.07), B: adult female paratype (VNUF R. 2022.09), and C: juvenile male paratype (VNUF R. 2023.22). Photos: Vinh Quang Luu." figureDoi="http://doi.org/10.5281/zenodo.13209727" httpUri="https://zenodo.org/record/13209727/files/figure.png" pageId="31" pageNumber="102">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FFB2FF89FF13FEC3F17BFE30" pageId="31" pageNumber="102" type="materials_examined">
<paragraph id="0E6E36FDFFB2FF89FF13FEC3F7DDFEA4" blockId="31.[151,1437,283,2002]" pageId="31" pageNumber="102">
<materialsCitation id="BEB93CA0FFB2FF89FF13FEC3F7DDFEA4" collectingDate="2022-06-11" collectionCode="VNUF, R, V" collectorName="Adult &amp; Vinh Q. Luu" country="Vietnam" county="Van Ho District" elevation="1065" location="Nguyen" longitude="104.71333" municipality="Tan Xuan Commune" pageId="31" pageNumber="102" specimenCount="1" stateProvince="Son La" typeStatus="holotype">
<emphasis id="3CA5EAEFFFB2FF89FF13FEC3F258FEEC" bold="true" box="[151,267,283,309]" pageId="31" pageNumber="102">
<typeStatus id="D16A885FFFB2FF89FF13FEC3F255FEEC" box="[151,262,283,309]" pageId="31" pageNumber="102" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectorName id="A324532BFFB2FF89FE96FEC3F201FEEC" box="[274,338,283,309]" pageId="31" pageNumber="102">Adult</collectorName>
female
<collectionCode id="68C0AE38FFB2FF89FE2FFEC4F2A4FEEF" box="[427,503,284,310]" pageId="31" pageNumber="102">VNUF</collectionCode>
<collectionCode id="68C0AE38FFB2FF89FE7AFEC4F142FEEF" box="[510,529,284,310]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="102">R</collectionCode>
.2022.07 (Field no. TX2.22.07) collected on
<date id="7A6F103DFFB2FF89FB80FEC3F7CBFEEC" box="[1028,1176,283,310]" pageId="31" pageNumber="102" value="2022-06-11">
<collectingDate id="6A2BE9D5FFB2FF89FB80FEC3F7CBFEEC" box="[1028,1176,283,310]" pageId="31" pageNumber="102" value="2022-06-11">11 June 2022</collectingDate>
</date>
at 1940 hrs by
<collectorName id="A324532BFFB2FF89FAC0FEC3F394FE83" pageId="31" pageNumber="102">Vinh Q. Luu</collectorName>
, Nghia
<collectionCode id="68C0AE38FFB2FF89FE9FFE98F27CFE83" box="[283,303,320,346]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="102" type="Museum">V</collectionCode>
. Ha, Huong
<collectionCode id="68C0AE38FFB2FF89FE3CFE98F29FFE83" box="[440,460,320,346]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="102" type="Museum">V</collectionCode>
. Bui, Phuong
<collectionCode id="68C0AE38FFB2FF89FDE1FE98F12AFE83" box="[613,633,320,346]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="102" type="Museum">V</collectionCode>
.
<location id="0B0E6026FFB2FF89FD07FE98F18FFE83" LSID="urn:lsid:plazi:treatment:867887EBFFB2FFB4FF13FF4FF0C4FEBC:0B0E6026FFB2FF89FD07FE98F18FFE83" box="[643,732,320,346]" country="Vietnam" county="Van Ho District" longitude="104.71333" municipality="Tan Xuan Commune" name="Nguyen" pageId="31" pageNumber="102" stateProvince="Son La">Nguyen</location>
at the karst mountain of
<location id="0B0E6026FFB2FF89FC69FE98F726FE80" LSID="urn:lsid:plazi:treatment:867887EBFFB2FFB4FF13FF4FF0C4FEBC:0B0E6026FFB2FF89FC69FE98F726FE80" box="[1005,1141,319,346]" country="Vietnam" county="Van Ho District" longitude="104.71333" municipality="Tan Xuan Commune" name="Bun Village" pageId="31" pageNumber="102" stateProvince="Son La">Bun Village</location>
,
<collectingMunicipality id="EE0AAC87FFB2FF89FBFAFE98F63BFE80" box="[1150,1384,319,346]" pageId="31" pageNumber="102">Tan Xuan Commune</collectingMunicipality>
,
<collectingCounty id="E70F4E71FFB2FF89FAF5FE98F244FEA4" pageId="31" pageNumber="102">Van Ho District</collectingCounty>
,
<collectingRegion id="CC15F81FFFB2FF89FEA6FEBBF2B3FEA4" box="[290,480,355,382]" country="Vietnam" name="Son La" pageId="31" pageNumber="102">Son La Province</collectingRegion>
,
<collectingCountry id="76C6766DFFB2FF89FE6FFEBBF119FEA4" box="[491,586,355,381]" name="Vietnam" pageId="31" pageNumber="102">Vietnam</collectingCountry>
(20°40'352&quot;N 104°42'048&quot;E;
<quantity id="C9299B18FFB2FF89FC1AFEBBF0A0FEA7" box="[926,1011,355,382]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="31" pageNumber="102" unit="m" value="1065.0">
<elevation id="85FCD1CEFFB2FF89FC1AFEBBF0A0FEA7" box="[926,1011,355,382]" metricMagnitude="3" metricUnit="m" metricValue="1.065" pageId="31" pageNumber="102" unit="m" value="1065.0">1065 m</elevation>
</quantity>
in elevation).
