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<document ID-DOI="http://doi.org/10.5281/zenodo.6115602" ID-GBIF-Dataset="85cd7166-7b23-496d-8b10-7f1dcf2c4d60" ID-GBIF-Taxon="114920415" ID-ISSN="1175-5326" ID-Zenodo-Dep="6115602" ID-ZooBank="A625A5A9-EE80-45E0-A9BE-7A183B0996B1" checkinTime="1447441568197" checkinUser="donat" docAuthor="Robert A. Johnson" docDate="2015" docId="971D8786FF8EFFD665D41483D0BB2497" docLanguage="en" docName="29902.pdf.imf" docOrigin="Zootaxa 4029 (1)" docTitle="Pogonomyrmex abdominalis" docType="treatment" docVersion="5" lastPageId="110" lastPageNumber="111" masterDocId="6B24FFFEFFE9FFB865431775D1342613" masterDocTitle="A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part I" masterLastPageNumber="142" masterPageNumber="1" pageId="103" pageNumber="104" updateTime="1645066743641" updateUser="ExternalLinkService">
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<mods:title>A taxonomic revision of South American species of the seed-harvester ant genus Pogonomyrmex (Hymenoptera: Formicidae). Part I</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Robert A. Johnson</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>4029</mods:number>
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<mods:number>1</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.4029.1.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="ZooBank">A625A5A9-EE80-45E0-A9BE-7A183B0996B1</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6115602" ID-GBIF-Taxon="114920415" ID-Zenodo-Dep="6115602" LSID="urn:lsid:plazi:treatment:971D8786FF8EFFD665D41483D0BB2497" httpUri="http://treatment.plazi.org/id/971D8786FF8EFFD665D41483D0BB2497" lastPageId="110" lastPageNumber="111" pageId="103" pageNumber="104">
<subSubSection box="[151,497,1014,1040]" pageId="103" pageNumber="104" type="nomenclature">
<paragraph box="[151,497,1014,1040]" pageId="103" pageNumber="104">
<emphasis bold="true" box="[151,497,1014,1040]" italics="true" pageId="103" pageNumber="104">
<heading bold="true" box="[151,497,1014,1040]" fontSize="11" level="1" pageId="103" pageNumber="104" reason="1">
<taxonomicName box="[151,497,1014,1040]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="abdominalis">Pogonomyrmex abdominalis</taxonomicName>
</heading>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[151,330,1049,1074]" pageId="103" pageNumber="104" type="description">
<paragraph box="[151,330,1049,1074]" pageId="103" pageNumber="104">
(
<figureCitation box="[159,322,1049,1074]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStart-2="FIGURE 55" captionStartId-0="105.[151,249,1978,2000]" captionStartId-1="106.[151,249,1637,1659]" captionStartId-2="107.[151,249,1978,2000]" captionTargetBox-0="[412,1171,193,1956]" captionTargetBox-1="[314,1263,193,1606]" captionTargetBox-2="[395,1187,193,1956]" captionTargetId-0="figure@105.[412,1175,193,1956]" captionTargetId-1="figure@106.[314,1273,193,1615]" captionTargetId-2="figure@107.[395,1192,193,1956]" captionTargetPageId-0="105" captionTargetPageId-1="106" captionTargetPageId-2="107" captionText-0="FIGURE 53. Photograph of Pogonomyrmex abdominalis Santschi worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0235283). Photographs by Will Ericson from www. AntWeb. org." captionText-1="FIGURE 54. Photograph from posterior margin of head looking down onto lateral clypeal lobes: (A) Pogonomyrmex naegelii Forel—black arrows show small lobe that projects dorsally from anterior margin of antennal fossa (CASENT 0914811), and (B) Pogonomyrmex abdominalis Santschi—red arrows show absence of small lobe on anterior margin of antennal fossa (CASENT 0914813). Photographs by Michele Esposito from www. AntWeb. org." captionText-2="FIGURE 55. Photograph of Pogonomyrmex abdominalis Santschi alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0914378). Photographs by Michele Esposito from www. AntWeb. org." pageId="103" pageNumber="104">Figures 5355</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="reference_group">
<paragraph pageId="103" pageNumber="104">
<taxonomicName box="[151,577,1114,1136]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="naegelii" subGenus="Ephebomyrmex">
<emphasis box="[151,310,1115,1136]" italics="true" pageId="103" pageNumber="104">Pogonomyrmex</emphasis>
(
<emphasis box="[324,481,1114,1136]" italics="true" pageId="103" pageNumber="104">Ephebomyrmex</emphasis>
)
<emphasis box="[495,577,1114,1136]" italics="true" pageId="103" pageNumber="104">naegelii</emphasis>
</taxonomicName>
st.
