treatments-xml/data/CF/3A/F0/CF3AF04A5A77FFA6FF1AFE04311B0DA6.xml
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<mods:title id="BC40423FBADBA12186BBDEE0579EB0B9">On Verhoeffs Otostigmus subgenus Malaccopleurus, the nudus group of Otostigmus subgenus Otostigmus Porat, 1876, and Digitipes Attems, 1930, with a description of the foetus stadium larva in O. sulcipes Verhoeff, 1937, (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<mods:namePart id="862380CE576E85B90C1F4BCD585DA057">Lewis, John G. E.</mods:namePart>
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<treatment id="CF3AF04A5A77FFA6FF1AFE04311B0DA6" ID-DOI="http://doi.org/10.5281/zenodo.5271641" ID-GBIF-Taxon="119644167" ID-Zenodo-Dep="5271641" LSID="urn:lsid:plazi:treatment:CF3AF04A5A77FFA6FF1AFE04311B0DA6" httpUri="http://treatment.plazi.org/id/CF3AF04A5A77FFA6FF1AFE04311B0DA6" lastPageId="4" lastPageNumber="229" pageId="2" pageNumber="227">
<subSubSection id="0F8912D75A77FFA0FF1AFE0436550FA1" box="[151,635,437,463]" pageId="2" pageNumber="227" type="nomenclature">
<paragraph id="472C415C5A77FFA0FF1AFE0436550FA1" blockId="2.[151,635,437,496]" box="[151,635,437,463]" pageId="2" pageNumber="227">
<heading id="1C64F6305A77FFA0FF1AFE0436550FA1" bold="true" box="[151,635,437,463]" fontSize="11" level="1" pageId="2" pageNumber="227" reason="1">
<taxonomicName id="80933ADF5A77FFA0FF1AFE0436550FA1" ID-CoL="4B7ZG" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,635,437,463]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A77FFA0FF1AFE0436550FA1" bold="true" box="[151,635,437,463]" pageId="2" pageNumber="227">
<emphasis id="75E79D4E5A77FFA0FF1AFE04350E0FA1" bold="true" box="[151,288,437,463]" italics="true" pageId="2" pageNumber="227">Otostigmus</emphasis>
(
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<collectingCountry id="3F8401CC5A77FFA0FEBCFE0435660FA1" box="[305,328,437,463]" name="Iceland" pageId="2" pageNumber="227">O</collectingCountry>
.
</emphasis>
)
<emphasis id="75E79D4E5A77FFA0FED3FE0435900FA1" bold="true" box="[350,446,437,463]" italics="true" pageId="2" pageNumber="227">sulcipes</emphasis>
<bibRefCitation id="23023CAD5A77FFA0FE48FE0436550FA1" author="Verhoeff, K. W." box="[453,635,437,463]" firstAuthor="Verhoeff" journalOrPublisher="Bulletin of the Raffles Museum" pageId="2" pageNumber="227" pagination="198 - 270" part="13" refId="ref15522" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270 + pls. 13 - 24." title="Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A77FFA0FF1AFE6635370F9E" box="[151,281,471,496]" pageId="2" pageNumber="227" type="description">
<paragraph id="472C415C5A77FFA0FF1AFE6635370F9E" blockId="2.[151,635,437,496]" box="[151,281,471,496]" pageId="2" pageNumber="227">
(
<figureCitation id="DFA85DD95A77FFA0FF12FE66353E0F9E" box="[159,272,471,496]" captionStart-0="FIGURES 1 8" captionStart-1="FIGURES 916" captionStartId-0="3.[151,264,1856,1878]" captionStartId-1="5.[151,264,1734,1756]" captionTargetBox-0="[183,1395,205,1814]" captionTargetBox-1="[282,1300,207,1715]" captionTargetId-0="figure@3.[176,1406,193,1832]" captionTargetId-1="figure@5.[281,1310,193,1726]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." captionText-1="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri-0="https://zenodo.org/record/232675/files/figure.png" httpUri-1="https://zenodo.org/record/232676/files/figure.png" pageId="2" pageNumber="227">Figs 116</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A77FFA0FF1AFD9031180C1D" pageId="2" pageNumber="227" type="reference_group">
<paragraph id="472C415C5A77FFA0FF1AFD9036950C3A" blockId="2.[151,1435,544,627]" pageId="2" pageNumber="227">
<treatmentCitationGroup id="678366725A77FFA0FF1AFD9036950C3A" pageId="2" pageNumber="227">
