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<document ID-DOI="http://doi.org/10.5281/zenodo.6468403" ID-GBIF-Dataset="93b27db0-ab0a-40d9-862c-1a5c68678186" ID-ISSN="1639-4798" ID-Zenodo-Dep="6468403" checkinTime="1629124948625" checkinUser="carolina" docAuthor="Werff, Henk Van Der" docDate="2013" docId="C71C7428FFF6FFD9F58762A7FD4CCD86" docLanguage="en" docName="Adansonia.35.2.235-279.pdf" docOrigin="Adansonia (3) 35 (2)" docSource="http://dx.doi.org/10.5252/a2013n2a5" docStyle="DocumentStyle:B98EC8A323C0F63E1C55336AD58D9273.2:Adansonia.2011-2014.journal_article" docStyleId="B98EC8A323C0F63E1C55336AD58D9273" docStyleName="Adansonia.2011-2014.journal_article" docStyleVersion="2" docTitle="Ocotea humbertii Kosterm." docType="treatment" docVersion="7" lastPageId="19" lastPageNumber="253" masterDocId="3B250C50FFE4FFCAF5556014FFCBCF4B" masterDocTitle="A revision of the genus Ocotea Aubl. (Lauraceae) in Madagascar and the Comoro Islands" masterLastPageNumber="279" masterPageNumber="235" pageId="18" pageNumber="253" updateTime="1650309971595" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Ocotea Aubl. (Lauraceae) in Madagascar and the Comoro Islands</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Werff, Henk Van Der</mods:namePart>
<mods:affiliation>Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 (USA)</mods:affiliation>
<mods:nameIdentifier type="email">henk.vanderwerff@mobot.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Adansonia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="series">
<mods:title>3</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2013-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>35</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>235</mods:start>
<mods:end>279</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.5252/a2013n2a5</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/a2013n2a5</mods:identifier>
<mods:identifier type="GBIF-Dataset">93b27db0-ab0a-40d9-862c-1a5c68678186</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
<mods:identifier type="Zenodo-Dep">5206484</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6468403" ID-Zenodo-Dep="6468403" LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86" httpUri="http://treatment.plazi.org/id/C71C7428FFF6FFD9F58762A7FD4CCD86" lastPageId="19" lastPageNumber="253" pageId="18" pageNumber="253">
<subSubSection box="[210,550,690,718]" pageId="18" pageNumber="253" type="nomenclature">
<paragraph blockId="18.[210,550,690,749]" box="[210,550,690,718]" pageId="18" pageNumber="253">
<heading box="[210,550,690,718]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="253" reason="3">
15.
<taxonomicName authority="Kosterm." authorityName="Kosterm." box="[252,550,690,718]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="253" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis bold="true" box="[252,443,690,717]" italics="true" pageId="18" pageNumber="253">Ocotea humbertii</emphasis>
Kosterm.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[340,420,722,749]" pageId="18" pageNumber="253" type="description">
<paragraph blockId="18.[210,550,690,749]" box="[340,420,722,749]" pageId="18" pageNumber="253">
(