<taxonomicName authority="Santschi, 1929: 278" authorityName="Santschi" authorityPageNumber="278" authorityYear="1929" box="[612,942,1114,1136]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[612,737,1114,1136]" italics="true" pageId="103" pageNumber="104">abdominalis</emphasis>
Santschi, 1929: 278
</taxonomicName>
(worker, queen). Syntypes examined: 8 workers, 4 queens [MACN], ARGENTINA, Córdoba: Sierras de Córdoba, Alta Gracia La Granja, #1710 (Charles Bruch leg.). See also Gallardo, 1932: 112, figs. 6, 7. MACN worker here designated
<emphasis bold="true" box="[887,1043,1176,1198]" pageId="103" pageNumber="104">LECTOTYPE</emphasis>
[CASENT0235285].
</paragraph>
<paragraph box="[151,1127,1206,1229]" pageId="103" pageNumber="104">
<taxonomicName authority="Santschi" authorityName="Santschi" box="[151,718,1206,1229]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="abdominalis" subGenus="Ephebomyrmex">
<emphasis box="[151,310,1207,1228]" italics="true" pageId="103" pageNumber="104">Pogonomyrmex</emphasis>
(
<emphasis box="[324,482,1206,1229]" italics="true" pageId="103" pageNumber="104">Ephebomyrmex</emphasis>
)
<emphasis box="[497,622,1206,1229]" italics="true" pageId="103" pageNumber="104">abdominalis</emphasis>
Santschi
</taxonomicName>
: Kusnezov, 1951: 250, raised to species.
</paragraph>
<paragraph box="[151,1129,1237,1259]" pageId="103" pageNumber="104">
<emphasis box="[151,440,1237,1259]" italics="true" pageId="103" pageNumber="104">
Ephebomyrmex
<taxonomicName authority="Santschi" authorityName="Santschi" box="[315,553,1237,1259]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="abdominalis">abdominalis (Santschi)</taxonomicName>
</emphasis>
: Kempf, 1972: 106, first combination in
<emphasis box="[965,1123,1237,1259]" italics="true" pageId="103" pageNumber="104">Ephebomyrmex</emphasis>
.
</paragraph>
<paragraph box="[151,1150,1267,1290]" pageId="103" pageNumber="104">
<taxonomicName authority="Santschi" authorityName="Santschi" box="[151,537,1267,1290]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[151,441,1267,1290]" italics="true" pageId="103" pageNumber="104">Pogonomyrmex abdominalis</emphasis>
Santschi
</taxonomicName>
: Bolton, 1995: 339, revived combination in
<emphasis box="[984,1150,1269,1290]" italics="true" pageId="103" pageNumber="104">
<taxonomicName box="[984,1146,1269,1290]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="genus">Pogonomyrmex</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="103" pageNumber="104" type="diagnosis">
<paragraph pageId="103" pageNumber="104">
<emphasis bold="true" box="[151,378,1334,1359]" pageId="103" pageNumber="104">Worker. Diagnosis.</emphasis>
Within the
<emphasis box="[510,628,1334,1358]" italics="true" pageId="103" pageNumber="104">
<taxonomicName box="[510,628,1334,1358]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="naegelii">P. naegelii</taxonomicName>
</emphasis>
-group, the combination of: (1) approximately 810 coarse, longitudinal rugae between frontal lobes, (2) lacking small lobe that projects dorsally from anterior margin of antennal fossa, (3) peduncle of petiole and anterior surface of petiolar node meet at or near a right angle, (4) in dorsal view, posterior surface of petiolar node distinctly wider than distance between tips of superior propodeal spines, and (5) longest hairs on mesosoma approaching to slightly&gt;MOD uniquely characterize this species (
<emphasis bold="true" box="[1117,1284,1476,1501]" pageId="103" pageNumber="104">
<figureCitation box="[1117,1284,1476,1501]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStartId-0="105.[151,249,1978,2000]" captionStartId-1="106.[151,249,1637,1659]" captionTargetBox-0="[412,1171,193,1956]" captionTargetBox-1="[314,1263,193,1606]" captionTargetId-0="figure@105.[412,1175,193,1956]" captionTargetId-1="figure@106.[314,1273,193,1615]" captionTargetPageId-0="105" captionTargetPageId-1="106" captionText-0="FIGURE 53. Photograph of Pogonomyrmex abdominalis Santschi worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0235283). Photographs by Will Ericson from www. AntWeb. org." captionText-1="FIGURE 54. Photograph from posterior margin of head looking down onto lateral clypeal lobes: (A) Pogonomyrmex naegelii Forel—black arrows show small lobe that projects dorsally from anterior margin of antennal fossa (CASENT 0914811), and (B) Pogonomyrmex abdominalis Santschi—red arrows show absence of small lobe on anterior margin of antennal fossa (CASENT 0914813). Photographs by Michele Esposito from www. AntWeb. org." pageId="103" pageNumber="104">Figures 5354</figureCitation>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104" type="description">
<paragraph pageId="103" pageNumber="104">
<emphasis bold="true" box="[199,374,1512,1536]" pageId="103" pageNumber="104">Measurements</emphasis>
—lectotype (
<emphasis box="[525,539,1512,1536]" italics="true" pageId="103" pageNumber="104">n</emphasis>
= 31). HL 1.25 (1.171.41); HW 1.26 (1.141.33); MOD 0.26 (0.250.30); OMD 0.28 (0.240.33); SL 0.84 (0.870.99); PNW 0.82 (0.750.92); HFL 1.09 (1.031.27); ML 1.38 (1.331.55); PW 0.38 (0.340.43); PPW 0.49 (0.460.54). Indices: SI 66.67 (70.0885.09); CI 100.80 (92.37105.69); OI 20.63 (20.4924.56); HFI 86.51 (83.46103.31).