</materialsCitation>
</paragraph>
<paragraph id="0E6E36FDFFB2FF89FF43FE5FF17BFE30" blockId="31.[151,1437,283,2002]" pageId="31" pageNumber="102">
<materialsCitation id="BEB93CA0FFB2FF89FF43FE5FF6CFFE78" box="[199,1436,391,418]" collectingDate="2022-06-11" collectionCode="VNUF, R" collectorName="Adult &amp; Vinh Q. Luu" country="Vietnam" county="Van Ho District" elevation="1065" location="Nguyen" longitude="104.71333" municipality="Tan Xuan Commune" pageId="31" pageNumber="102" specimenCount="1" stateProvince="Son La" typeStatus="paratype">
<emphasis id="3CA5EAEFFFB2FF89FF43FE5FF26CFE78" bold="true" box="[199,319,391,417]" pageId="31" pageNumber="102">
<typeStatus id="D16A885FFFB2FF89FF43FE5FF26CFE78" box="[199,319,391,417]" pageId="31" pageNumber="102" type="paratype">Paratypes</typeStatus>
</emphasis>
. Adult female
<collectionCode id="68C0AE38FFB2FF89FE64FE50F17FFE7B" box="[480,556,392,418]" pageId="31" pageNumber="102">VNUF</collectionCode>
<collectionCode id="68C0AE38FFB2FF89FDB5FE50F117FE7B" box="[561,580,392,418]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="102">R</collectionCode>
. 2022.09 (Field no. TX2.22.09) bears the same collection data as the holotype
</materialsCitation>
<materialsCitation id="BEB93CA0FFB2FF89FF13FE73F176FE30" collectingDate="2023-08-29" collectionCode="VNUF, R" collectorName="Vinh Q. Luu &amp; Le D. Phan" country="Vietnam" county="Van Ho District" elevation="1065" location="Nguyen" longitude="104.71333" municipality="Tan Xuan Commune" pageId="31" pageNumber="102" specimenCount="1" stateProvince="Son La" typeStatus="paratype">
and juvenile male
<collectionCode id="68C0AE38FFB2FF89FEE0FE74F2E3FE1F" box="[356,432,428,454]" pageId="31" pageNumber="102">VNUF</collectionCode>
<collectionCode id="68C0AE38FFB2FF89FE33FE74F299FE1F" box="[439,458,428,454]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="102">R</collectionCode>
.2023.22 (Field no. TX23.22) by
<collectorName id="A324532BFFB2FF89FCB9FE73F09FFE1F" box="[829,972,427,454]" pageId="31" pageNumber="102">Vinh Q. Luu</collectorName>
,
<collectorName id="A324532BFFB2FF89FC53FE74F705FE1C" box="[983,1110,427,454]" pageId="31" pageNumber="102">Le D. Phan</collectorName>
collected on
<date id="7A6F103DFFB2FF89FB6FFE73F6CFFE1C" box="[1259,1436,427,453]" pageId="31" pageNumber="102" value="2023-08-29">
<collectingDate id="6A2BE9D5FFB2FF89FB6FFE73F6CFFE1C" box="[1259,1436,427,453]" pageId="31" pageNumber="102" value="2023-08-29">29 August 2023</collectingDate>
</date>
at the same locality as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FFB2FF89FF43FE2BF0CEFC40" pageId="31" pageNumber="102" type="diagnosis">
<paragraph id="0E6E36FDFFB2FF89FF43FE2BF0CEFC40" blockId="31.[151,1437,283,2002]" pageId="31" pageNumber="102">
<emphasis id="3CA5EAEFFFB2FF89FF43FE2BF269FDD4" bold="true" box="[199,314,499,525]" pageId="31" pageNumber="102">Diagnosis</emphasis>
.
<taxonomicName id="C9D14D7EFFB2FF89FEC1FE2BF1F7FDD4" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[325,676,499,525]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="31" pageNumber="102" phylum="Chordata" rank="species" species="vanhoensis" status="sp. nov.">
<emphasis id="3CA5EAEFFFB2FF89FEC1FE2BF1F7FDD4" box="[325,676,499,525]" italics="true" pageId="31" pageNumber="102">Hemiphyllodactylus vanhoensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFFB2FF89FD2CFE2BF1ADFDD4" bold="true" box="[680,766,499,525]" pageId="31" pageNumber="102">
<taxonomicNameLabel id="27965794FFB2FF89FD2CFE2BF1ADFDD4" box="[680,766,499,525]" pageId="31" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from all other species of
<taxonomicName id="C9D14D7EFFB2FF89FB38FE2BF6CFFDD4" box="[1212,1436,499,525]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="31" pageNumber="102" phylum="Chordata" rank="genus">
<emphasis id="3CA5EAEFFFB2FF89FB38FE2BF6CFFDD4" box="[1212,1436,499,525]" italics="true" pageId="31" pageNumber="102">Hemiphyllodactylus</emphasis>
</taxonomicName>
by possessing the unique combination of having a maximum SVL of
<quantity id="C9299B18FFB2FF89FC67FDCFF703FDEB" box="[995,1104,535,562]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="31" pageNumber="102" unit="mm" value="46.5">46.5 mm</quantity>
; 7