<treatmentCitation id="C632674D5A77FFA0FF1AFD9036DD0C58" author="Verhoeff" box="[151,755,544,566]" page="204" pageId="2" pageNumber="227" year="1937">
<taxonomicName id="80933ADF5A77FFA0FF1AFD9036EC0C58" ID-CoL="4B7ZG" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,706,544,566]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="sulcipes" subGenus="Malaccopleurus">
<emphasis id="75E79D4E5A77FFA0FF1AFD9035250C58" box="[151,267,545,566]" italics="true" pageId="2" pageNumber="227">Otostigmus</emphasis>
(
<emphasis id="75E79D4E5A77FFA0FE97FD9135900C58" box="[282,446,544,566]" italics="true" pageId="2" pageNumber="227">Malaccopleurus</emphasis>
)
<emphasis id="75E79D4E5A77FFA0FE43FD91360E0C58" box="[462,544,544,566]" italics="true" pageId="2" pageNumber="227">sulcipes</emphasis>
<bibRefCitation id="23023CAD5A77FFA0FDAAFD9136EC0C58" author="Verhoeff, K. W." box="[551,706,544,566]" firstAuthor="Verhoeff" journalOrPublisher="Bulletin of the Raffles Museum" pageId="2" pageNumber="227" pagination="198 - 270" part="13" refId="ref15522" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270 + pls. 13 - 24." title="Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</taxonomicName>
, 204
</treatmentCitation>
(key), 205, Pl. 12,
<figureCitation id="DFA85DD95A77FFA0FC36FD91303A0C58" box="[955,1044,544,566]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 5, 6</figureCitation>
. Gunong Brinchang, Pahang in
<quantity id="806BECB95A77FFA0FAD1FD9135390C3A" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="2" pageNumber="227" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1500 1800 m</quantity>
.
<collectingCountry id="3F8401CC5A77FFA0FEADFD8F35580C3A" box="[288,374,574,596]" name="Malaysia" pageId="2" pageNumber="227">Malacca</collectingCountry>
[
<collectingCountry id="3F8401CC5A77FFA0FE0FFD8F35FD0C3A" box="[386,467,574,596]" name="Malaysia" pageId="2" pageNumber="227">Melaka</collectingCountry>
, Peninsular
<collectingCountry id="3F8401CC5A77FFA0FDC3FD8F369F0C3A" box="[590,689,574,596]" name="Malaysia" pageId="2" pageNumber="227">Malaysia</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="472C415C5A77FFA0FF1AFDEF31180C1D" blockId="2.[151,1435,544,627]" box="[151,1334,605,627]" pageId="2" pageNumber="227">
<treatmentCitationGroup id="678366725A77FFA0FF1AFDEF31180C1D" box="[151,1334,605,627]" pageId="2" pageNumber="227">
<treatmentCitation id="C632674D5A77FFA0FF1AFDEF37290C1D" author="Verhoeff" box="[151,775,605,627]" page="204" pageId="2" pageNumber="227" year="1937">
<taxonomicName id="80933ADF5A77FFA0FF1AFDEF36F90C1D" ID-CoL="4B7ZJ" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[151,727,605,627]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="trisulcatus" subGenus="Malaccopleurus">
<emphasis id="75E79D4E5A77FFA0FF1AFDEF35250C1D" box="[151,267,606,627]" italics="true" pageId="2" pageNumber="227">Otostigmus</emphasis>
(
<emphasis id="75E79D4E5A77FFA0FE94FDEC35930C1D" box="[281,445,605,627]" italics="true" pageId="2" pageNumber="227">Malaccopleurus</emphasis>
)
<emphasis id="75E79D4E5A77FFA0FE41FDEC36190C1D" box="[460,567,605,627]" italics="true" pageId="2" pageNumber="227">trisulcatus</emphasis>
<bibRefCitation id="23023CAD5A77FFA0FDB0FDEC36F90C1D" author="Verhoeff, K. W." box="[573,727,605,627]" firstAuthor="Verhoeff" journalOrPublisher="Bulletin of the Raffles Museum" pageId="2" pageNumber="227" pagination="198 - 270" part="13" refId="ref15522" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270 + pls. 13 - 24." title="Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore" type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
</taxonomicName>
, 204
</treatmentCitation>
(key), 207, Pl. 12,
<figureCitation id="DFA85DD95A77FFA0FC44FDEC30290C1D" box="[969,1031,605,627]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 7</figureCitation>
.