</paragraph>
<paragraph lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104">
<emphasis bold="true" box="[199,365,1655,1680]" pageId="103" pageNumber="104">Redescription</emphasis>
. Head subquadrate to quadrate (CI = 92.37105.69), widest just posterior to eye; posterior margin flat to weakly concave. Longitudinal rugae on cephalic dorsum prominent, occasionally weakly rugoreticulate especially near posterior margin; approximately 810 coarse, longitudinal rugae between frontal lobes; in full-face view, medial rugae not diverging toward posterior corners of head. In profile, area posterior to eyes rugose to moderately rugoreticulate, rugae not converging toward vertex. Cephalic interrugae moderately to strongly granulate, dull to weakly shining; vertex rugose. Anterior margin of clypeus moderately to strongly concave, dorsal surface with numerous subparallel, longitudinal rugae; lacking small lobe that projects dorsally from anterior margin of antennal fossa. Mandible with six teeth; mandibular dorsum coarsely rugose. Up to several moderately long, curved, bristle-like, yellow-brown to brownish hairs project from anterior margin of clypeus and basolateral margin of mandibles. MOD ranging from 0.200.25x HL. In profile, eyes situated anterior to middle of head, OMD = 0.801.19x MOD. Antennal scapes moderately long (SI = 66.6785.09), failing to reach vertex by 1.01.5x length of basal funicular segment; entire scape with strong longitudinal striae, dull to weakly shining; basal flange of scape flattened and well-developed with carinate margin. Psammophore poorly-developed, consisting of short to medium-length hairs scattered across ventral side of head.
</paragraph>
<paragraph pageId="104" pageNumber="105">Mesosomal profile convex; all mesosomal surfaces with highly irregular rugae, rugoreticulate, or vermiculate. Mesoepinotal sulcus not impressed. Dorsum of promesonotum and sides of pronotum rugoreticulate to vermiculate. Mesopleura with irregular rugae angling posterodorsally to rugoreticulate. Dorsum of propodeum with wavy to irregular transverse rugae that traverse anteroventrally, becoming more irregular on sides. Superior propodeal spines moderately long, acuminate, connected by well-defined keel; spine length approximately 0.7 0.8x distance between their bases. Inferior propodeal spines well-developed, acuminate, length approximately 0.5 1.0x that of superior spines, base wider than length of superior spines; inferior and superior spines connected by weak crest. Propodeal spiracles ovoid to circular facing posterad. Interrugae on mesosoma moderately granulate, weakly shining to smooth and strongly shining. Legs moderately coriarious, weakly shining.</paragraph>
<paragraph pageId="104" pageNumber="105">Peduncle of petiole about as long as petiolar node, anteroventral margin with a weakly to strongly-developed triangular process. In profile, petiolar node asymmetrical with anterior surface shorter than posterior surface; apex weakly rounded to subangulate; anterior surface meeting peduncle at or near a right angle. In dorsal view, petiolar node longer than wide, widest near middle, narrowing to spatulate to rounded anterior margin; maximal width of posterior surface greater than distance between tips of superior propodeal spines; dorsum and sides strongly rugoreticulate to vermiculate, interrugae granulate, dull to weakly shining. Dorsum of postpetiole convex in profile; in dorsal view, postpetiole robust, widest at or near posterior margin, narrowing from near middle to weakly truncate anterior margin; maximal width slightly greater than length; dorsum and sides weakly to moderately rugoreticulate or with several irregular longitudinal to oblique rugae, interrugae moderately to strongly granulate, dull to weakly shining. Ventral process of postpetiole large, bulbous, height similar to dorsal portion of postpetiole. First gastral tergum weakly to strongly coriarious, dull to weakly shining to smooth and strongly shining; anterior margin sometimes with weak longitudinal striae.</paragraph>
<paragraph pageId="104" pageNumber="105">
Erect yellow-brown to brownish pilosity moderately abundant on head, variable in length, mostly short to medium-length, often with one or more longer hairs that approximate MOD. Moderately abundant suberect to semidecumbent pilosity on scape, abundant decumbent hairs on funicular segments. Legs with moderately abundant semidecumbent brownish setae. Mesosoma, petiolar node, postpetiole, gastral terga with moderately dense, erect setae, mostly similar in length, longest approaching to slightly&gt;MOD; hairs on propodeum less dense. Concolorous tan to tannish-brown with darker gaster (
<figureCitation box="[758,925,1220,1245]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStartId-0="105.[151,249,1978,2000]" captionStartId-1="106.[151,249,1637,1659]" captionTargetBox-0="[412,1171,193,1956]" captionTargetBox-1="[314,1263,193,1606]" captionTargetId-0="figure@105.[412,1175,193,1956]" captionTargetId-1="figure@106.[314,1273,193,1615]" captionTargetPageId-0="105" captionTargetPageId-1="106" captionText-0="FIGURE 53. Photograph of Pogonomyrmex abdominalis Santschi worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0235283). Photographs by Will Ericson from www. AntWeb. org." captionText-1="FIGURE 54. Photograph from posterior margin of head looking down onto lateral clypeal lobes: (A) Pogonomyrmex naegelii Forel—black arrows show small lobe that projects dorsally from anterior margin of antennal fossa (CASENT 0914811), and (B) Pogonomyrmex abdominalis Santschi—red arrows show absence of small lobe on anterior margin of antennal fossa (CASENT 0914813). Photographs by Michele Esposito from www. AntWeb. org." pageId="104" pageNumber="105">
<emphasis bold="true" box="[758,925,1220,1245]" pageId="104" pageNumber="105">Figures 5354</emphasis>
</figureCitation>
).