<emphasis id="3CA5EAEFFFB2FF89FBF4FDC0F72DFDE8" box="[1136,1150,536,561]" italics="true" pageId="31" pageNumber="102">–</emphasis>
10 chin scales; enlarged postmentals; two or three circumnasal scales; two or three intersupranasals (=postrostrals); 10
<emphasis id="3CA5EAEFFFB2FF89FB5EFDE4F7BBFD8C" box="[1242,1256,572,597]" italics="true" pageId="31" pageNumber="102">–</emphasis>
12 supralabials; 10 or 11 infralabials; 15
<emphasis id="3CA5EAEFFFB2FF89FE2FFDB8F2EAFDA0" box="[427,441,608,633]" italics="true" pageId="31" pageNumber="102">–</emphasis>
19 longitudinally arranged dorsal scales at midbody contained within one eye diameter and 8
<emphasis id="3CA5EAEFFFB2FF89FF51FD5CF3B0FD44" box="[213,227,644,669]" italics="true" pageId="31" pageNumber="102">–</emphasis>
11 ventral scales; four to five subdigital lamellae on the first finger; four subdigital lamellae on the first toe; no pore
<emphasis id="3CA5EAEFFFB2FF89FF6AFD70F3A4FD18" box="[238,247,680,705]" italics="true" pageId="31" pageNumber="102">-</emphasis>
bearing, femoroprecloacal scales in adult females; 21 continuous, pore
<emphasis id="3CA5EAEFFFB2FF89FB9BFD70F77BFD18" box="[1055,1064,680,705]" italics="true" pageId="31" pageNumber="102">-</emphasis>
bearing, femoroprecloacal scales (pitted scales in the juvenile male); no plate
<emphasis id="3CA5EAEFFFB2FF89FDF9FD14F1D5FD3C" box="[637,646,716,741]" italics="true" pageId="31" pageNumber="102">-</emphasis>
like subcaudal scales; adult females not yellow; dark and light pre
<emphasis id="3CA5EAEFFFB2FF89FAE0FD14F63EFD3C" box="[1380,1389,716,741]" italics="true" pageId="31" pageNumber="102">-</emphasis>
and postorbital stripes in parallel extends from posterior nares to forelimb insertion; dorsolateral light
<emphasis id="3CA5EAEFFFB2FF89FB7FFD28F657FCD0" box="[1275,1284,752,777]" italics="true" pageId="31" pageNumber="102">-</emphasis>
colored spots on trunk; dark dorsolateral or ventrolateral stripe on trunk; dark paravertebral markings on trunk; light
<emphasis id="3CA5EAEFFFB2FF89FABBFCCCF61BFCF4" box="[1343,1352,788,813]" italics="true" pageId="31" pageNumber="102">-</emphasis>
colored postsacral marking bearing anteriorly projecting arms; and caecum and gonads unpigmented. These characters are scored across all
<taxonomicName id="C9D14D7EFFB2FF89FEE4FC83F1C8FCAC" box="[352,667,859,885]" pageId="31" pageNumber="102">
<emphasis id="3CA5EAEFFFB2FF89FEE4FC83F113FCAC" box="[352,576,859,885]" italics="true" pageId="31" pageNumber="102">Hemiphyllodactylus</emphasis>
species
</taxonomicName>
in
<tableCitation id="43530346FFB2FF89FD40FC83F049FCAF" box="[708,794,859,886]" captionStart="TABLE 5" captionStartId="16.[152,244,703,729]" captionTargetPageId="16" captionText="TABLE 5. Significant p-values from the results of the TukeyHSD tests analyses comparing the normalized morphometric and meristic characters of Hemiphyllodactylus" httpUri="http://table.plazi.org/id/5AAE6675FFBDFF86FF1CFD67F3AFFCF8" pageId="31" pageNumber="102" tableUuid="5AAE6675FFBDFF86FF1CFD67F3AFFCF8">Table 5</tableCitation>
and from all other species of
<taxonomicName id="C9D14D7EFFB2FF89FBF8FC83F60FFCAC" box="[1148,1372,859,885]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="31" pageNumber="102" phylum="Chordata" rank="genus">
<emphasis id="3CA5EAEFFFB2FF89FBF8FC83F60FFCAC" box="[1148,1372,859,885]" italics="true" pageId="31" pageNumber="102">Hemiphyllodactylus</emphasis>
</taxonomicName>
from southern
<collectingCountry id="76C6766DFFB2FF89FF7BFCA7F216FC40" box="[255,325,895,921]" name="China" pageId="31" pageNumber="102">China</collectingCountry>
,
<collectingCountry id="76C6766DFFB2FF89FECBFCA7F2FDFC40" box="[335,430,895,921]" name="Vietnam" pageId="31" pageNumber="102">Vietnam</collectingCountry>
and
<collectingCountry id="76C6766DFFB2FF89FE60FC58F149FC43" box="[484,538,896,922]" name="Laos" pageId="31" pageNumber="102">Laos</collectingCountry>
(clades
<quantity id="C9299B18FFB2FF89FDFCFCA7F1F0FC40" box="[632,675,895,921]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="31" pageNumber="102" unit="in" value="6.0">6 in</quantity>
Agung
<emphasis id="3CA5EAEFFFB2FF89FD7FFC58F069FC40" box="[763,826,895,921]" italics="true" pageId="31" pageNumber="102">et al.,</emphasis>
(2022)).
