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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.333.5324" ID-GBIF-Dataset="5acfbea6-2243-4afa-8db4-f3991e2f40bf" ID-PMC="PMC3800832" ID-Pensoft-Pub="1313-2970-333-31" ID-PubMed="24146558" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-333-31" ModsDocOrigin="ZooKeys 333" ModsDocTitle="Review of the grassland leafhopper genus Exitianus Ball (Hemiptera, Cicadellidae, Deltocephalinae, Chiasmini) from China" checkinTime="1451246913390" checkinUser="pensoft" docAuthor="Duan, Yani &amp; Zhang, Yalin" docDate="2013" docId="3EB3AA6FCBBA39F16702F684104110AE" docLanguage="en" docName="ZooKeys 333: 31-43" docOrigin="ZooKeys 333" docSource="http://dx.doi.org/10.3897/zookeys.333.5324" docTitle="Exitianus indicus" docType="treatment" docVersion="3" lastPageNumber="40" masterDocId="FFE88A51FFCAFFA1124E473A791EFF8A" masterDocTitle="Review of the grassland leafhopper genus Exitianus Ball (Hemiptera, Cicadellidae, Deltocephalinae, Chiasmini) from China" masterLastPageNumber="43" masterPageNumber="31" pageNumber="36" updateTime="1668156481080" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the grassland leafhopper genus Exitianus Ball (Hemiptera, Cicadellidae, Deltocephalinae, Chiasmini) from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Duan, Yani</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Yalin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>333</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>43</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.333.5324</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.333.5324</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-333-31</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152047770" LSID="urn:lsid:plazi:treatment:3EB3AA6FCBBA39F16702F684104110AE" httpUri="http://treatment.plazi.org/id/3EB3AA6FCBBA39F16702F684104110AE" lastPageId="9" lastPageNumber="40" pageId="5" pageNumber="36">
<subSubSection pageId="5" pageNumber="36" type="nomenclature">
<paragraph pageId="5" pageNumber="36">
<taxonomicName LSID="http://species-id.net/wiki/Exitianus_indicus" authority="Distant" class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">
<pageBreakToken pageId="5" pageNumber="36" start="start">Exitianus</pageBreakToken>
indicus (Distant)
</taxonomicName>
Plate II, Figures 3-6
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="36" type="reference_group">
<paragraph pageId="5" pageNumber="36">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Athysanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Athysanus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Athysanus indicus</taxonomicName>
Distant, 1908: 344.
</paragraph>
<paragraph pageId="5" pageNumber="36">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Athysanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Athysanus atkinsoni" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="atkinsoni">Athysanus atkinsoni</taxonomicName>
;
<bibRefCitation pageId="5" pageNumber="36">Distant 1908</bibRefCitation>
: 345. Synonymized by
<bibRefCitation author="Ross, HH" journalOrPublisher="Bulletin of the British Museum Entomology" pageId="12" pageNumber="43" pagination="1 - 30" title="The evolution and dispersal of the grassland leafhopper genus Exitianus, with keys to the Old World species (Cicadellidae-Hemiptera)." volume="22" year="1968">Ross 1968</bibRefCitation>
: 12.
</paragraph>
<paragraph pageId="5" pageNumber="36">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Exitianus indicus</taxonomicName>
;
<bibRefCitation author="Ross, HH" journalOrPublisher="Bulletin of the British Museum Entomology" pageId="12" pageNumber="43" pagination="1 - 30" title="The evolution and dispersal of the grassland leafhopper genus Exitianus, with keys to the Old World species (Cicadellidae-Hemiptera)." volume="22" year="1968">Ross 1968</bibRefCitation>
: 12, figs 9
<normalizedToken originalValue="10,26">-10,26-</normalizedToken>
30, 69.
</paragraph>
<paragraph pageId="5" pageNumber="36">
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus major" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="major">Exitianus major</taxonomicName>
;
<bibRefCitation author="Ahmed, M" journalOrPublisher="Great Basin Naturalist Memoirs" pageId="11" pageNumber="42" pagination="10 - 17" title="Some new cicadellids from grasslands of Karachi, Pakistan (Homoptera, Cicadellidae)." volume="12" year="1988">Ahmed et al. 1988</bibRefCitation>
: 10, fig. 1. Synonymized by
<bibRefCitation author="Khatri, I" journalOrPublisher="Zootaxa" pageId="11" pageNumber="42" pagination="1 - 47" title="The Deltocephalinae leafhoppers of Pakistan (Hemiptera, Cicadellidae)." volume="2365" year="2010">Khatri and Webb 2010</bibRefCitation>
: 10.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="36" type="description">
<paragraph pageId="5" pageNumber="36">Description.</paragraph>
<paragraph pageId="5" pageNumber="36">Length. Male: 4.0-4.5mm; female: 4.0-5.2mm.</paragraph>
<paragraph pageId="5" pageNumber="36">
Yellow-brown with variable transverse arcuate brown band on vertex (Plate II:
<normalizedToken originalValue="AE">A-E</normalizedToken>
); frontoclypeus with faint brown lateral arcs (Plate II: F). Scutellum with faint brown basal triangles (Plate II:
<normalizedToken originalValue="AE">A-E</normalizedToken>
).