</paragraph>
<paragraph pageId="104" pageNumber="105">
<emphasis bold="true" box="[199,412,1256,1281]" pageId="104" pageNumber="105">Queen. Diagnosis.</emphasis>
This caste is diagnosed by: (1) caste-specific morphology of the mesosoma related to wingbearing and presence of ocelli on head, (2) small (HW &lt;1.35 mm), (3) petiolar node and postpetiole wide (petiolar node&gt; 0.50 mm, postpetiole&gt; 0.70 mm), (4) posterior surface of petiolar node granulate-punctate, and (5) longitudinal rugae on mesoscutum and mesoscutellum (
<figureCitation box="[776,890,1364,1389]" captionStart="FIGURE 55" captionStartId="107.[151,249,1978,2000]" captionTargetBox="[395,1187,193,1956]" captionTargetId="figure@107.[395,1192,193,1956]" captionTargetPageId="107" captionText="FIGURE 55. Photograph of Pogonomyrmex abdominalis Santschi alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT 0914378). Photographs by Michele Esposito from www. AntWeb. org." pageId="104" pageNumber="105">
<emphasis bold="true" box="[776,890,1364,1389]" pageId="104" pageNumber="105">Figure 55</emphasis>
</figureCitation>
).
</paragraph>
<paragraph pageId="104" pageNumber="105">
<emphasis bold="true" box="[199,374,1401,1425]" pageId="104" pageNumber="105">Measurements</emphasis>
—(
<emphasis box="[411,425,1401,1425]" italics="true" pageId="104" pageNumber="105">n</emphasis>
= 4). HL 1.191.35; HW 1.241.30; MOD 0.300.32; OMD 0.270.31; SL 0.690.75; PNW 1.001.03; HFL 0.920.95; ML 1.561.65; PW0.540.57; PPW 0.710.76. Indices: SI 54.3357.69; CI 95.38106.72; OI 23.0825.20; HFI 72.4476.61.
</paragraph>
<paragraph box="[199,392,1508,1533]" pageId="104" pageNumber="105">
<emphasis bold="true" box="[199,267,1509,1533]" pageId="104" pageNumber="105">Male.</emphasis>
<taxonomicName box="[273,388,1508,1533]" family="Polemoniaceae" genus="Unknown" higherTaxonomySource="IPNI" kingdom="Pantae" order="Polemoniales" pageId="104" pageNumber="118" rank="genus">Unknown</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="materials_examined">
<paragraph pageId="104" pageNumber="105">
<emphasis bold="true" box="[199,857,1544,1569]" pageId="104" pageNumber="105">
Additional material examined. ARGENTINA:
<emphasis bold="true" box="[758,857,1544,1569]" italics="true" pageId="104" pageNumber="105">Córdoba</emphasis>
</emphasis>
: Alta Gracia la Granja, Sierras de Córdoba, no date (MACN); Estancia Santo Domingo, 24 Feb, 2009 (RAJC; RGPC).
<emphasis bold="true" box="[906,1030,1580,1605]" italics="true" pageId="104" pageNumber="105">Entre Ríos</emphasis>
: Parque Nacional El Palmar, Jan 5 6, 2006 (RGPC); El Palmar, 25 m, Feb 7, 1999 (RAJC); Victoria, 80, Jan 6, 2008 &amp; Jan 12, 2011 (FML; MCZ; RAJC).
<emphasis bold="true" box="[246,351,1652,1677]" italics="true" pageId="104" pageNumber="105">Misiones</emphasis>
: Loreto, no date (MACN); Estación Experimental Loreto, no date (FML).
<emphasis bold="true" box="[1232,1428,1652,1677]" pageId="104" pageNumber="105">
BOLIVIA:
<emphasis bold="true" box="[1373,1428,1652,1677]" italics="true" pageId="104" pageNumber="105">Beni</emphasis>
</emphasis>
: Reyes, Nov 22, 1921 (USNM).