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FFB2FFB7FF43FC7BF6CFF978" lastPageId="33" lastPageNumber="104" pageId="31" pageNumber="102" type="description">
<paragraph id="0E6E36FDFFB2FF89FF43FC7BF7BEF808" blockId="31.[151,1437,283,2002]" pageId="31" pageNumber="102">
<emphasis id="3CA5EAEFFFB2FF89FF43FC7BF2B8FC64" bold="true" box="[199,491,931,957]" pageId="31" pageNumber="102">
Description of
<typeStatus id="D16A885FFFB2FF89FEF9FC7BF2B5FC64" box="[381,486,931,957]" pageId="31" pageNumber="102" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female SVL
<quantity id="C9299B18FFB2FF89FD56FC7BF06FFC67" box="[722,828,931,958]" metricMagnitude="-2" metricUnit="m" metricValue="4.43" pageId="31" pageNumber="102" unit="mm" value="44.3">44.3 mm</quantity>
; head triangular in dorsal profile, depressed, distinct from neck; lores flat; rostrum moderate in length (SnEye/SVL 0.09); prefrontal region weakly concave; canthus rostralis smoothly rounded, barely discernable; snout moderate, rounded in dorsal profile; eye large; ear opening elliptical, small; eye to ear distance greater than diameter of eye; rostral wider than high, bordered posteriorly by large supranasals; two intersupranasals; external nares bordered anteriorly by rostral, dorsally by supranasal, posteriorly by three postnasals, ventrally by first supralabial; 11 (R, L) rectangular supralabials tapering to below posterior margin of eye; 11 (R, L) rectangular infralabials tapering to below posterior margin of eye; scales of rostrum, lores, top of head, and occiput small, raised, those of rostrum largest; dorsal superciliaries flat, mostly square, subimbricate, largest anteriorly; mental triangular, bordered laterally by first infralabials and posteriorly by two parallelogram
<emphasis id="3CA5EAEFFFB2FF89FE0EFB30F2C0FAD8" box="[394,403,1256,1281]" italics="true" pageId="31" pageNumber="102">-</emphasis>
shaped postmentals; each postmental in contact with first infralabial and bordered laterally by a smaller sublabial; seven chin scales; gular scales small, subimbricate, grading posteriorly into slightly larger, subimbricate throat and even larger pectoral scales which grade into slightly larger, subimbricate ventrals. Body somewhat elongate (Trunk/SVL 0.53), dorsoventrally compressed; dorsal scales small, granular, 19 dorsal scales at midbody contained within one eye diameter; ventral scales flat, subimbricate much larger than dorsal scales, eight ventral scales contained within one eye diameter; precloacal scales larger than abdominal scales; 21 continuous, pore
<emphasis id="3CA5EAEFFFB2FF89FF4DFA18F381FA00" box="[201,210,1472,1497]" italics="true" pageId="31" pageNumber="102">-</emphasis>
bearing, femoroprecloacal scales (pitted scales in the juvenile); single enlarged tubercle on lateral margin of tail base; forelimbs short, robust in stature, covered with flat, subimbricate scales dorsally and ventrally; palmar scales slightly raised, subimbricate; all digits except digit I well
<emphasis id="3CA5EAEFFFB2FF89FCAEF9D0F060F9F8" box="[810,819,1544,1569]" italics="true" pageId="31" pageNumber="102">-</emphasis>
developed; digit I vestigial, clawless; distal subdigital lamellae of digits II–V undivided, angular and U
<emphasis id="3CA5EAEFFFB2FF89FD3AF9F4F194F99C" box="[702,711,1580,1605]" italics="true" pageId="31" pageNumber="102">-</emphasis>
shaped, lamellae proximal to these transversely expanded; distal lamellar formula of digits II–V (left) 4-4-4-4; four transversely expanded lamellae on digit I; claws on digits II–V well developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; hind limbs short, more robust than forelimbs, covered with slightly raised, juxtaposed scales dorsally and by larger, flat subimbricate scales anteriorly and ventrally; plantar scales slightly raised, subimbricate; all digits except digit I well
<emphasis id="3CA5EAEFFFB2FF89FEE5F938F239F920" box="[353,362,1760,1785]" italics="true" pageId="31" pageNumber="102">-</emphasis>
developed; digit I vestigial, clawless; distal subdigital lamellae of digits II–V undivided, angular and U
<emphasis id="3CA5EAEFFFB2FF89FF5EF8DCF3B0F8C4" box="[218,227,1796,1821]" italics="true" pageId="31" pageNumber="102">-</emphasis>
shaped, lamellae proximal to these transversely expanded; distal lamellar formula of digits II–V (left) 4-5-5- 5; four transversely expanded lamellae on digit I; claws on digits II–V well
<emphasis id="3CA5EAEFFFB2FF89FC61F8F0F0BDF898" box="[997,1006,1832,1857]" italics="true" pageId="31" pageNumber="102">-</emphasis>
developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; tail original, complete; caudal scales occurring in whorls; dorsal caudal scales larger than dorsal body scales, flat, subcycloid, subimbricate; ventrolateral caudals slightly enlarged, weakly flared anteriorly giving a fringe
<emphasis id="3CA5EAEFFFB2FF89FB9BF84CF77BF874" box="[1055,1064,1940,1965]" italics="true" pageId="31" pageNumber="102">-</emphasis>
like appearance; subcaudals flat, slightly larger than dorsal caudals, not plate
<emphasis id="3CA5EAEFFFB2FF89FD04F860F1DAF808" box="[640,649,1976,2001]" italics="true" pageId="31" pageNumber="102">-</emphasis>
like. Raw morphological data are presented in
<tableCitation id="43530346FFB2FF89FB17F86FF7BBF808" box="[1171,1256,1975,2001]" captionStart="TABLE 8" captionStartId="29.[152,244,151,175]" captionTargetPageId="29" captionText="TABLE 8. Morphological characters of Hemiphyllodactylus houaphanensis sp. nov. from Houaphan Province, Laos and Hemiphyllodactylus vanhonensis sp. nov. from Son La Province, Vietnam (measurements in mm, other abbreviations defined in the text, * partly regenerated tail)." httpUri="http://table.plazi.org/id/5AAE6675FFB0FF8BFF1CFF4FF127FF2E" pageId="31" pageNumber="102" tableUuid="5AAE6675FFB0FF8BFF1CFF4FF127FF2E">Table 8</tableCitation>
.