</paragraph>
<paragraph pageId="5" pageNumber="36">Malegenitalia. Pygofer side usually with 2-3 apical brown or black macrosetae (Figs 3-4). Style preapical lobe narrowly triangular, apophysis abruptly tapered at apex (Fig. 5E). Aedeagal shaft strongly laterally compressed, gonopore large with rim forming concave margin in lateral view; with pair of small dorsobasal processes (Figs 5G, 6).</paragraph>
<paragraph pageId="5" pageNumber="36">Female. Sternite VII with posterior margin with a shallow notch in mid-line (Plate II: G).</paragraph>
<caption pageId="5" pageNumber="36">
<paragraph pageId="5" pageNumber="36">
Plate II.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Exitianus indicus</taxonomicName>
.
<normalizedToken originalValue="AE">A-E</normalizedToken>
habitus, dorsal view F face G the end of female abdomen, ventral view.
</paragraph>
</caption>
<caption pageId="5" pageNumber="36">
<paragraph pageId="5" pageNumber="36">
Figure 3.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Exitianus indicus</taxonomicName>
.
<normalizedToken originalValue="AJ">A-J</normalizedToken>
male pygofer side, lateral view from the same specimen, respectively.
</paragraph>
</caption>
<caption pageId="5" pageNumber="36">
<paragraph pageId="5" pageNumber="36">
Figure 4.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Exitianus indicus</taxonomicName>
.
<normalizedToken originalValue="AH">A-H</normalizedToken>
male pygofer side, lateral view.
</paragraph>
</caption>
<caption pageId="5" pageNumber="36">
<paragraph pageId="5" pageNumber="36">
Figure 5.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Exitianus indicus</taxonomicName>
.
<normalizedToken originalValue="AC">A-C</normalizedToken>
subgenital plate, ventral view D valve, ventral view E style, dorsal view F connective, dorsal view G aedeagus, dorsal view.
</paragraph>
</caption>
<caption pageId="5" pageNumber="36">
<paragraph pageId="5" pageNumber="36">
Figure 6.
<taxonomicName class="Insecta" family="Cicadellidae" genus="Exitianus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exitianus indicus" order="Hemiptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="indicus">Exitianus indicus</taxonomicName>
.
<normalizedToken originalValue="AG">A-G</normalizedToken>
aedeagus, lateral view A, B (after Ross, 1968)
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="40" pageId="5" pageNumber="36" type="material examined">
<paragraph pageId="5" pageNumber="36">Material examined.</paragraph>
<paragraph lastPageId="9" lastPageNumber="40" pageId="5" pageNumber="36">
China: Jilin Prov.: 1 male, Lingjiang City, 30.VII.1983, coll. Wu Zhengliang &amp; Hua Baozhen; Zhejiang Prov.: 21 males, 10 females, Wuyanling, 400m, 1.VIII.2005, coll. DY; Fujian Prov.: 1 male, Wuyishan, Nantan, 18.VIII.2002, coll. SQ; 2 males, Guangze County, 24.VIII.2002, coll. SQ; 1 male, Wuyishan Sanggang, 19.VII.2006, coll. YM; 41 males, 10 females, Shanghang County, Buyun Town, 21.VII.2009, coll. CY; 3 males, 1 female, Wuyishan, Tongmucun, 7.VIII.2009, coll. CY; Jiangxi Prov.: 34 males,
<pageBreakToken pageId="6" pageNumber="37" start="start">10</pageBreakToken>
females, Anfu County, 10.VIII.2002; 4 males, 4 females, Ruijin City, Baying Town, 280m, 15.VIII.2004, coll. WC &amp; YM; 7 males, 6 females, Pingxiang City, Lianhua County, 4.VIII.2002, coll. SQ; Henan Prov.: 1 male, Jigongshan, 11.VII.1997, coll. SQ; 1 male, Ruijin City, Baying Town, 280m, 15.VIII.2004, coll. WC &amp; YM; 1 male, Baiyunshan, 17.VIII.2008, coll. LL; Hubei Prov.: 1 male, Wudangshan, 22.VII.2001, coll. Huang Min &amp; Zhang Guiling; Hunan Prov.: 1 male, Chenzhou City, 28.VIII.1985, coll. ZY &amp; Chai Yonghui; 9 males, 3 females, Changde City, Huanan Factory, 22.VII.2002, coll. SQ; 6