<emphasis id="75E79D4E5A77FFA0FB9FFDEC305D0C1D" bold="true" box="[1042,1139,605,627]" pageId="2" pageNumber="227">Syn. nov.</emphasis>
<collectingCountry id="3F8401CC5A77FFA0FBF7FDEC311D0C1D" box="[1146,1331,605,627]" name="Malaysia" pageId="2" pageNumber="227">Melaka, Malaysia</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A77FFA0FF1AFD2E30DB0DFE" pageId="2" pageNumber="227" type="diagnosis">
<paragraph id="472C415C5A77FFA0FF1AFD2E30DB0DFE" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
<emphasis id="75E79D4E5A77FFA0FF1AFD2E353F0CD6" bold="true" box="[151,273,671,696]" pageId="2" pageNumber="227">Diagnosis.</emphasis>
Body slender. Length up to
<quantity id="806BECB95A77FFA0FDDAFD1136850CD9" box="[599,683,672,696]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="227" unit="mm" value="47.0">47 mm</quantity>
. Antennal articles 18, the basal three glabrous dorsally. Forcipular coxosternal tooth-plates typically with 4+4 teeth. Tergites with complete paramedian sutures from 5, marginate from 7, 8 or 9, without keels or spinules. Sternite paramedian sutures almost complete, a median longitudinal and small round posterior median depression present on most sternites or absent. Coxopleural process with two apical spines. Coxopleural pore-field narrow, not expanded anteriorly. Legs without or leg 1 or legs 1 and 2 with a tarsal spur. Ultimate leg prefemur, femur and tibia with a low longitudinal dorsal ridge demarcated by grooves, prefemoral spines VL 3, VM 2 or 3, M 2, 3 or 4, DM 2 or 3, CS 1 (four rows, 1012 spines in toto).
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A77FFA0FF4AFC2D316B0D93" pageId="2" pageNumber="227" type="materials_examined">
<paragraph id="472C415C5A77FFA0FF4AFC2D316B0D93" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
<emphasis id="75E79D4E5A77FFA0FF4AFC2D35DE0DDB" bold="true" box="[199,496,924,949]" pageId="2" pageNumber="227">
<typeStatus id="9828FFFE5A77FFA0FF4AFC2D352C0DDB" box="[199,258,924,949]" pageId="2" pageNumber="227">Type</typeStatus>
material examined.
</emphasis>
<materialsCitation id="F7FB4B015A77FFA0FE7AFC2C316B0D93" ID-GBIF-Occurrence="3924454302" collectionCode="ZMB" elevation="908" location="Malacca" municipality="Malacca" pageId="2" pageNumber="227" specimenCount="1" typeStatus="syntype">
<collectionCode id="2182D9995A77FFA0FE7AFC2C361A0DDA" box="[503,564,925,948]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="227">ZMB</collectionCode>
13364
<typeStatus id="9828FFFE5A77FFA0FD04FC2D36C90DDA" box="[649,743,924,948]" pageId="2" pageNumber="227" type="syntype">syntype</typeStatus>
(s).
<taxonomicName id="80933ADF5A77FFA0FC81FC2D37A70DDA" box="[780,905,924,948]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A77FFA0FC81FC2D37A70DDA" box="[780,905,924,948]" italics="true" pageId="2" pageNumber="227">O. sulcipes</emphasis>
</taxonomicName>
Malacca (one vial and 2 slides, 3613 &amp; 3614). In vial:
<taxonomicName id="80933ADF5A77FFA0FF76FC7135CF0DB6" box="[251,481,960,984]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A77FFA0FF76FC7135CF0DB6" box="[251,481,960,984]" italics="true" pageId="2" pageNumber="227">Otostigmus sulcipes</emphasis>
</taxonomicName>
Malacca, Verh. Slides:
<taxonomicName id="80933ADF5A77FFA0FC88FC7130190DB6" authority="Verh." authorityName="Verh." box="[773,1079,959,984]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="227" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A77FFA0FC88FC7137C20DB6" box="[773,1004,960,984]" italics="true" pageId="2" pageNumber="227">Otostigmus sulcipes</emphasis>
Verh.