<figureCitation box="[347,411,722,749]" captionStart="FIG" captionStartId="16.[677,688,1208,1226]" captionTargetBox="[677,1211,215,1158]" captionTargetId="figure-434@16.[677,1210,215,1158]" captionTargetPageId="16" captionText="FIG. 5. — Distribution of Ocotea faucherei (Danguy) Kosterm. (N), O. humbertii Kosterm. (p), O. humblotii Baill. (l), O. involuta Kosterm. (Ì) and O. zahamenensis van der Werff, sp. nov. («)." figureDoi="http://doi.org/10.5281/zenodo.5206494" httpUri="https://zenodo.org/record/5206494/files/figure.png" pageId="18" pageNumber="253">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[136,465,780,804]" pageId="18" pageNumber="253" type="reference_group">
<paragraph blockId="18.[113,648,780,1206]" box="[136,465,780,804]" pageId="18" pageNumber="253">
<bibRefCitation author="KOSTERMANS A. J. G. H." box="[136,461,780,804]" journalOrPublisher="Notulae systematicae" pageId="18" pageNumber="253" pagination="67 - 128" part="8" refId="ref26754" refString="KOSTERMANS A. J. G. H. 1939. - Enumeratio Lauracearum Madagascariensium et ex Insulis Mascarenis. (Revisio Lauracearum VI). Notulae systematicae 8 (2): 67 - 128." title="Enumeratio Lauracearum Madagascariensium et ex Insulis Mascarenis. (Revisio Lauracearum VI)" type="journal article" year="1939">
<emphasis box="[136,328,780,804]" italics="true" pageId="18" pageNumber="253">Notulae systematicae</emphasis>
8: 86 (1939)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="253" type="materials_examined">
<paragraph blockId="18.[113,648,780,1206]" pageId="18" pageNumber="253">
<materialsCitation collectingDate="1934-01" collectionCode="P" collectorName="Humbert" country="Madagascar" elevation="1850" location="Massif de l'Andohahela" pageId="18" pageNumber="253" specimenCode="P00541569" specimenCount="1" typeStatus="holotype">
TYPUS. —
<emphasis bold="true" box="[235,363,827,851]" pageId="18" pageNumber="253">
<collectingCountry box="[235,363,827,851]" name="Madagascar" pageId="18" pageNumber="253">Madagascar</collectingCountry>
</emphasis>
.
<location LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86:4A6A93E5FFF6FFD8F42B632FFD4ACC18" box="[382,641,827,851]" country="Madagascar" name="Massif de l'Andohahela" pageId="18" pageNumber="253">Massif de l’Andohahela</location>
,
<quantity box="[113,261,854,878]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="18" pageNumber="253" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">
<elevation box="[113,261,854,878]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="18" pageNumber="253" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800-1900 m</elevation>
</quantity>
,
<date box="[275,347,854,878]" pageId="18" pageNumber="253" value="1934-01">
<collectingDate box="[275,347,854,878]" pageId="18" pageNumber="253" value="1934-01">I.1934</collectingDate>
</date>
, fls,
<emphasis box="[400,567,854,878]" italics="true" pageId="18" pageNumber="253">
<collectorName box="[400,491,854,878]" pageId="18" pageNumber="253">Humbert</collectorName>
13613
</emphasis>
(
<typeStatus box="[583,642,854,878]" pageId="18" pageNumber="253">holo-</typeStatus>
,
<collectionCode box="[115,130,881,904]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="18" pageNumber="253" type="Herbarium">P</collectionCode>
[
<specimenCode box="[136,246,880,904]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="18" pageNumber="253" type="Herbarium">P00541569</specimenCode>
]!