</paragraph>
<caption id="5AAE6675FF8DFFB6FF1CF84FF033F82E" ID-DOI="http://doi.org/10.5281/zenodo.13209727" ID-Zenodo-Dep="13209727" httpUri="https://zenodo.org/record/13209727/files/figure.png" pageId="32" pageNumber="103" startId="32.[152,255,1943,1967]" targetBox="[284,1303,182,1918]" targetPageId="32" targetType="figure">
<paragraph id="0E6E36FDFF8DFFB6FF1CF84FF033F82E" blockId="32.[151,1437,1943,2040]" pageId="32" pageNumber="103">
<emphasis id="3CA5EAEFFF8DFFB6FF1CF84FF24BF876" bold="true" box="[152,280,1943,1967]" pageId="32" pageNumber="103">FIGURE 9.</emphasis>
Dorsal views of
<taxonomicName id="C9D14D7EFF8DFFB6FE47F84FF057F876" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[451,772,1943,1967]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="32" pageNumber="103" phylum="Chordata" rank="species" species="vanhoensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8DFFB6FE47F84FF057F876" box="[451,772,1943,1967]" italics="true" pageId="32" pageNumber="103">Hemiphyllodactylus vanhoensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8DFFB6FC8DF84FF00AF876" bold="true" box="[777,857,1943,1967]" pageId="32" pageNumber="103">
<taxonomicNameLabel id="27965794FF8DFFB6FC8DF84FF00AF876" box="[777,857,1943,1967]" pageId="32" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Bun Village, Tan Xuan Commune, Van Ho District, Son La Province, Vietnam. A: adult female holotype (VNUF R.2022.07), B: adult female paratype (VNUF R. 2022.09), and C: juvenile male paratype (VNUF R. 2023.22). Photos: Vinh Quang Luu.
</paragraph>
</caption>
<caption id="5AAE6675FF8CFFB7FF1CFB49F075FB15" ID-DOI="http://doi.org/10.5281/zenodo.13209729" ID-Zenodo-Dep="13209729" httpUri="https://zenodo.org/record/13209729/files/figure.png" pageId="33" pageNumber="104" startId="33.[152,255,1169,1193]" targetBox="[151,1436,181,1145]" targetPageId="33" targetType="figure">
<paragraph id="0E6E36FDFF8CFFB7FF1CFB49F075FB15" blockId="33.[151,1436,1169,1229]" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF1CFB49F277FB70" bold="true" box="[152,292,1169,1193]" pageId="33" pageNumber="104">FIGURE 10.</emphasis>
Habitat of
<taxonomicName id="C9D14D7EFF8CFFB7FE11FB49F1B0FB70" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[405,739,1169,1193]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="33" pageNumber="104" phylum="Chordata" rank="species" species="vanhonensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8CFFB7FE11FB49F1B0FB70" box="[405,739,1169,1193]" italics="true" pageId="33" pageNumber="104">Hemiphyllodactylus vanhonensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8CFFB7FD6DFB49F06BFB70" bold="true" box="[745,824,1169,1193]" pageId="33" pageNumber="104">
<taxonomicNameLabel id="27965794FF8CFFB7FD6DFB49F06BFB70" box="[745,824,1169,1193]" pageId="33" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from at the Bun Village, Tan Xuan Commune, Van Ho District, Son La Province, Vietnam. Photo: Vinh Quang Luu.
</paragraph>
</caption>
<paragraph id="0E6E36FDFF8CFFB7FF43FAC7F2D3FA58" blockId="33.[151,1437,1311,2022]" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF43FAC7F2CDFAE0" bold="true" box="[199,414,1311,1337]" pageId="33" pageNumber="104">
Variation (
<figureCitation id="96EA2A78FF8CFFB7FECCFAC7F2DDFAE0" box="[328,398,1311,1337]" captionStart="FIGURE 9" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[284,1303,182,1918]" captionTargetId="figure-36@32.[284,1304,181,1919]" captionTargetPageId="32" captionText="FIGURE 9. Dorsal views of Hemiphyllodactylus vanhoensis sp. nov. from Bun Village, Tan Xuan Commune, Van Ho District, Son La Province, Vietnam. A: adult female holotype (VNUF R.2022.07), B: adult female paratype (VNUF R. 2022.09), and C: juvenile male paratype (VNUF R. 2023.22). Photos: Vinh Quang Luu." figureDoi="http://doi.org/10.5281/zenodo.13209727" httpUri="https://zenodo.org/record/13209727/files/figure.png" pageId="33" pageNumber="104">Fig. 9</figureCitation>
).
</emphasis>
The color patterns of the
<typeStatus id="D16A885FFF8CFFB7FD42FAF8F062FAE3" box="[710,817,1312,1338]" pageId="33" pageNumber="104" type="paratype">paratypes</typeStatus>
generally match that of the
<typeStatus id="D16A885FFF8CFFB7FBF6FAC7F78BFAE0" box="[1138,1240,1311,1337]" pageId="33" pageNumber="104" type="holotype">holotype</typeStatus>
. The dark, dorsal pattern of VNUF R.2022.09 is not as bold as that of the
<typeStatus id="D16A885FFF8CFFB7FC95FA9BF024FA84" box="[785,887,1347,1373]" pageId="33" pageNumber="104" type="holotype">holotype</typeStatus>
. Variation in meristic and morphometric data are presented in
<tableCitation id="43530346FF8CFFB7FEA2FABFF228FA58" box="[294,379,1383,1409]" captionStart="TABLE 8" captionStartId="29.[152,244,151,175]" captionTargetPageId="29" captionText="TABLE 8. Morphological characters of Hemiphyllodactylus houaphanensis sp. nov. from Houaphan Province, Laos and Hemiphyllodactylus vanhonensis sp. nov. from Son La Province, Vietnam (measurements in mm, other abbreviations defined in the text, * partly regenerated tail)." httpUri="http://table.plazi.org/id/5AAE6675FFB0FF8BFF1CFF4FF127FF2E" pageId="33" pageNumber="104" tableUuid="5AAE6675FFB0FF8BFF1CFF4FF127FF2E">Table 8</tableCitation>
.