</taxonomicName>
Pahang
<quantity id="806BECB95A77FFA0FB29FC0E31070DB9" box="[1188,1321,959,984]" metricMagnitude="2" metricUnit="m" metricValue="9.075" metricValueMax="18.0" metricValueMin="0.15" pageId="2" pageNumber="227" unit="m" value="907.5" valueMax="1800.0" valueMin="15.0">
<elevation id="CCBEA66F5A77FFA0FB29FC0E31070DB9" box="[1188,1321,959,984]" metricMagnitude="2" metricUnit="m" metricValue="9.075" metricValueMax="18.0" metricValueMin="0.15" pageId="2" pageNumber="227" unit="m" value="907.5" valueMax="1800.0" valueMin="15.0">151800 m</elevation>
</quantity>
<collectingMunicipality id="A748DB265A77FFA0FABBFC0E31B70DB6" box="[1334,1433,959,984]" pageId="2" pageNumber="227">Malacca</collectingMunicipality>
. Syntypus 3613 with segments 1821 and ultimate legs. 3614 with head, mandibles and first four segments.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A77FFA0FF4AFBB636F409A7" pageId="2" pageNumber="227" type="description">
<paragraph id="472C415C5A77FFA0FF4AFBB637B90A06" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
<emphasis id="75E79D4E5A77FFA0FF4AFBB636790A4E" bold="true" box="[199,599,1031,1056]" pageId="2" pageNumber="227">
Description of
<typeStatus id="9828FFFE5A77FFA0FEFBFBB635FC0A4E" box="[374,466,1031,1056]" pageId="2" pageNumber="227" type="syntype">syntype</typeStatus>
in ethanol.
</emphasis>
(Verhoeffs data in parentheses where relevant). Slender (very drawn out). Length
<quantity id="806BECB95A77FFA0FF62FB9D356E0A2A" box="[239,320,1068,1093]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="227" unit="mm" value="35.0">35 mm</quantity>
(
<quantity id="806BECB95A77FFA0FEDDFB9D35FF0A2A" box="[336,465,1068,1093]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="2" pageNumber="227" unit="mm" value="39.5" valueMax="43.0" valueMin="36.0">3643 mm</quantity>
). Trunk greyish orange with faint turquoise grey paramedian and marginal stripes on segments 720 (greyish yellow with four green longitudinal stripes).
</paragraph>
<paragraph id="472C415C5A77FFA0FF4AFBC535EB0A96" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
Antennae of 18 articles, the basal three glabrous dorsally (the two basal little haired). Forcipular coxosternal tooth-plates as long as wide, each with four teeth, the central two the larger (
<figureCitation id="DFA85DD95A77FFA0FB95FB2630750ADE" box="[1048,1115,1175,1200]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 1</figureCitation>
) (base obtuse without deep incision (i.e. not separated by a deep groove into two groups of two teeth)). Trochanteroprefemoral process with a single low medial denticle.
</paragraph>
<paragraph id="472C415C5A77FFA0FF4AFAB5371F0B2E" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">Tergites elongated with complete paramedian sutures from 5, marginate from 8 (7 or 8). Tergite of ultimate legbearing segment with shallow posterior median depression.</paragraph>
<paragraph id="472C415C5A77FFA0FF4AFAFD30A80B9A" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
Sternites 219 with almost complete paramedian sutures in sulci and ending in depressions (pits) and with a median longitudinal and small round posterior median depression on most sternites (
<figureCitation id="DFA85DD95A77FFA0FBF3FADE30E80BE6" box="[1150,1222,1391,1416]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 2</figureCitation>
) (three deep sulci (
<emphasis id="75E79D4E5A77FFA0FF2DFA2534D00BC2" box="[160,254,1428,1452]" italics="true" pageId="2" pageNumber="227">Furchen</emphasis>
) and a posterior pit). Sternite of ultimate leg-bearing segment with sides converging posteriorly and posterior margin slightly concave. Coxopleural process with two (one) apical spines. The pore-field narrow (
<figureCitation id="DFA85DD95A77FFA0FADCFA0631BD0BBE" box="[1361,1427,1463,1488]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 3</figureCitation>
) (pore-field narrow, scarcely widened anteriorly where it is no more than six pores across).