<emphasis bold="true" box="[514,699,1688,1713]" italics="true" pageId="104" pageNumber="105">
Dpto.
<taxonomicName box="[586,699,1688,1713]" family="Polemoniaceae" genus="Unknown" higherTaxonomySource="IPNI" kingdom="Pantae" order="Polemoniales" pageId="104" pageNumber="105" rank="genus">Unknown</taxonomicName>
</emphasis>
: San Antonio, Nov 22, 1921 (USNM).
<emphasis bold="true" box="[1153,1428,1688,1713]" pageId="104" pageNumber="105">
BRAZIL:
<emphasis bold="true" box="[1278,1428,1688,1713]" italics="true" pageId="104" pageNumber="105">Mato Grosso</emphasis>
</emphasis>
: Belo Horizonte, Nov 4, no year (CASC; MCZ).
<emphasis bold="true" box="[716,875,1724,1749]" italics="true" pageId="104" pageNumber="105">Minas Gerais</emphasis>
: 6 km E Mariana Mtns, 1000 m, May 16, 1971 (MCZ).
<emphasis bold="true" box="[244,457,1760,1785]" italics="true" pageId="104" pageNumber="105">Río Grande do Sul</emphasis>
: Porto Alegre, Mar 24, 1971 (MZUSP).
<emphasis bold="true" box="[909,1028,1760,1785]" italics="true" pageId="104" pageNumber="105">São Paulo</emphasis>
: Fazanda Itaquera, Nova Europa, no date (MZUSP); Pindamonhangaba, Aug 25, 1961 (MZUSP).
<emphasis bold="true" box="[876,1081,1796,1821]" italics="true" pageId="104" pageNumber="105">
Estado
<taxonomicName authority="Pirafron" authorityName="Pirafron" box="[967,1197,1796,1821]" family="Polemoniaceae" genus="Unknown" higherTaxonomySource="IPNI" kingdom="Pantae" order="Polemoniales" pageId="104" pageNumber="105" rank="genus">Unknown: Pirafron</taxonomicName>
</emphasis>
?, no date (MZUSP) (
<emphasis bold="true" box="[160,294,1832,1857]" pageId="104" pageNumber="105">
<figureCitation box="[160,273,1832,1857]" captionStart="FIGURE 56" captionStartId="108.[151,249,1946,1968]" captionTargetBox="[420,1168,183,1922]" captionTargetId="figure@108.[420,1176,183,1922]" captionTargetPageId="108" captionText="FIGURE 56. Geographic distribution of: (A) Pogonomyrmex abdominalis Santschi, (B) Pogonomyrmex naegelii Forel, and (C) Pogonomyrmex tenuipubens Santschi. The larger black circle in each panel (when present) denotes the type locality. For P. naegelii, Argentina province borders are in blue, Brazil state borders are in green." pageId="104" pageNumber="105">Figure 56</figureCitation>
A
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="104" pageNumber="105" type="etymology">
<paragraph pageId="104" pageNumber="105">
<emphasis bold="true" box="[199,334,1869,1893]" pageId="104" pageNumber="105">Etymology.</emphasis>
The specific epithet,
<taxonomicName box="[577,715,1868,1892]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[577,715,1868,1892]" italics="true" pageId="104" pageNumber="105">abdominalis</emphasis>
</taxonomicName>
(from Latin,
<emphasis box="[867,969,1868,1892]" italics="true" pageId="104" pageNumber="105">abdomen</emphasis>
- = abdomen), appears to be derived from the enlarged postpetiole on this species, which Santschi described as wider than long for the worker and even larger and more pronounced in the queen.
</paragraph>
</subSubSection>
<caption pageId="105" pageNumber="106" targetBox="[412,1171,193,1956]" targetPageId="105">
<paragraph pageId="105" pageNumber="106">
<emphasis bold="true" box="[151,288,1978,2000]" pageId="105" pageNumber="106">FIGURE 53.</emphasis>
Photograph of
<taxonomicName authority="Santschi" authorityName="Santschi" box="[447,830,1978,2000]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[447,738,1978,2000]" italics="true" pageId="105" pageNumber="106">Pogonomyrmex abdominalis</emphasis>
Santschi
</taxonomicName>
worker: (
<emphasis bold="true" box="[932,950,1978,2000]" pageId="105" pageNumber="106">A</emphasis>
) frontal view of head, (
<emphasis bold="true" box="[1194,1211,1979,2000]" pageId="105" pageNumber="106">B</emphasis>
) lateral view of body, and (
<emphasis bold="true" box="[202,220,2009,2031]" pageId="105" pageNumber="106">C</emphasis>
) dorsal view of body (CASENT0235283). Photographs by Will Ericson from www.AntWeb.org.
