310 lines
42 KiB
XML
310 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.100798" ID-Pensoft-Pub="1313-2970-1168-295" ID-Pensoft-UUID="F19054F21F3D516FA0292C896F566182" ID-ZooBank="745FFCD5E3E84DC1B76CC82AD1AC5C64" ModsDocID="1313-2970-1168-295" checkinTime="1688439520051" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Zhao, Lingchen & Zhang, Feng" docDate="2023" docId="7499EA7A3D1557788DC98A6A384841B2" docLanguage="en" docName="ZooKeys 1168: 295-327" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.100798" docTitle="Catatemnus tibetanus Hou & Zhao & Zhang 2023, sp. nov." docType="treatment" docUuid="35D3757B-4668-4BA4-A512-7DB9EC52A87E" docUuidSource="ZooBank" docVersion="1" id="F19054F21F3D516FA0292C896F566182" lastPageNumber="295" masterDocId="F19054F21F3D516FA0292C896F566182" masterDocTitle="First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)" masterLastPageNumber="327" masterPageNumber="295" pageNumber="295" updateTime="1688439520051" updateUser="pensoft">
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<mods:titleInfo id="8FD78CC366C4975B699B747DB1F14FE3">
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<mods:title id="3D4AA455FE471F75D8BE5CE5509D94D6">First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)</mods:title>
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<mods:name id="BA58A366A2590A816C442C976B77B7D5" type="personal">
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<mods:namePart id="4F797D514468198127A03070FA8EE89B">Hou, Yanmeng</mods:namePart>
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<mods:affiliation id="12C0CB33138B1B46B515897237A2DFD3">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="2344B8CE31377D490ADEA5C06D78BAEE">Zhao, Lingchen</mods:namePart>
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<mods:nameIdentifier id="00720D83621D76638B7CA1267C2F6AC5" type="ORCID">https://orcid.org/0000-0001-8477-836X</mods:nameIdentifier>
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<mods:affiliation id="CC4C2D0103D16B9FBD7C4ABBA15CCEC8">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China & Changzhi Tunliu District No. 6 Middle School, Changzhi, Shanxi 046100, China</mods:affiliation>
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<mods:namePart id="900A316C0BAA9DFDEF1AEC402245A9BF">Zhang, Feng</mods:namePart>
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<mods:nameIdentifier id="BACA399DD075A8F367EF2ECE15FD84B7" type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="723A11999CE570B25B2EEA768F4A2FB5">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier id="2E8E634AB463C09DF77C5239F2026D34" type="email">dudu06042001@163.com</mods:nameIdentifier>
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<mods:title id="B2CA7695E2CD3EBB97761FF7418D447C">ZooKeys</mods:title>
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<mods:date id="378B0A52B3B37309E5D5422167F440A6">2023</mods:date>
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<mods:number id="41AD9E854D6149A410EF69AD956DA60D">2023-07-03</mods:number>
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<mods:number id="1C886774E4804FB39F9F2C7F4CE359FF">1168</mods:number>
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<mods:classification id="F5E25B364223EDB2E5BC119EE28FC246">journal article</mods:classification>
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<mods:identifier id="027EF36490683E92DA1F2CA77814F7BE" type="DOI">http://dx.doi.org/10.3897/zookeys.1168.100798</mods:identifier>
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<treatment id="7499EA7A3D1557788DC98A6A384841B2" LSID="urn:lsid:zoobank.org:act:35D3757B-4668-4BA4-A512-7DB9EC52A87E" httpUri="http://treatment.plazi.org/id/7499EA7A3D1557788DC98A6A384841B2" lastPageNumber="295" pageId="0" pageNumber="295">
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<subSubSection id="2AE7AE6A1D98F270E0A9624B8ECDBD62" pageId="0" pageNumber="295" type="nomenclature">
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<paragraph id="E3FD36B60F5F1928E8F7E40C8C6FCAAE" pageId="0" pageNumber="295">
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<taxonomicName id="756604344AD6884C33C017CD8B7AD821" LSID="https://zoobank.org/35D3757B-4668-4BA4-A512-7DB9EC52A87E" authority="Hou & Zhao & Zhang, 2023" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tibetanus" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tibetanus" status="sp. nov.">Catatemnus tibetanus</taxonomicName>
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<taxonomicNameLabel id="D88343A3E1A8CF3105751F9B1759DA39" pageId="0" pageNumber="295">sp. nov.</taxonomicNameLabel>
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<subSubSection id="05333D3E6508802BC6628A959F6F6BFF" pageId="0" pageNumber="295" type="description">
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<paragraph id="37D9089303FEC096DB3525C906CF155B" pageId="0" pageNumber="295">