</paragraph>
<paragraph id="472C415C5A77FFA0FF4AF9B0368B08EB" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
Leg 1 with a tarsal spur, 221 without. Ultimate legs slender, prefemur and femur with a dorsal longitudinal ridge demarcated by a groove on each side and pitted (
<figureCitation id="DFA85DD95A77FFA0FD70F995376C0852" box="[765,834,1572,1597]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 4</figureCitation>
), the tibia with a weak longitudinal ridge. Prefemoral spines VL 3, VM 2, M 3, DM 2/3, CS 1 (with 11 spines, 3 ventral, 5 medial, 2 dorsomedial). Prefemora of penultimate legs also with a weak dorsal ridge.
</paragraph>
<paragraph id="472C415C5A77FFA0FF4AF93E3170097B" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
<emphasis id="75E79D4E5A77FFA0FF4AF93E36F008C6" bold="true" box="[199,734,1679,1704]" pageId="2" pageNumber="227">
Description of
<typeStatus id="9828FFFE5A77FFA0FEFAF93E35FD08C6" box="[375,467,1679,1704]" pageId="2" pageNumber="227" type="syntype">syntype</typeStatus>
on slides 3613 &amp; 3614
</emphasis>
. Antennae of 18 articles. Articles1 and 2 with a few fine setae dorsally increasing on 3 (
<figureCitation id="DFA85DD95A77FFA0FE34F90535D008A3" box="[441,510,1716,1741]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 5</figureCitation>
), dense on 4.Ventrally with scattered setae on articles 1 and 2, dense on article 3 (
<figureCitation id="DFA85DD95A77FFA0FF12F96634C4089E" box="[159,234,1751,1776]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 6</figureCitation>
). Presumably Verhoeffs “the two basal little haired” refers to the ventral surfaces. Each forcipular coxosternal tooth-plate with four teeth. Trochanteroprefemoral process with one or two low medial denticles.
</paragraph>
<paragraph id="472C415C5A77FFA0FF4AF8AE36F409A7" blockId="2.[151,1437,671,1993]" pageId="2" pageNumber="227">
Sternite of ultimate leg-bearing segment with sides converging posteriorly and posterior margin slightly concave. Coxopleural process with two apical spines. This preparation shows the narrow pore-field, scarcely widened anteriorly where it is no more than six pores across (
<figureCitation id="DFA85DD95A77FFA0FCE4F8D6379E09EE" box="[873,944,1895,1920]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 7</figureCitation>
). Leg 1 with a tarsal spur,
<date id="332D679C5A77FFA0FB62F8D931B209EE" box="[1263,1436,1895,1920]" pageId="2" pageNumber="227" value="1921-02-04" valueMax="1921-02-18" valueMin="1921-02-04">24 and 1821</date>
without. Ultimate leg prefemur with 11 spines VL 3, VM 2 or 3, M 2 or 3, DM 2, CS1 (
<figureCitation id="DFA85DD95A77FFA0FBE1F83D309E09CA" box="[1132,1200,1932,1957]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1856,1878]" captionTargetBox="[183,1395,205,1814]" captionTargetId="figure@3.[176,1406,193,1832]" captionTargetPageId="3" captionText="FIGURES 1 8. Otostigmus sulcipes. 1. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol. 2. Sternite 11, the same. 3. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view. 4. Left ultimate leg prefemur and femur dorsal, the same. 5. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view. 6. Antennal articles 1 to 3, the same, ventral view. 7. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view. 8. Ultimate leg prefemur, the same, ventral view. Scale bars = 1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm." httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="2" pageNumber="227">Fig. 8</figureCitation>
). The dorsal grooves on the ultimate leg not seen in this cleared material.
</paragraph>
</subSubSection>
<caption id="13EC11D45A76FFA1FF1AF8F136870981" httpUri="https://zenodo.org/record/232675/files/figure.png" pageId="3" pageNumber="228" targetBox="[183,1395,205,1814]" targetPageId="3">
<paragraph id="472C415C5A76FFA1FF1AF8F136870981" blockId="3.[151,1436,1856,2031]" pageId="3" pageNumber="228">
<emphasis id="75E79D4E5A76FFA1FF1AF8F135150938" bold="true" box="[151,315,1856,1878]" pageId="3" pageNumber="228">FIGURES 18.</emphasis>
<emphasis id="75E79D4E5A76FFA1FECCF8F0363A0938" box="[321,532,1856,1878]" italics="true" pageId="3" pageNumber="228">
<taxonomicName id="80933ADF5A76FFA1FECCF8F0363F0938" box="[321,529,1856,1878]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="3" pageNumber="228" phylum="Arthropoda" rank="species" species="sulcipes">Otostigmus sulcipes</taxonomicName>
.