</materialsCitation>
;
<materialsCitation box="[269,642,880,904]" collectingDate="1934-01" collectionCode="MO, P" collectorName="Humbert" country="Madagascar" elevation="1850" location="Massif de l'Andohahela" pageId="18" pageNumber="253" specimenCode="P00541570, P00541571" specimenCount="1" typeStatus="isotype">
<typeStatus box="[269,305,880,904]" pageId="18" pageNumber="253">iso-</typeStatus>
,
<collectionCode box="[313,359,880,904]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="18" pageNumber="253" type="Herbarium">MO</collectionCode>
!,
<collectionCode box="[370,385,881,904]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="18" pageNumber="253" type="Herbarium">P</collectionCode>
[
<specimenCode box="[391,501,880,904]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="18" pageNumber="253" type="Herbarium">P00541570</specimenCode>
,
<specimenCode box="[509,620,880,904]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="18" pageNumber="253" type="Herbarium">P00541571</specimenCode>
]!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[113,648,780,1206]" pageId="18" pageNumber="253">
SPECIMENS EXAMINED. —
<materialsCitation collectionCode="P" collectorName="Humbert" location="Massif de l'Andohahela" pageId="18" pageNumber="253" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86:4A6A93E5FFF6FFD8F43263B4FD8DCCF3" box="[359,582,928,952]" name="Massif de l'Andohahela" pageId="18" pageNumber="253">Massif de l’Andohahela</location>
,
<emphasis italics="true" pageId="18" pageNumber="253">
<collectorName pageId="18" pageNumber="253">Humbert</collectorName>
13613
</emphasis>
(
<collectionCode box="[223,238,954,978]" pageId="18" pageNumber="253">P</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectionCode="P" location="Rakotomalaza" municipality="Andohahela" pageId="18" pageNumber="253" specimenCode="P01991759" specimenCount="1" stateProvince="Toliara">
<collectingRegion box="[252,318,954,978]" country="Madagascar" name="Toliara" pageId="18" pageNumber="253">Toliara</collectingRegion>
, RNI 11
<collectingMunicipality box="[400,513,954,978]" pageId="18" pageNumber="253">Andohahela</collectingMunicipality>
,
<emphasis italics="true" pageId="18" pageNumber="253">
<location LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86:4A6A93E5FFF6FFD8F75D63AEFD4CCC99" box="[520,647,954,978]" municipality="Andohahela" name="Rakotomalaza" pageId="18" pageNumber="253" stateProvince="Toliara">Rakotomalaza</location>
555
</emphasis>
(MO, P[
<specimenCode box="[246,362,981,1005]" pageId="18" pageNumber="253">P01991759</specimenCode>
])
</materialsCitation>
;
<materialsCitation collectionCode="P" location="Ivahona" municipality="Betroka" pageId="18" pageNumber="253" specimenCode="P06241566" specimenCount="1" stateProvince="Toliara">
<collectingRegion box="[391,462,981,1005]" country="Madagascar" name="Toliara" pageId="18" pageNumber="253">Toliara</collectingRegion>
.
<collectingMunicipality box="[472,551,981,1005]" pageId="18" pageNumber="253">Betroka</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86:4A6A93E5FFF6FFD8F76763C1FD48CCA6" box="[562,643,981,1005]" municipality="Betroka" name="Ivahona" pageId="18" pageNumber="253" stateProvince="Toliara">Ivahona</location>
,
<location LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86:4A6A93E5FFF6FFD8F52663FBFF1FCB4C" box="[115,212,1007,1031]" municipality="Betroka" name="Befarafara" pageId="18" pageNumber="253" stateProvince="Toliara">Befarafara</location>
,
<emphasis box="[218,427,1007,1031]" italics="true" pageId="18" pageNumber="253">
<location LSID="urn:lsid:plazi:treatment:C71C7428FFF6FFD9F58762A7FD4CCD86:4A6A93E5FFF6FFD8F58F63FBFE8CCB4C" box="[218,327,1007,1031]" municipality="Betroka" name="Andrianjafy" pageId="18" pageNumber="253" stateProvince="Toliara">Andrianjafy</location>
et al. 1093
</emphasis>
(MO, P[
<specimenCode box="[513,625,1007,1031]" pageId="18" pageNumber="253">
P
<date box="[526,625,1007,1031]" pageId="18" pageNumber="253" value="1566-06-24">06241566</date>
</specimenCode>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="253" type="distribution">
<paragraph blockId="18.[113,648,780,1206]" pageId="18" pageNumber="253">
DISTRIBUTION. — This species is known from two collections made in the Massif of Andohahela at
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="18" pageNumber="253" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800- 1900 m</quantity>
elevation and one collection near Betroka at
<quantity box="[115,192,1134,1158]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="18" pageNumber="253" unit="m" value="1360.0">1360 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection box="[113,546,1182,1206]" pageId="18" pageNumber="253" type="biology_ecology">
<paragraph blockId="18.[113,648,780,1206]" box="[113,546,1182,1206]" pageId="18" pageNumber="253">PHENOLOGY. — Flowers and fruits: January.</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5206496" ID-Zenodo-Dep="5206496" httpUri="https://zenodo.org/record/5206496/files/figure.png" pageId="18" pageNumber="253" startId="18.[677,688,1197,1215]" targetBox="[677,1211,215,1155]" targetPageId="18">
<paragraph blockId="18.[677,1210,1196,1260]" pageId="18" pageNumber="253">
FIG. 6. — Distribution of
<taxonomicName authority="Kosterm." authorityName="Kosterm." box="[891,1117,1196,1215]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="253" phylum="Tracheophyta" rank="species" species="brevipes">
<emphasis box="[891,1031,1196,1215]" italics="true" pageId="18" pageNumber="253">Ocotea brevipes</emphasis>
Kosterm.