</paragraph>
<paragraph id="0E6E36FDFF8CFFB7FF43FA53F6CFF978" blockId="33.[151,1437,1311,2022]" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF43FA53F29AFA7C" bold="true" box="[199,457,1419,1445]" pageId="33" pageNumber="104">
Coloration in life (
<figureCitation id="96EA2A78FF8CFFB7FE24FA53F29AFA7C" box="[416,457,1419,1445]" captionStart="FIGURE 9" captionStartId="32.[152,255,1943,1967]" captionTargetBox="[284,1303,182,1918]" captionTargetId="figure-36@32.[284,1304,181,1919]" captionTargetPageId="32" captionText="FIGURE 9. Dorsal views of Hemiphyllodactylus vanhoensis sp. nov. from Bun Village, Tan Xuan Commune, Van Ho District, Son La Province, Vietnam. A: adult female holotype (VNUF R.2022.07), B: adult female paratype (VNUF R. 2022.09), and C: juvenile male paratype (VNUF R. 2023.22). Photos: Vinh Quang Luu." figureDoi="http://doi.org/10.5281/zenodo.13209727" httpUri="https://zenodo.org/record/13209727/files/figure.png" pageId="33" pageNumber="104">Fig</figureCitation>
</emphasis>
.
<emphasis id="3CA5EAEFFF8CFFB7FE52FA53F2A7FA7C" bold="true" box="[470,500,1419,1445]" pageId="33" pageNumber="104">9).</emphasis>
Ground color of top of head, body, and limbs dark
<emphasis id="3CA5EAEFFF8CFFB7FBA2FA54F77CFA7C" box="[1062,1071,1420,1445]" italics="true" pageId="33" pageNumber="104">-</emphasis>
brown; top of head overlain with dark, cluster-shaped pattern; dark and light pre
<emphasis id="3CA5EAEFFF8CFFB7FD25FA68F1F9FA10" box="[673,682,1456,1481]" italics="true" pageId="33" pageNumber="104">-</emphasis>
and postorbital stripes in parallel extending from posterior nares to forelimb insertion; dark, semi
<emphasis id="3CA5EAEFFF8CFFB7FE62FA0CF2BCFA34" box="[486,495,1492,1517]" italics="true" pageId="33" pageNumber="104">-</emphasis>
reticulate paravertebral markings extend from nape to base of tail transforming into a distinct black, post
<emphasis id="3CA5EAEFFF8CFFB7FEFFFA20F2D7F9C8" box="[379,388,1528,1553]" italics="true" pageId="33" pageNumber="104">-</emphasis>
sacral marking; lightcolored dorsolateral spots appear along the trunk; faint yellow post
<emphasis id="3CA5EAEFFF8CFFB7FAD6FA20F608F9C8" box="[1362,1371,1528,1553]" italics="true" pageId="33" pageNumber="104">-</emphasis>
sacral marking bears light
<emphasis id="3CA5EAEFFF8CFFB7FEF3F9C4F2D3F9EC" box="[375,384,1564,1589]" italics="true" pageId="33" pageNumber="104">-</emphasis>
colored anteriorly projecting arms; flanks with faint, dark stripes; limbs bearing irregularly shaped diffuse dark markings and bands; gular and belly regions generally cream, except for darker lateral areas and faint stippling in scales; pigmentation density increases posteriorly; ground color of dorsal caudal region brown, bearing eight dark bands, irregular posteriorly; subcaudal region dull
<emphasis id="3CA5EAEFFF8CFFB7FC1EF950F0F0F978" box="[922,931,1672,1697]" italics="true" pageId="33" pageNumber="104">-</emphasis>
yellow with scattered black and white scales.
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FF8CFFB7FF43F973F033F930" pageId="33" pageNumber="104" type="distribution">
<paragraph id="0E6E36FDFF8CFFB7FF43F973F033F930" blockId="33.[151,1437,1311,2022]" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF43F973F232F91C" bold="true" box="[199,353,1707,1733]" pageId="33" pageNumber="104">Distribution.</emphasis>
<taxonomicName id="C9D14D7EFF8CFFB7FEEFF973F1B3F91C" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[363,736,1707,1733]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="33" pageNumber="104" phylum="Chordata" rank="species" species="vanhonensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8CFFB7FEEFF973F1B3F91C" box="[363,736,1707,1733]" italics="true" pageId="33" pageNumber="104">Hemiphyllodactylus vanhonensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8CFFB7FD6FF973F01BF91C" bold="true" box="[747,840,1707,1733]" pageId="33" pageNumber="104">
<taxonomicNameLabel id="27965794FF8CFFB7FD6FF973F01BF91C" box="[747,840,1707,1733]" pageId="33" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from at the Bun Village, Tan Xuan Commune, Van Ho District,
<collectingRegion id="CC15F81FFF8CFFB7FE53F917F1C7F930" box="[471,660,1743,1770]" country="Vietnam" name="Son La" pageId="33" pageNumber="104">Son La Province</collectingRegion>
,
<collectingCountry id="76C6766DFF8CFFB7FD1BF917F1ADF930" box="[671,766,1743,1769]" name="Vietnam" pageId="33" pageNumber="104">Vietnam</collectingCountry>
(
<figureCitation id="96EA2A78FF8CFFB7FC89F917F01CF930" box="[781,847,1743,1769]" captionStart="FIGURE 1" captionStartId="1.[152,255,1510,1534]" captionTargetBox="[151,1435,705,1487]" captionTargetId="figure-286@1.[151,1435,705,1487]" captionTargetPageId="1" captionText="FIGURE 1. Type localities of the species of Hemiphyllodactylus of clade 6 (Agung et al., 2022)." figureDoi="http://doi.org/10.5281/zenodo.13209711" httpUri="https://zenodo.org/record/13209711/files/figure.png" pageId="33" pageNumber="104">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FF8CFFB7FF43F92BF299F8A3" pageId="33" pageNumber="104" type="biology_ecology">
<paragraph id="0E6E36FDFF8CFFB7FF43F92BF299F8A3" blockId="33.[151,1437,1311,2022]" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF43F92BF2DFF8D4" bold="true" box="[199,396,1779,1805]" pageId="33" pageNumber="104">Natural History.</emphasis>
<taxonomicName id="C9D14D7EFF8CFFB7FE17F92BF056F8D4" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[403,773,1779,1805]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="33" pageNumber="104" phylum="Chordata" rank="species" species="vanhonensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8CFFB7FE17F92BF056F8D4" box="[403,773,1779,1805]" italics="true" pageId="33" pageNumber="104">Hemiphyllodactylus vanhonensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8CFFB7FC89F92BF034F8D4" bold="true" box="[781,871,1779,1805]" pageId="33" pageNumber="104">
<taxonomicNameLabel id="27965794FF8CFFB7FC89F92BF034F8D4" box="[781,871,1779,1805]" pageId="33" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is an upland species and all individuals from Bun Village were found at night between 1900 hrs and 2030 hrs on leaves 4.5–5.0 m above the ground at the entrance of a karstic cave (
<figureCitation id="96EA2A78FF8CFFB7FEE4F8E3F2E7F88C" box="[352,436,1851,1877]" captionStart="FIGURE 10" captionStartId="33.[152,255,1169,1193]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-17@33.[151,1436,181,1145]" captionTargetPageId="33" captionText="FIGURE 10. Habitat of Hemiphyllodactylus vanhonensis sp. nov. is known from at the Bun Village, Tan Xuan Commune, Van Ho District, Son La Province, Vietnam. Photo: Vinh Quang Luu." figureDoi="http://doi.org/10.5281/zenodo.13209729" httpUri="https://zenodo.org/record/13209729/files/figure.png" pageId="33" pageNumber="104">Fig. 10</figureCitation>
). The surrounding habitat was undisturbed forest. Relative temperature was 26.1–27.6
<superScript id="F9A49BB5FF8CFFB7FA05F8E3F6CEF88C" attach="right" box="[1409,1437,1851,1877]" fontSize="6" pageId="33" pageNumber="104">oC</superScript>
and humidity was 63–65%.