</emphasis>
<emphasis id="75E79D4E5A76FFA1FD96F8F136090938" bold="true" box="[539,551,1856,1878]" pageId="3" pageNumber="228">1</emphasis>
. Forcipular coxopleural tooth-plates and left forcipular trochanteroprefemoral process, syntype ZMB 13364 Malacca, spm. in ethanol.
<emphasis id="75E79D4E5A76FFA1FD0BF8EF36BD091A" bold="true" box="[646,659,1886,1908]" pageId="3" pageNumber="228">2</emphasis>
. Sternite 11, the same.
<emphasis id="75E79D4E5A76FFA1FC04F8EF37BB091A" bold="true" box="[905,917,1886,1908]" pageId="3" pageNumber="228">3</emphasis>
. Coxopleuron and sternite of ultimate leg-bearing segment, the same, ventral view.
<emphasis id="75E79D4E5A76FFA1FE65F8CC35DA09FD" bold="true" box="[488,500,1917,1939]" pageId="3" pageNumber="228">4</emphasis>
. Left ultimate leg prefemur and femur dorsal, the same.
<emphasis id="75E79D4E5A76FFA1FBB9F8CC306E09FD" bold="true" box="[1076,1088,1917,1939]" pageId="3" pageNumber="228">5</emphasis>
. Antennal article 3, syntype ZMB 13364, slide 3614, dorsal view.
<emphasis id="75E79D4E5A76FFA1FE57F82D35C809DC" bold="true" box="[474,486,1948,1970]" pageId="3" pageNumber="228">6</emphasis>
. Antennal articles 1 to 3, the same, ventral view.
<emphasis id="75E79D4E5A76FFA1FC53F82D37C409DC" bold="true" box="[990,1002,1948,1970]" pageId="3" pageNumber="228">7</emphasis>
. Coxopleuron and sternite of ultimate legbearing segment syntype ZMB 13364, slide 3613, ventral view.
<emphasis id="75E79D4E5A76FFA1FCADF80B371D09BE" bold="true" box="[800,819,1978,2000]" pageId="3" pageNumber="228">8.</emphasis>
Ultimate leg prefemur, the same, ventral view. Scale bars =1 mm except Figs 1, 3, 5 &amp; 6 = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="0F8912D75A71FFA6FF4AFF26379C0E96" pageId="4" pageNumber="229" type="discussion">
<paragraph id="472C415C5A71FFA6FF4AFF26379C0E96" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="229">
<emphasis id="75E79D4E5A71FFA6FF4AFF26351A0EDE" bold="true" box="[199,308,151,176]" pageId="4" pageNumber="229">Remarks</emphasis>
. The characteristics of
<taxonomicName id="80933ADF5A71FFA6FDB4FF2937370EDE" box="[569,793,152,176]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="229" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A71FFA6FDB4FF2937370EDE" box="[569,793,152,176]" italics="true" pageId="4" pageNumber="229">Otostigmus sulcipes</emphasis>
</taxonomicName>
are seen in a number of
<taxonomicName id="80933ADF5A71FFA6FBA1FF29316A0EDE" authority="OtostIgmus" authorityName="OtostIgmus" box="[1068,1348,152,176]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="75E79D4E5A71FFA6FBA1FF2930850EC1" box="[1068,1195,152,175]" italics="true" pageId="4" pageNumber="229">Otostigmus</emphasis>
(
<emphasis id="75E79D4E5A71FFA6FB36FF2931140EC1" box="[1211,1338,152,175]" italics="true" pageId="4" pageNumber="229">Otostigmus</emphasis>
)
</taxonomicName>
species and it, and the other two “
<emphasis id="75E79D4E5A71FFA6FE48FF0D36550EBA" box="[453,635,188,212]" italics="true" pageId="4" pageNumber="229">Malaccopleurus</emphasis>
” species are here assigned to the subgenus
<emphasis id="75E79D4E5A71FFA6FBFCFF0D30D60EBD" box="[1137,1272,188,211]" italics="true" pageId="4" pageNumber="229">
<taxonomicName id="80933ADF5A71FFA6FBFCFF0D30DA0EBD" box="[1137,1268,188,211]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="229" phylum="Arthropoda" rank="genus">Otostigmus</taxonomicName>
.