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<figureCitation id="67E824FBD6F42E7007367C92FCF8CFBD" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Catatemnus tibetanus sp. nov., holotype male A habitus (dorsal view) B carapace (dorsal view) C left pedipalp (dorsal view) D left chela (lateral view) E left chela fingers (lateral view) F genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 1.00 mm (A); 0.20 mm (B-I)." figureDoi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295">Figs 16</figureCitation>
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<figureCitation id="9F6344C134966AA0527325FC2079880D" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">, 17 (西藏猫伪蝎)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="FB150059DF250BDD3F7DB4144D924677" pageId="0" pageNumber="295" type="type material">
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<paragraph id="590F4D29DB3898E1B1A9C3EA254CC42E" pageId="0" pageNumber="295">Type material.</paragraph>
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<paragraph id="69EF66177E9DCC4DEC8C3CFDF1266E4B" pageId="0" pageNumber="295">
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<emphasis id="3D732D577BC5C411996D17DCCC095904" bold="true" italics="true" pageId="0" pageNumber="295">Holotype</emphasis>
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: China • ♂; Xizang Autonomous Region (Tibet), Motuo County, Beibeng Township, Jiangxin Village;
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<geoCoordinate id="82E2EB47435256F182FF25E13CB4A46E" degrees="29" direction="north" minutes="14.199" orientation="latitude" precision="1" value="29.23665">29°14.199'N</geoCoordinate>
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,
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<geoCoordinate id="303819C5A6F4B5F551D53E146EFC2830" degrees="95" direction="east" minutes="10.740" orientation="longitude" precision="1" value="95.179">95°10.740'E</geoCoordinate>
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; 850 m a.s.l.; 09 Aug. 2015; Yejie Lin leg.; Ps.-MHBU-XZ15080901.
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</paragraph>
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<paragraph id="C2A2A7945CB13E753B74674538F7910E" pageId="0" pageNumber="295">
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<emphasis id="D19680577F7CAD55E0FCA0C5E3143E8A" bold="true" pageId="0" pageNumber="295">Diagnosis</emphasis>
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(♂). This new species is characterized by: two distinct developed eyespots, 1.32
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<normalizedToken id="2D72B88FCE7172DBD924BE8F0B4AC9CD" originalValue="×">x</normalizedToken>
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longer than broad; palpal slightly stout, femur 2.44
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<normalizedToken id="8852A38EE497A5E7E395EA11B6BF7B9B" originalValue="×">x</normalizedToken>
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, chela with pedicel 2.82
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<normalizedToken id="EF2A3E49E8595ED7331491AAA03ADF26" originalValue="×">x</normalizedToken>
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, chela without pedicel 2.62
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<normalizedToken id="B32B3CC596C760F0A421DDE715529A3E" originalValue="×">x</normalizedToken>
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longer than broad; distance between ist and isb farther to distance between est and esb; retrolateral surface of trochanter, prolateral surface of patella, femur and hand and base of chelal fingers granular.
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</paragraph>
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</subSubSection>
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<subSubSection id="2AABB2C63DD6DD72DCBC21D11E5C4894" pageId="0" pageNumber="295" type="description">
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<paragraph id="D7AD73ACD6E7683DC9D8E6E7F2DBB88A" pageId="0" pageNumber="295">Description.</paragraph>
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<paragraph id="8ACDCABC8F23A656B3457AC265BE9AFC" pageId="0" pageNumber="295">
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<emphasis id="5F7D5A3C0271582E798312404854ADC9" bold="true" pageId="0" pageNumber="295">Adult male</emphasis>
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(female unknown) (Figs
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<figureCitation id="BB7363034C30E0AEB7156E5B505D80BF" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Catatemnus tibetanus sp. nov., holotype male A habitus (dorsal view) B carapace (dorsal view) C left pedipalp (dorsal view) D left chela (lateral view) E left chela fingers (lateral view) F genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 1.00 mm (A); 0.20 mm (B-I)." figureDoi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295">16</figureCitation>
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,
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<figureCitation id="C50CD6FB0AA2362E7A0AF10B155CD8B6" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17</figureCitation>
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).