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FF8CFFB7FF43F85BF3ADF818" pageId="33" pageNumber="104" type="etymology">
<paragraph id="0E6E36FDFF8CFFB7FF43F85BF3ADF818" blockId="33.[151,1437,1311,2022]" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF43F85BF21DF844" bold="true" box="[199,334,1923,1949]" pageId="33" pageNumber="104">Etymology.</emphasis>
The specific epithet is a toponym referring to the
<typeStatus id="D16A885FFF8CFFB7FCF1F85CF0F6F847" box="[885,933,1924,1950]" pageId="33" pageNumber="104">type</typeStatus>
locality of Van Ho District,
<collectingRegion id="CC15F81FFF8CFFB7FB59F85BF6CBF844" box="[1245,1432,1923,1950]" country="Vietnam" name="Son La" pageId="33" pageNumber="104">Son La Province</collectingRegion>
,
<collectingCountry id="76C6766DFF8CFFB7FF13F87FF3A9F818" box="[151,250,1959,1985]" name="Vietnam" pageId="33" pageNumber="104">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="46CB6576FF8CFFB4FF43F813F0C4FEBC" lastPageId="34" lastPageNumber="105" pageId="33" pageNumber="104" type="discussion">
<paragraph id="0E6E36FDFF8CFFB4FF43F813F0C4FEBC" blockId="33.[151,1437,1311,2022]" lastBlockId="34.[151,1437,151,357]" lastPageId="34" lastPageNumber="105" pageId="33" pageNumber="104">
<emphasis id="3CA5EAEFFF8CFFB7FF43F813F235F83C" bold="true" box="[199,358,1995,2021]" pageId="33" pageNumber="104">Comparisons</emphasis>
. The molecular analyses indicate that
<taxonomicName id="C9D14D7EFF8CFFB7FC97F813F725F83C" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[787,1142,1995,2021]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="33" pageNumber="104" phylum="Chordata" rank="species" species="vanhoensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8CFFB7FC97F813F725F83C" box="[787,1142,1995,2021]" italics="true" pageId="33" pageNumber="104">Hemiphyllodactylus vanhoensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8CFFB7FBFAF813F784F83C" bold="true" box="[1150,1239,1995,2021]" pageId="33" pageNumber="104">
<taxonomicNameLabel id="27965794FF8CFFB7FBFAF813F784F83C" box="[1150,1239,1995,2021]" pageId="33" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the genetically distinct sister species of
<taxonomicName id="C9D14D7EFF8FFFB4FE34FF4FF101FF68" box="[432,594,151,177]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="34" pageNumber="105" phylum="Chordata" rank="species" species="bonkowskii">
<emphasis id="3CA5EAEFFF8FFFB4FE34FF4FF101FF68" box="[432,594,151,177]" italics="true" pageId="34" pageNumber="105">H. bonkowskii</emphasis>
</taxonomicName>
(
<figureCitation id="96EA2A78FF8FFFB4FDE7FF4FF1F9FF68" box="[611,682,151,177]" captionStart="FIGURE 2" captionStartId="21.[152,255,1218,1242]" captionTargetBox="[159,1427,554,1190]" captionTargetId="figure-170@21.[158,1430,549,1195]" captionTargetPageId="21" captionText="FIGURE 2. Maximum likelihood consensus tree of clade 6 of Hemiphyllodactylus (not including H. harterti) with Bayesian posterior probabilities (BPP) and Ultrafast Bootstrap support (UFB) values at the nodes, respectively." figureDoi="http://doi.org/10.5281/zenodo.13209713" httpUri="https://zenodo.org/record/13209713/files/figure.png" pageId="34" pageNumber="105">Fig. 2</figureCitation>
). The new species is most closely related to
<taxonomicName id="C9D14D7EFF8FFFB4FB2FFF4FF61EFF68" box="[1195,1357,151,177]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="34" pageNumber="105" phylum="Chordata" rank="species" species="bonkowskii">
<emphasis id="3CA5EAEFFF8FFFB4FB2FFF4FF61EFF68" box="[1195,1357,151,177]" italics="true" pageId="34" pageNumber="105">H. bonkowskii</emphasis>
</taxonomicName>
with a mean uncorrected pairwise sequence divergence of 3.3% between them (
<tableCitation id="43530346FF8FFFB4FC52FF63F763FF0C" box="[982,1072,187,213]" captionStart="TABLE 2" captionStartId="5.[151,243,640,666]" captionTargetPageId="5" captionText="TABLE 2. Uncorrected genetic P-distances (%) in ND2 gen of Hemiphyllodactylus. Intraspecific p-distance are in bold font, n/a = data not applicable." httpUri="http://table.plazi.org/id/5AAE6675FFA8FF93FF13FD58F5B7FD43" pageId="34" pageNumber="105" tableUuid="5AAE6675FFA8FF93FF13FD58F5B7FD43">Table 2</tableCitation>
).