</emphasis>
It is proposed below that
<taxonomicName id="80933ADF5A71FFA6FE9EFF5135850E96" box="[275,427,224,248]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="229" phylum="Arthropoda" rank="species" species="trisulcatus">
<emphasis id="75E79D4E5A71FFA6FE9EFF5135850E96" box="[275,427,224,248]" italics="true" pageId="4" pageNumber="229">
<collectingCountry id="3F8401CC5A71FFA6FE9EFF5135040E99" box="[275,298,224,247]" name="Iceland" pageId="4" pageNumber="229">O</collectingCountry>
. trisulcatus
</emphasis>
</taxonomicName>
is a junior subjective synonym of
<emphasis id="75E79D4E5A71FFA6FCA3FF51379C0E96" box="[814,946,224,248]" italics="true" pageId="4" pageNumber="229">
<taxonomicName id="80933ADF5A71FFA6FCA3FF5137800E96" box="[814,942,224,248]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="229" phylum="Arthropoda" rank="species" species="sulcipes">
<collectingCountry id="3F8401CC5A71FFA6FCA3FF51376B0E99" box="[814,837,224,247]" name="Iceland" pageId="4" pageNumber="229">O</collectingCountry>
. sulcipes
</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A71FFA6FF4AFEB5366B0FE6" pageId="4" pageNumber="229" type="materials_examined">
<paragraph id="472C415C5A71FFA6FF4AFEB5366B0FE6" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="229">
<emphasis id="75E79D4E5A71FFA6FF4AFEB536290F73" bold="true" box="[199,519,260,285]" pageId="4" pageNumber="229">Other material examined.</emphasis>
<materialsCitation id="F7FB4B015A71FFA6FD9EFEB5366B0FE6" ID-GBIF-Occurrence="3924454301" collectionCode="ZSM" pageId="4" pageNumber="229" specimenCount="1">
<collectionCode id="2182D9995A71FFA6FD9EFEB5367E0F72" box="[531,592,260,284]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="229">ZSM</collectionCode>
. Two foetus stage larvae in ethanol labelled
<taxonomicName id="80933ADF5A71FFA6FBE0FEB534F10F2E" authority="Verh . Colln." authorityName="Verh . Colln." class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="229" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A71FFA6FBE0FEB5317D0F72" box="[1133,1363,260,284]" italics="true" pageId="4" pageNumber="229">Otostigmus sulcipes</emphasis>
Verh. Colln.
</taxonomicName>
, two on slides labelled
<taxonomicName id="80933ADF5A71FFA6FE6EFE9937AA0F2E" authority="Verh. Fotus. The" authorityName="Verh. Fotus. The" box="[483,900,295,320]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="4" pageNumber="229" phylum="Arthropoda" rank="species" species="sulcipes">
<emphasis id="75E79D4E5A71FFA6FE6EFE9936ED0F2E" box="[483,707,296,320]" italics="true" pageId="4" pageNumber="229">Otostigmus sulcipes</emphasis>
Verh. Fötus. The
</taxonomicName>
first (spm. 1) without a registration number, the second (spm. 2) Reg. Nr.A 20030600 with head, forcipules and segment 1, segments 19, 20 and 21, detached ultimate legs and four trunk segments.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0F8912D75A71FFA6FF4AFE25311B0DA6" pageId="4" pageNumber="229" type="description">
<paragraph id="472C415C5A71FFA6FF4AFE2534D70CC6" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="229">
<emphasis id="75E79D4E5A71FFA6FF4AFE2536800FC3" bold="true" box="[199,686,404,429]" pageId="4" pageNumber="229">Composite description of foetus stadium.</emphasis>
(Verhoeffs data in parentheses where relevant). Length
<quantity id="806BECB95A71FFA6FAA7FE2531B60FC2" box="[1322,1432,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="4" pageNumber="229" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
. Unpigmented apart from pale orange brown yolk-filled mid-gut. Antennae of 17 articles, the elongated penultimate showing a subdivision into two beneath the foetal cuticle (
<figureCitation id="DFA85DD95A71FFA6FCB1FE6D37AC0F9A" box="[828,898,476,501]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 9</figureCitation>
) (18 segmented but 16 and 17 still not clearly divided). With a few scattered setae, the basal articles wider than long. Cephalic plate slightly wider than long, the sides converging posteriorly (
<figureCitation id="DFA85DD95A71FFA6FE67FD95366E0C52" box="[490,576,548,573]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 10</figureCitation>
). A possible indication of ocelli as round subcuticular structures in spm. 2 (
<figureCitation id="DFA85DD95A71FFA6FF12FDF634DC0C0E" box="[159,242,583,608]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 11</figureCitation>
) but not spm. 1. Clypeus without setae. Labral mid-piece present but without tooth. Second maxillary telopodite with a rudimentary pretarsus but without a dorsal brush and very sparsely setose (
<figureCitation id="DFA85DD95A71FFA6FB3EFDDD31290CEA" box="[1203,1287,620,645]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 12</figureCitation>
). No spur on article 2.