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<emphasis id="98788BCFB17C1F66E11A4543A0C76A67" bold="true" italics="true" pageId="0" pageNumber="295">Color</emphasis>
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: anterior half of carapace and pedipalps reddish brown, posterior half of carapace yellowish brown; tergites brown.
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<emphasis id="C335E7A69A1312E810E48F2F32D1AD26" bold="true" italics="true" pageId="0" pageNumber="295">Carapace</emphasis>
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(Figs
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<figureCitation id="509BB6CFD2D949FB1AA85A7EBF737777" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Catatemnus tibetanus sp. nov., holotype male A habitus (dorsal view) B carapace (dorsal view) C left pedipalp (dorsal view) D left chela (lateral view) E left chela fingers (lateral view) F genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 1.00 mm (A); 0.20 mm (B-I)." figureDoi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295">16B</figureCitation>
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,
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<figureCitation id="5E1FC3608C618D9DF97CE5C7A40E9353" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17A</figureCitation>
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): length distinctly longer than breadth (1.32
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<normalizedToken id="A39116A2B6F4301D2EBF7248D8E001C2" originalValue="×">x</normalizedToken>
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); anterior half darker than posterior half; surface smooth, but with a medial furrow; with two distinct eyespots situated near anterior margin of carapace; anterior margin with four setae, posterior margin with eight setae, 40 in total, each seta acicular and very slightly curved.
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<emphasis id="6C51CD6C14793E879686CDCD8D857ADB" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
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(Fig.
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<figureCitation id="39CF894A94F34B234A0C88771062B7A4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17B-D</figureCitation>
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): much smaller than carapace length; surface smooth; four setae (sbs absent; es shorter than others) and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand; movable finger with one galeal seta; bs dentate apically, is and ls long and acute. Fixed finger with five large retrorse teeth and two or three small apical teeth, movable finger with a long broadly rounded subapical lobe; galea present, short and apically with five branchlets (Fig.
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<figureCitation id="5FDAD3753A0148F6D664289F77CC0962" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17C</figureCitation>
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). Serrula interior connected to fixed finger for entire length, proximally modified to form velum, serrula exterior with 27 lamellae, the basal one longest; lamina exterior present. Rallum comprising four blades, the basal two blades shorter than others, the distal one dentated anteriorly, remainder smooth (Fig.
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<figureCitation id="7C841C6561D5667B450A6F16B57017DA" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17D</figureCitation>
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).
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<emphasis id="D5D5BA8A89EB9BE1CAE883AF34CCA3D8" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
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(Figs
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<figureCitation id="CC3BDB2EF7AE7EA3C853E1F3CC0D1F23" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Catatemnus tibetanus sp. nov., holotype male A habitus (dorsal view) B carapace (dorsal view) C left pedipalp (dorsal view) D left chela (lateral view) E left chela fingers (lateral view) F genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 1.00 mm (A); 0.20 mm (B-I)." figureDoi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295">16C-E</figureCitation>
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,
|
||
<figureCitation id="7914AD3BAE98A5955F84D33D5F576E08" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17E, F</figureCitation>
|
||
): stout, trochanter 1.74
|
||
<normalizedToken id="EA00B2CEDB513902CA23439D01E05241" originalValue="×">x</normalizedToken>
|
||
, femur 2.44
|
||
<normalizedToken id="65BB59D7709CA1482E6942DC1D07626E" originalValue="×">x</normalizedToken>
|
||
, patella 1.98
|
||
<normalizedToken id="6614EB7D9D32B8ADA9DCD7BA3DEC77A5" originalValue="×">x</normalizedToken>
|
||
, chela with pedicel (without pedicel) 2.82
|
||
<normalizedToken id="80E69B3E49B8BCB69FA8A14FA6E3F1F6" originalValue="×">x</normalizedToken>
|
||
(2.62
|
||
<normalizedToken id="65E766ED0A6942236E0C968DB101FA72" originalValue="×">x</normalizedToken>
|
||
), hand without pedicel 1.50
|
||
<normalizedToken id="FC271194868A20796A536B5EA7C93BB6" originalValue="×">x</normalizedToken>
|
||
longer than broad; movable chelal finger 0.66
|
||
<normalizedToken id="CB0CAC4169B82D3B592F12A05E85F1EE" originalValue="×">x</normalizedToken>
|
||
longer than hand with pedicel. Setae generally long and acuminate. Retrolateral surface of trochanter, prolateral surface of patella, femur, and hand granular; trochanter with two well-developed conical tubercles. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria: eb and esb situated at base of fixed finger on retrolateral face, esb slightly distal to eb; ib and isb situated at base of fixed finger on prolateral face, isb slightly distal to ib; est in the middle of fixed finger; et near sub-distal of fixed finger; est midway between et and esb; it distal to est and proximal to et; ist situated very slightly distal to est; it closer to ist than to fingertip; distance between est and esb shorter to that of ist and isb; b and sb situated at base of movable finger on retrolateral face; t in the middle of movable finger and at same level as est; sb closer to b than to st; st midway between t and sb (Fig.