<taxonomicName id="C9D14D7EFF8FFFB4FBC2FF63F7B5FF0C" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[1094,1254,187,213]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="34" pageNumber="105" phylum="Chordata" rank="species" species="vanhoensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8FFFB4FBC2FF63F7B5FF0C" box="[1094,1254,187,213]" italics="true" pageId="34" pageNumber="105">H. vanhoensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8FFFB4FB6BFF63F618FF0C" bold="true" box="[1263,1355,187,213]" pageId="34" pageNumber="105">
<taxonomicNameLabel id="27965794FF8FFFB4FB6BFF63F618FF0C" box="[1263,1355,187,213]" pageId="34" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs significantly from
<taxonomicName id="C9D14D7EFF8FFFB4FEE8FF07F15DFF20" box="[364,526,223,249]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="34" pageNumber="105" phylum="Chordata" rank="species" species="bonkowskii">
<emphasis id="3CA5EAEFFF8FFFB4FEE8FF07F15DFF20" box="[364,526,223,249]" italics="true" pageId="34" pageNumber="105">H. bonkowskii</emphasis>
</taxonomicName>
in mean values of SnEye (0.58 vs. 0.67,
<emphasis id="3CA5EAEFFF8FFFB4FC6AFF07F0AFFF21" box="[1006,1020,223,248]" italics="true" pageId="34" pageNumber="105">p</emphasis>
= &lt;0.001), SnW (0.12 vs. 0.31,
<emphasis id="3CA5EAEFFF8FFFB4FAF1FF07F6D0FF21" box="[1397,1411,223,248]" italics="true" pageId="34" pageNumber="105">p</emphasis>
= &lt;0.001), VS (9.67 vs. 14.33,
<emphasis id="3CA5EAEFFF8FFFB4FE5AFEDBF2BFFEC5" box="[478,492,259,284]" italics="true" pageId="34" pageNumber="105">p</emphasis>
= &lt;0.001), DS (17.33 vs. 25.17,
<emphasis id="3CA5EAEFFF8FFFB4FCE4FEDBF03DFEC5" box="[864,878,259,284]" italics="true" pageId="34" pageNumber="105">p</emphasis>
= &lt;0.001), SL1F (4.50 vs. 5.00,
<emphasis id="3CA5EAEFFF8FFFB4FB64FEDBF7BDFEC5" box="[1248,1262,259,284]" italics="true" pageId="34" pageNumber="105">p</emphasis>
= &lt;0.001); and SL1T (4.67 vs. 4.83,
<emphasis id="3CA5EAEFFF8FFFB4FEFAFEFFF2DFFE99" box="[382,396,295,320]" italics="true" pageId="34" pageNumber="105">p</emphasis>
= &lt;0.001). Statistically significant and discrete categorical differences between
<taxonomicName id="C9D14D7EFF8FFFB4FB7BFEFFF6CFFE98" authorityName="Luu &amp; Hoang &amp; Ha &amp; Grismer &amp; Murdoch &amp; Sitthivong &amp; Phimpasone &amp; Grismer" authorityYear="2024" box="[1279,1436,295,321]" class="Squamata" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" pageId="34" pageNumber="105" phylum="Chordata" rank="species" species="vanhoensis" status="sp. nov.">
<emphasis id="3CA5EAEFFF8FFFB4FB7BFEFFF6CFFE98" box="[1279,1436,295,321]" italics="true" pageId="34" pageNumber="105">H. vanhoensis</emphasis>
</taxonomicName>
<emphasis id="3CA5EAEFFF8FFFB4FF13FE93F3A2FEBC" bold="true" box="[151,241,331,357]" pageId="34" pageNumber="105">
<taxonomicNameLabel id="27965794FF8FFFB4FF13FE93F3A2FEBC" box="[151,241,331,357]" pageId="34" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and all other species in clade 6 are presented in
<tableCitation id="43530346FF8FFFB4FC8AFE93F03FFEBC" box="[782,876,330,357]" captionStart="TABLE 5" captionStartId="16.[152,244,703,729]" captionTargetPageId="16" captionText="TABLE 5. Significant p-values from the results of the TukeyHSD tests analyses comparing the normalized morphometric and meristic characters of Hemiphyllodactylus" httpUri="http://table.plazi.org/id/5AAE6675FFBDFF86FF1CFD67F3AFFCF8" pageId="34" pageNumber="105" tableUuid="5AAE6675FFBDFF86FF1CFD67F3AFFCF8">Tables 5</tableCitation>
&amp;
<tableCitation id="43530346FF8FFFB4FC0DFE93F0C4FEBC" box="[905,919,331,357]" captionStart="TABLE 6" captionStartId="18.[151,248,464,492]" captionTargetPageId="18" captionText="TABLE 6. Summary statistics of transformed morphometric and meristic characters among the Hemiphyllodactyus species." httpUri="http://table.plazi.org/id/5AAE6675FFBFFF84FF13FE08F555FE32" pageId="34" pageNumber="105" tableUuid="5AAE6675FFBFFF84FF13FE08F555FE32">6</tableCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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