</paragraph>
<paragraph id="472C415C5A71FFA6FF4AFD05314E0C81" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="229">
(Forcipules with all parts clearly developed. Poison gland present but without secretory duct). Forcipular coxosternal tooth-plates rounded, bi- or tri-lobed (
<figureCitation id="DFA85DD95A71FFA6FD45FD6637350C9E" box="[712,795,727,752]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 13</figureCitation>
). Forcipular trochanteroprefemoral process present.
</paragraph>
<paragraph id="472C415C5A71FFA6FF4AFD4D37660D32" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="229">
Tergites without sutures or sulci. Only the tergite of the ultimate leg-bearing segment marginate. Sternites without sutures or sulci. sternite of ultimate leg-bearing segment with sides converging posteriorly. Coxopleural process digitiform. No indication of spines or pores (
<figureCitation id="DFA85DD95A71FFA6FD6AFCF537150D32" box="[743,827,836,861]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="472C415C5A71FFA6FF4AFCD931BB0DCB" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="229">
Legs with divided tarsi, a terminal whorl of setae on each article. Claws present but without accessory spines. Ultimate leg with four rows of unsclerotised spines 3,2,1,3 and 3,2,2,3 (
<figureCitation id="DFA85DD95A71FFA6FC3AFC3D30240DCB" box="[951,1034,908,933]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 15</figureCitation>
). Detail of spines shown in
<figureCitation id="DFA85DD95A71FFA6FACDFC3D31BC0DCB" box="[1344,1426,908,933]" captionStart="FIGURES 916" captionStartId="5.[151,264,1734,1756]" captionTargetBox="[282,1300,207,1715]" captionTargetId="figure@5.[281,1310,193,1726]" captionTargetPageId="5" captionText="FIGURES 916. Foetus stadium Otostigmus sulcipes Malacca. 9. Distal four articles showing article 16 subdivided into two beneath the foetal cuticle spm 1. 10. Cephalic plate spm. 1. 11. Detail of cephalic plate showing possible ocelli spm. 2. 12. Pretarsus of second maxillary telopodite spm.1. 13. Forcipular coxosternal tooth-plates spm. 1. 14. Coxopleuron and sternite of ultimate leg-bearing segment spm. 1, ventral view. 15. Ultimate leg prefemur spm. 2. 16. Detail of ultimate leg prefemur showing developing spines. FIGURES 1722. Otostigmus trisulcatus 17. Forcipular coxopleural tooth-plates syntype 13365. 18. Left forcipular trochanteroprefemoral process, the same. 19. Sternite 14, the same. 20. Coxopleuron and sternite of ultimate leg-bearing segment and ultimate leg prefemoral, the same, ventral view (coxal pores not shown). 21. Right coxopleuron syntype ZMB 13365, slide 4002. 22. Ultimate leg prefemur syntype 13365, dorsal view. Scale bars = 0.5 mm except Fig. 12, 13 &amp; 16 = 0.1 mm." httpUri="https://zenodo.org/record/232676/files/figure.png" pageId="4" pageNumber="229">Fig. 16</figureCitation>
.
</paragraph>
<paragraph id="472C415C5A71FFA6FF4AFC1E311B0DA6" blockId="4.[151,1437,151,968]" box="[199,1333,943,968]" pageId="4" pageNumber="229">Spiracles round, on hump-like stigmatopleurite (tracheae seen as weak filaments so must be air-free).</paragraph>
</subSubSection>
</treatment>
</document>