</paragraph>
</caption>
<caption pageId="106" pageNumber="107" targetBox="[314,1263,193,1606]" targetPageId="106">
<paragraph pageId="106" pageNumber="107">
<emphasis bold="true" box="[151,288,1637,1659]" pageId="106" pageNumber="107">FIGURE 54.</emphasis>
Photograph from posterior margin of head looking down onto lateral clypeal lobes: (A)
<taxonomicName box="[1189,1436,1637,1659]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[1189,1436,1637,1659]" italics="true" pageId="106" pageNumber="107">Pogonomyrmex naegelii</emphasis>
</taxonomicName>
Forel—black arrows show small lobe that projects dorsally from anterior margin of antennal fossa (CASENT0914811), and (B)
<taxonomicName box="[151,449,1698,1720]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[151,449,1698,1720]" italics="true" pageId="106" pageNumber="107">Pogonomyrmex abdominalis</emphasis>
</taxonomicName>
Santschi—red arrows show absence of small lobe on anterior margin of antennal fossa (CASENT0914813). Photographs by Michele Esposito from www.AntWeb.org.
</paragraph>
</caption>
<subSubSection lastPageId="109" lastPageNumber="110" pageId="106" pageNumber="107" type="discussion">
<paragraph lastPageId="109" lastPageNumber="110" pageId="106" pageNumber="107">
<emphasis bold="true" box="[199,332,1795,1820]" pageId="106" pageNumber="107">Discussion.</emphasis>
<emphasis box="[354,691,1795,1820]" italics="true" pageId="106" pageNumber="107">
<taxonomicName box="[354,691,1795,1820]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="abdominalis">Pogonomyrmex abdominalis</taxonomicName>
</emphasis>
co-occurs with the other two
<taxonomicName box="[1117,1251,1795,1820]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[1117,1131,1796,1819]" italics="true" pageId="106" pageNumber="107">P</emphasis>
.
<emphasis box="[1161,1251,1795,1820]" italics="true" pageId="106" pageNumber="107">naegelii</emphasis>
</taxonomicName>
-group species.
<taxonomicName box="[151,472,1831,1856]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[151,472,1831,1856]" italics="true" pageId="106" pageNumber="107">Pogonomyrmex abdominalis</emphasis>
</taxonomicName>
can be distinguished from
<emphasis box="[778,940,1831,1856]" italics="true" pageId="106" pageNumber="107">
<taxonomicName box="[778,940,1831,1856]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="tenuipubens">P. tenuipubens</taxonomicName>
</emphasis>
based on the following characters: (1) hairs on the mesosoma and psammophore are coarse and much longer than width of cephalic interrugae, and (2) approximately 810 coarse rugae between frontal lobes. In
<taxonomicName box="[814,976,1903,1928]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="tenuipubens">
<emphasis box="[814,976,1903,1928]" italics="true" pageId="106" pageNumber="107">P. tenuipubens</emphasis>
</taxonomicName>
: (1) the abundant hairs on the mesosoma and psammophore are thin and delicate, their maximal length is similar to the width of cephalic interrugae, and (2) more than 15 fine rugae are present between the frontal lobes.
<taxonomicName box="[846,1167,1975,2000]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[846,1167,1975,2000]" italics="true" pageId="106" pageNumber="107">Pogonomyrmex abdominalis</emphasis>
</taxonomicName>
is distinguished from
<emphasis italics="true" pageId="106" pageNumber="107">
<taxonomicName class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="naegelii">P. naegelii</taxonomicName>
</emphasis>
based on the following characters: (1) usually larger (HW = 1.141.33 mm;
<figureCitation box="[1101,1215,2011,2036]" captionStart="FIGURE 57" captionStartId="109.[151,249,1498,1520]" captionTargetBox="[162,1430,204,1468]" captionTargetId="figure@109.[151,1436,193,1476]" captionTargetPageId="109" captionText="FIGURE 57. Bivariate plots for morphological comparisons of Pogonomyrmex abdominalis Santschi, P. naegelii Forel, and P. tenuipubens Santschi: (A) head width versus head length, (B) head width versus petiolar node width, (C) head width versus postpetiole width, and (D) petiolar node width versus postpetiole width." pageId="106" pageNumber="107">
<emphasis bold="true" box="[1101,1215,2011,2036]" pageId="106" pageNumber="107">Figure 57</emphasis>
</figureCitation>
), (2) lacking small lobe that projects dorsally from anterior margin of antennal fossa, (3) peduncle of petiole and anterior surface of petiolar node meet at or near a right angle, and (4) the posterior surface of petiolar node wider than distance between tips of superior propodeal spines. In
<taxonomicName box="[658,775,1698,1722]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[658,775,1698,1722]" italics="true" pageId="109" pageNumber="110">P. naegelii</emphasis>
</taxonomicName>
: (1) the body is usually smaller (HW = 1.051.23 mm), (2) a small lobe projects dorsally from the anterior margin of the antennal fossa (
<emphasis bold="true" box="[1007,1120,1734,1758]" pageId="109" pageNumber="110">
<figureCitation box="[1007,1120,1734,1758]" captionStart="FIGURE 54" captionStartId="106.[151,249,1637,1659]" captionTargetBox="[314,1263,193,1606]" captionTargetId="figure@106.[314,1273,193,1615]" captionTargetPageId="106" captionText="FIGURE 54. Photograph from posterior margin of head looking down onto lateral clypeal lobes: (A) Pogonomyrmex naegelii Forel—black arrows show small lobe that projects dorsally from anterior margin of antennal fossa (CASENT 0914811), and (B) Pogonomyrmex abdominalis Santschi—red arrows show absence of small lobe on anterior margin of antennal fossa (CASENT 0914813). Photographs by Michele Esposito from www. AntWeb. org." pageId="109" pageNumber="110">Figure 54</figureCitation>
</emphasis>
), (3) the peduncle of petiole and anterior surface of petiolar node meet at an obtuse angle, and (4) the width of the posterior surface of petiolar node is similar to or slightly greater than the distance between the tips of the superior propodeal spines.