</taxonomicName>
(
<emphasis bold="true" box="[1127,1141,1196,1214]" pageId="18" pageNumber="253">N</emphasis>
),
<emphasis italics="true" pageId="18" pageNumber="253">O. foveolata</emphasis>
Kosterm. (
<emphasis bold="true" box="[830,848,1219,1236]" italics="true" pageId="18" pageNumber="253">p</emphasis>
),
<taxonomicName authority="van der Werff" authorityName="van der Werff" box="[865,1141,1219,1237]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="253" phylum="Tracheophyta" rank="species" species="longipedicellata">
<emphasis box="[865,1021,1219,1237]" italics="true" pageId="18" pageNumber="253">O. longipedicellata</emphasis>
van der Werff
</taxonomicName>
(
<emphasis bold="true" box="[1153,1167,1219,1237]" pageId="18" pageNumber="253">l</emphasis>
) and
<taxonomicName authority="Kosterm." authorityName="Kosterm." box="[677,868,1242,1260]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="253" phylum="Tracheophyta" rank="species" species="sessiliflora">
<emphasis box="[677,787,1242,1260]" italics="true" pageId="18" pageNumber="253">O. sessiliflora</emphasis>
Kosterm.
</taxonomicName>
(
<emphasis bold="true" box="[878,895,1242,1259]" pageId="18" pageNumber="253">Ì</emphasis>
).
</paragraph>
</caption>
<subSubSection pageId="18" pageNumber="253" type="description">
<paragraph blockId="18.[111,648,1262,1700]" box="[113,258,1262,1288]" pageId="18" pageNumber="253">DESCRIPTION</paragraph>
<paragraph blockId="18.[111,648,1262,1700]" lastBlockId="18.[675,1211,1325,1573]" pageId="18" pageNumber="253">
Tree, to
<quantity box="[200,255,1293,1319]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="18" pageNumber="253" unit="m" value="18.0">18 m</quantity>
tall. Twigs angular, when very young brown-tomentellous, but soon glabrescent; terminal buds brown-tomentellous. Leaves alternate, firmly chartaceous, 5.5-9 ×
<quantity box="[346,472,1388,1415]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="18" pageNumber="253" unit="cm" value="2.9" valueMax="3.5" valueMin="2.3">2.3-3.5 cm</quantity>
, base acute, tip obtuse or acute, midrib slightly raised, lateral veins immersed on both surfaces, fine reticulation raised on lower surface, very slightly raised on upper surface, glabrous on both surfaces except for some curled reddish hairs along midrib on lower surface, lateral veins not very prominent, 5-6 on each side; domatia present, as deep pockets or cavities with a slit-like opening. Petioles
<quantity box="[301,388,1641,1668]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="18" pageNumber="253" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6-8 mm</quantity>
long, flat on upper side, with similar indument as twigs. Inflorescences in axils of deciduous bracts near tips of twigs, glabrous or pubescent, to
<quantity box="[849,901,1356,1383]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="253" unit="cm" value="2.0">2 cm</quantity>
long, few-flowered.Tepals 6, glabrous outside, pubescent inside,
<emphasis box="[1044,1061,1389,1414]" italics="true" pageId="18" pageNumber="253">c.</emphasis>
<quantity box="[1067,1151,1388,1415]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="253" unit="mm" value="1.5">1.5 mm</quantity>
long. Stamens 9, 4-celled, filaments with a few hairs, anthers glabrous, staminodia 3, stipitiform with a few hairs. Ovary and inside of receptacle glabrous. Cupule bowl-shaped, shallow,