<pageBreakToken pageId="7" pageNumber="38" start="start">males</pageBreakToken>
, Zhangjiajie, 25.VII.2002; Guangdong Prov.: 1 male, Dianbai County, 13.IV.1983, coll. ZY; 1 male, Shenzhen, 18.IV.1983, coll. ZY; 1 male, Dinghushan, 7.VII.1985, coll. ZY; Guangxi Prov.: 10 males, 4 females, Huaping, 9.VIII.2000, coll. LZ; 1 male, Huaping, 26.VIII.2000, coll. LZ; 5 males, Fangchenggang City, Naqin Town, 1-3.VIII.2001, coll. He Zhiqiang; 13 males, 6 females, Xinzhai, 18.VIII.2005, coll. YM &amp; KJ;1 male, Yuanbaoshan, 12.VIII.2006, coll. YM &amp; KJ; Hainan Prov.: 10 males, 5 females, Yacheng, 6.V.1983, coll. ZY; 1 male, Wuzhishan, 720m, 31.VII.2009, coll. Gaoxia; 1 male, Bawangling, 9.VIII.2010, coll. WY; 1 male, Tongguling, 26.VIII.2010, coll. WY; Sichuan Prov.:
<pageBreakToken pageId="8" pageNumber="39" start="start">2</pageBreakToken>
males, 9 females, Kangding, 2500m, 8.XI.1999, coll. Qin Daozheng; 2 males, 8 females, Zhubalong, 2450m, 11.VII.2011, coll. Sun Qiang; Guizhou Prov.: 5 males, Guiyang City, Huaxi Park, 1100m, 25.VII.2001; Yunnan Prov.: 1 male, Daluo Town, 26.X.1987, coll. Feng Jinian &amp; Lili; 10 males, 8 females, Mengla County, Yaoqu Town, V.1991; 1 male, Tengchong County, 22.XI.1999; 2 males, Xishuangbanna, 11.VII.2003; 1 male, Mengla City, 21.VII.2005, coll. LL; 1 male, Simao City, 30.VII.2005, coll. LL; 31 males, 20 females, Tengchong County, Huoshan Park, 1930m, 14.VIII.2005, coll. YM &amp; KJ; 1 male, Dali, 29.VIII.2010, coll. ZM;1 male, Diqing, 14.VIII.2010, coll. ZM; 1 male, Jinghong City, 1.IX.2010, coll. ZM; 1 male, Daluo Town, 679m, 23.V.2011, coll. LL; 1 male, Lancang County, 25.V.2011, coll. LL; 1 male, Nansan Town, 29.V.2011, coll. LL; 1 male, Yingjiang County, Zhanxi Town, 1009m, 2.VI.2011, coll. LL; 1 male, Tengchong County, 1632m, 5.VI.2011, coll. LL; 4 males, Lushui County, Chenggan Town, 1013m, 6.VI.2011, coll. LL; 1 male, Zhenyuan County, 13.VI.2011, coll. LL; Shaanxi Prov.: 1 male, Liuba County, 20.VII.1995, coll. ZR; 3 males, 3 females, Liuba County,
<pageBreakToken pageId="9" pageNumber="40" start="start">20</pageBreakToken>
.VIII.1995, coll. ZR; 1 male, Zhouzhi County, 24.IX.2008, coll. LL; 1 male, Foping County, 1060m, 1.X.2008, coll. LL; Gansu Prov.: 1 male, Cheng County, 25.VII.2002, coll. WC &amp; Shang Suqin; 1 male, Ruijin City, Baying Town, 280m, 15.VIII.2004, coll. WC &amp; YM. All deposited in NWAFU and mainly collected at light. Abbreviations for collectors: DY: Duan Yani; SQ: Sun Qinxia; LL: Lu Lin; YM: Yang Meixia; CY: Cao Yanghui; ZY: Zhang Yalin; ZM: Zhang Meng; ZR: Zhang Wenzhu &amp; Ren Liyun; WC: Wei Cong; LZ: Liu Zhenjiang; KJ: Kang Juxia; WY: Wang Yang.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="40" type="distribution">
<paragraph pageId="9" pageNumber="40">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="40">Eastern Hemisphere.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="40" type="remarks">
<paragraph pageId="9" pageNumber="40">Remarks.</paragraph>
<paragraph pageId="9" pageNumber="40">Considerablevariation was found in the apical black macrosetae of the male pygofer and shape of the aedeagus in this species (see Figs 3, 4 &amp; 6).</paragraph>
</subSubSection>
</treatment>
</document>