</paragraph>
<caption pageId="107" pageNumber="108" targetBox="[395,1187,193,1956]" targetPageId="107">
<paragraph pageId="107" pageNumber="108">
<emphasis bold="true" box="[151,289,1978,2000]" pageId="107" pageNumber="108">FIGURE 55.</emphasis>
Photograph of
<taxonomicName authority="Santschi" authorityName="Santschi" box="[451,837,1978,2000]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[451,743,1978,2000]" italics="true" pageId="107" pageNumber="108">Pogonomyrmex abdominalis</emphasis>
Santschi
</taxonomicName>
alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body (CASENT0914378). Photographs by Michele Esposito from www.AntWeb.org.
</paragraph>
</caption>
<caption pageId="108" pageNumber="109" targetBox="[420,1168,183,1922]" targetPageId="108">
<paragraph pageId="108" pageNumber="109">
<emphasis bold="true" box="[151,289,1946,1968]" pageId="108" pageNumber="109">FIGURE 56.</emphasis>
Geographic distribution of: (
<emphasis bold="true" box="[595,613,1946,1968]" pageId="108" pageNumber="109">A</emphasis>
)
<taxonomicName authority="Santschi" authorityName="Santschi" box="[630,1019,1946,1968]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[630,923,1946,1968]" italics="true" pageId="108" pageNumber="109">Pogonomyrmex abdominalis</emphasis>
Santschi
</taxonomicName>
, (
<emphasis bold="true" box="[1040,1057,1947,1968]" pageId="108" pageNumber="109">B</emphasis>
)
<taxonomicName authority="Forel" authorityName="Forel" box="[1074,1387,1946,1968]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[1074,1323,1946,1968]" italics="true" pageId="108" pageNumber="109">Pogonomyrmex naegelii</emphasis>
Forel
</taxonomicName>
, and (
<emphasis bold="true" box="[159,177,1977,1999]" pageId="108" pageNumber="109">C</emphasis>
)
<taxonomicName authority="Santschi. The" authorityName="Santschi. The" box="[192,624,1977,1999]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="tenuipubens">
<emphasis box="[192,480,1977,1999]" italics="true" pageId="108" pageNumber="109">Pogonomyrmex tenuipubens</emphasis>
Santschi. The
</taxonomicName>
larger black circle in each panel (when present) denotes the type locality. For
<taxonomicName class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis italics="true" pageId="108" pageNumber="109">P. naegelii</emphasis>
</taxonomicName>
, Argentina province borders are in blue, Brazil state borders are in green.
</paragraph>
</caption>
<caption pageId="109" pageNumber="110" targetBox="[162,1430,204,1468]" targetPageId="109">
<paragraph pageId="109" pageNumber="110">
<emphasis bold="true" box="[151,287,1498,1520]" pageId="109" pageNumber="110">FIGURE 57.</emphasis>
Bivariate plots for morphological comparisons of
<taxonomicName authority="Santschi" authorityName="Santschi" box="[800,1185,1498,1520]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[800,1091,1498,1520]" italics="true" pageId="109" pageNumber="110">Pogonomyrmex abdominalis</emphasis>
Santschi
</taxonomicName>
,
<taxonomicName authority="Forel" authorityName="Forel" box="[1196,1364,1498,1520]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[1196,1302,1498,1520]" italics="true" pageId="109" pageNumber="110">P. naegelii</emphasis>
Forel
</taxonomicName>
, and
<taxonomicName authority="Santschi" authorityName="Santschi" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="tenuipubens">
<emphasis italics="true" pageId="109" pageNumber="110">P. tenuipubens</emphasis>
Santschi
</taxonomicName>
: (
<emphasis bold="true" box="[391,409,1529,1551]" pageId="109" pageNumber="110">A</emphasis>
) head width versus head length, (
<emphasis bold="true" box="[763,780,1530,1551]" pageId="109" pageNumber="110">B</emphasis>
) head width versus petiolar node width, (
<emphasis bold="true" box="[1214,1233,1529,1551]" pageId="109" pageNumber="110">C</emphasis>
) head width versus postpetiole width, and (
<emphasis bold="true" box="[390,408,1560,1581]" pageId="109" pageNumber="110">D</emphasis>
) petiolar node width versus postpetiole width.