<quantity box="[995,1068,1515,1541]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="253" unit="cm" value="1.3">1.3 cm</quantity>
wide,
<quantity box="[1136,1209,1514,1541]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="253" unit="cm" value="0.5">0.5 cm</quantity>
high, fruit ellipsoid,
<emphasis box="[896,913,1547,1572]" italics="true" pageId="18" pageNumber="253">c.</emphasis>
2 ×
<quantity box="[962,1041,1546,1573]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="253" unit="cm" value="1.5">1.5 cm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="254" pageId="18" pageNumber="253" type="discussion">
<paragraph blockId="18.[677,1209,1610,1700]" box="[677,777,1610,1636]" pageId="18" pageNumber="253">REMARKS</paragraph>
<paragraph blockId="18.[677,1209,1610,1700]" lastBlockId="19.[111,649,216,718]" lastPageId="19" lastPageNumber="254" pageId="18" pageNumber="253">
<taxonomicName box="[677,858,1641,1668]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="253" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis box="[677,858,1641,1668]" italics="true" pageId="18" pageNumber="253">Ocotea humbertii</emphasis>
</taxonomicName>
is readily recognized by the erect hairs on young twigs and terminal buds, the very short inflorescences, and the leaves with rather deep pocket-domatia. The
<typeStatus box="[330,375,248,274]" pageId="19" pageNumber="254">type</typeStatus>
collection has stunted and glabrous inflorescences, and the second collection has better developed, pubescent inflorescences. In other characters, such as indument on twigs and flowers, domatia
<typeStatus box="[346,395,375,401]" pageId="19" pageNumber="254">type</typeStatus>
, they are a good match.
<bibRefCitation author="KOSTERMANS A. J. G. H." box="[113,326,406,433]" firstAuthor="Kostermans" journalOrPublisher="Communication (Pengumuman) of the Forest Research Institute" pageId="19" pageNumber="254" pagination="1 - 44" part="57" refId="ref26822" refString="KOSTERMANS A. J. G. H. 1957. - Le genre Ocotea Aubl. (Lauracees) a Madagascar. Communication (Pengumuman) of the Forest Research Institute 57: 1 - 44." title="Le genre Ocotea Aubl. (Lauracees) a Madagascar" type="journal article" year="1957">Kostermans (1957)</bibRefCitation>
cited three other collections under
<taxonomicName box="[190,339,437,464]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="19" pageNumber="254" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis box="[190,339,437,464]" italics="true" pageId="19" pageNumber="254">O. humbertii</emphasis>
</taxonomicName>
; he already suggested that these could belong to a different species. I exclude these collections from
<taxonomicName box="[355,493,501,527]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="19" pageNumber="254" phylum="Tracheophyta" rank="species" species="humbertii">
<emphasis box="[355,493,501,527]" italics="true" pageId="19" pageNumber="254">O. humbertii</emphasis>
</taxonomicName>
, they differ in their appressed pubescence on the terminal buds, absence of tomentellous indument on young twigs and their distribution.
<emphasis box="[360,571,596,622]" italics="true" pageId="19" pageNumber="254">Rakotomalaza 1082</emphasis>
(MO) is provisionally placed here; its domatia are rather shallow and the leaves wider, but it has the typical indument of erect, short hairs on the young twigs.
</paragraph>
</subSubSection>
</treatment>
</document>
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