|
||
<figureCitation id="0D014473A801F61F94D49EB8283D74EB" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17F</figureCitation>
|
||
). Venom apparatus only present in fixed chelal finger, venom ducts curved and short, terminating in inflated nodus ramosus between et and est. Both chelal fingers with a row of acute teeth, spaced contiguously along the margin, slightly rounded proximally: fixed chelal finger with 26 teeth; movable chelal finger with 32 teeth (nearly as large as teeth on fixed chelal finger); without accessory teeth (Fig.
|
||
<figureCitation id="1917D85C38382F3696BE2FB7BF6BD19D" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17F</figureCitation>
|
||
). Sense spots absent; femur without long tactile setae. Movable chelal finger straight in lateral view (Figs
|
||
<figureCitation id="1D3E8C2373E23343FFC8060002A3A28E" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Catatemnus tibetanus sp. nov., holotype male A habitus (dorsal view) B carapace (dorsal view) C left pedipalp (dorsal view) D left chela (lateral view) E left chela fingers (lateral view) F genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 1.00 mm (A); 0.20 mm (B-I)." figureDoi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295">16E</figureCitation>
|
||
,
|
||
<figureCitation id="CC06EB7BE2ACC2E2755FD2A96B70632A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17F</figureCitation>
|
||
).
|
||
<emphasis id="F83F964F7C10FB860EA024A7FC0568C2" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
|
||
: typical, surface irregular grid decoration, all setae long and acuminate. Pleural membrane longitudinally striate, without setae. Tergites I-III and XI undivided, tergal chaetotaxy I-XI: 8: 7: 9: 6-6: 6-6: 6-5: 7-6: 6-7: 7-5: 7-6 + (2T): 12 (4T). Sternites divided, sternal chaetotaxy IV-XI: 6-5: 9-9: 8-7: 8-8: 7-7: 8-7: 8-7 + (4T): 12 + (4T). Anus (tergite XII and sternite XII) without raised rim. Anterior genital operculum with six setae on each side, posterior margin with four marginal setae on each side.
|
||
<emphasis id="FB279D11CAA06EBB5E7D27F601F569BF" bold="true" italics="true" pageId="0" pageNumber="295">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation id="2A04ED80D3C62739739B11B2E1867303" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Catatemnus tibetanus sp. nov., holotype male A habitus (dorsal view) B carapace (dorsal view) C left pedipalp (dorsal view) D left chela (lateral view) E left chela fingers (lateral view) F genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 1.00 mm (A); 0.20 mm (B-I)." figureDoi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295">16G-I</figureCitation>
|
||
,
|
||
<figureCitation id="510EDB51CE37180F6075C40495726D6A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Catatemnus tibetanus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) C galea D rallum E left pedipalp (dorsal view) F left chelal fingers (lateral view), with details of trichobothrial pattern G genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.20 mm (A, E, G-I); 0.10 mm (B-D, F). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal." figureDoi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295">17H, I</figureCitation>
|
||
): typical, fairly smooth, slightly stout; junction between femora and patellae I and II oblique. Femoropatella of leg IV 2.67
|
||
<normalizedToken id="AC9D9D6DE1286DAC4A4450665F584CC3" originalValue="×">x</normalizedToken>
|
||
longer than deep; tibia 3.53
|
||
<normalizedToken id="0AF6A71C2508A74105F89321B66F32F7" originalValue="×">x</normalizedToken>
|
||
longer than deep; with basal tactile setae on tarsal segment: tarsus 3.82
|
||
<normalizedToken id="7724A2EE948E60064CD3B5A88601F4A7" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS= 0.11); subterminal tarsal setae arcuate and acute. Arolium slightly shorter than claws, not divided; claws smooth.