</paragraph>
</caption>
<paragraph pageId="109" pageNumber="110">
In his description, Santschi listed Alta Gracia, Córdoba, as the type locality for
<taxonomicName box="[1139,1309,1842,1866]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[1139,1309,1842,1866]" italics="true" pageId="109" pageNumber="110">P. abdominalis</emphasis>
</taxonomicName>
, as did all subsequent authors (see Gallardo, 1932; Kempf, 1972; Kusnezov, 1951). However, labels on all syntypes say, “Alta Gracia La Granja, Sierras de Córdoba”. Alta Gracia and La Granja are two small towns that are located approximately 75 km apart along the eastern side of the Sierras de Córdoba. Thus, the labels suggest that the type locality was somewhere between these two towns.
</paragraph>
</subSubSection>
<subSubSection pageId="110" pageNumber="111" type="biology_ecology">
<paragraph pageId="110" pageNumber="111">
<emphasis bold="true" box="[199,294,151,176]" pageId="110" pageNumber="111">Biology.</emphasis>
<taxonomicName box="[302,623,151,176]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[302,623,151,176]" italics="true" pageId="110" pageNumber="111">Pogonomyrmex abdominalis</emphasis>
</taxonomicName>
is a solitary forager. Nests usually have a tumulus that is up to 15 cm in diameter. All series collected by the author were in open, grassy, park-like habitats. Colonies probably contain up to about 500 workers.
</paragraph>
<paragraph pageId="110" pageNumber="111">
<taxonomicName box="[199,528,260,284]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[199,528,260,284]" italics="true" pageId="110" pageNumber="111">Pogonomyrmex abdominalis</emphasis>
</taxonomicName>
appears to be a relatively uncommon species that has a wide geographic distribution. Little is known about the biology of
<taxonomicName box="[722,890,295,320]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[722,890,295,320]" italics="true" pageId="110" pageNumber="111">P. abdominalis</emphasis>
</taxonomicName>
but it is probably similar to that of
<taxonomicName box="[1308,1433,295,320]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[1308,1322,296,320]" italics="true" pageId="110" pageNumber="111">P</emphasis>
.
<emphasis box="[1339,1437,295,320]" italics="true" pageId="110" pageNumber="111">naegelii.</emphasis>
</taxonomicName>
Alate queens were collected as part of the type series (no date) and on 24 February, suggesting that mating flights occur during the austral summer. Additional queens need to be collected to determine if
<taxonomicName box="[1208,1379,367,392]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[1208,1379,367,392]" italics="true" pageId="110" pageNumber="111">P. abdominalis</emphasis>
</taxonomicName>
also produces ergatoid queens, as occurs in
<taxonomicName box="[611,732,404,428]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="naegelii">
<emphasis box="[611,732,404,428]" italics="true" pageId="110" pageNumber="111">P. naegelii</emphasis>
</taxonomicName>
and related North American species (Heinze, Hölldobler, &amp; Cover, 1992; Johnson
<emphasis box="[402,458,439,464]" italics="true" pageId="110" pageNumber="111">et al.</emphasis>
, 2007).
</paragraph>
</subSubSection>
<subSubSection pageId="110" pageNumber="111" type="distribution">
<paragraph pageId="110" pageNumber="111">
<taxonomicName box="[199,520,476,500]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[199,520,476,500]" italics="true" pageId="110" pageNumber="111">Pogonomyrmex abdominalis</emphasis>
</taxonomicName>
appears to be a lowland species that inhabits sites at elevations from 101000 m. This species occurs in the Dry Chaco, Humid Pampas, Uruguayan Savanna, Alto Paraná Atlantic Forest, Cerrado, Bahia Interior Forests, and Beni Savanna ecoregions as defined by Olson
<emphasis box="[997,1048,548,572]" italics="true" pageId="110" pageNumber="111">et al</emphasis>
. (2001) (
<emphasis bold="true" box="[1157,1293,548,572]" pageId="110" pageNumber="111">
<figureCitation box="[1157,1272,548,572]" captionStart="FIGURE 56" captionStartId="108.[151,249,1946,1968]" captionTargetBox="[420,1168,183,1922]" captionTargetId="figure@108.[420,1176,183,1922]" captionTargetPageId="108" captionText="FIGURE 56. Geographic distribution of: (A) Pogonomyrmex abdominalis Santschi, (B) Pogonomyrmex naegelii Forel, and (C) Pogonomyrmex tenuipubens Santschi. The larger black circle in each panel (when present) denotes the type locality. For P. naegelii, Argentina province borders are in blue, Brazil state borders are in green." pageId="110" pageNumber="111">Figure 56</figureCitation>
A
</emphasis>
). It is likely that
<taxonomicName box="[203,371,583,608]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis box="[203,371,583,608]" italics="true" pageId="110" pageNumber="111">P. abdominalis</emphasis>
</taxonomicName>
occurs in several additional ecoregions given the few records that are scattered over a very large geographic area.
</paragraph>
</subSubSection>
</treatment>
</document>