|
||
</paragraph>
|
||
<caption id="7B0585A469FD7E8AEB76368599054F9E" doi="10.3897/zookeys.1168.100798.figure16" httpUri="https://binary.pensoft.net/fig/872315" pageId="0" pageNumber="295" start="Figure 16" startId="F16">
|
||
<paragraph id="377A0EDFE6B2F154EF79C59A285BFFAF" pageId="0" pageNumber="295">
|
||
<emphasis id="F7DE48F9DF9C972BE6CAB55CB526406C" bold="true" pageId="0" pageNumber="295">Figure 16.</emphasis>
|
||
<taxonomicName id="28EABEB2ABEE7DCFD024AFF8988135F1" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tibetanus" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="C625F7C9E1102732F877D3AA35365A71" italics="true" pageId="0" pageNumber="295">Catatemnus tibetanus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis id="6AFD07AE5BDD5C4AA38795CABE2D3B9E" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
habitus (dorsal view)
|
||
<emphasis id="E84AC4EB6A7538850A5711055ECD1B6A" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="7CE96B23A542D6DC38849004259B730C" bold="true" pageId="0" pageNumber="295">C</emphasis>
|
||
left pedipalp (dorsal view)
|
||
<emphasis id="191DD0B4C5025A3324D6471CFDC0D397" bold="true" pageId="0" pageNumber="295">D</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis id="110DD975BAB8DD5A39993026F348F8F2" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
left chela fingers (lateral view)
|
||
<emphasis id="EE82213B1AFA9B7BDF6D8E52EFBCD80A" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
genital area (ventral view)
|
||
<emphasis id="A5AD353FE67C27A226A323836C575D96" bold="true" pageId="0" pageNumber="295">G</emphasis>
|
||
tarsus of left leg IV (lateral view)
|
||
<emphasis id="2C0DF0974F47699D0F70663AEF8D5F69" bold="true" pageId="0" pageNumber="295">H</emphasis>
|
||
left leg I (lateral view)
|
||
<emphasis id="64FA0F6EC863C7EDE9993A217DC1C78A" bold="true" pageId="0" pageNumber="295">I</emphasis>
|
||
left leg IV (lateral view). Scale bars: 1.00 mm (
|
||
<emphasis id="937F33A8E3FF7F82045908AD7F47ED8A" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
); 0.20 mm (
|
||
<emphasis id="05736505424DAB786F9B9A7A001A67D4" bold="true" pageId="0" pageNumber="295">B-I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CA4E9692AE48CDF6DE409624BB3F7A1F" doi="10.3897/zookeys.1168.100798.figure17" httpUri="https://binary.pensoft.net/fig/872316" pageId="0" pageNumber="295" start="Figure 17" startId="F17">
|
||
<paragraph id="780B8E1A1994E821DFDCBD5A97E55BF9" pageId="0" pageNumber="295">
|
||
<emphasis id="EE2282966819BA4C4EB478D1B1435D20" bold="true" pageId="0" pageNumber="295">Figure 17.</emphasis>
|
||
<taxonomicName id="C0C0542723B48ABD5B3E8BFC038146F4" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tibetanus" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="ACE66E5C0F467CE4FE2A4472EAE0FE09" italics="true" pageId="0" pageNumber="295">Catatemnus tibetanus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis id="DF02BE5CCF8779789FF06478C3223C27" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="E69A65754FC1BFD8F6DDC2EB566BBB7E" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
left chelicera (dorsal view)
|
||
<emphasis id="D6E8452780D83A433F42C893AFD4AFFC" bold="true" pageId="0" pageNumber="295">C</emphasis>
|
||
galea
|
||
<emphasis id="C1706EA9ECCA42FF72CD8BF64C86F24E" bold="true" pageId="0" pageNumber="295">D</emphasis>
|
||
rallum
|
||
<emphasis id="BF42D29E973EB32B24C9330BB87DDACF" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
left pedipalp (dorsal view)
|
||
<emphasis id="6938531656F231E0CFB299660ADD446F" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
left chelal fingers (lateral view), with details of trichobothrial pattern
|
||
<emphasis id="1EF15EDB67BFD40A256655BA058B8700" bold="true" pageId="0" pageNumber="295">G</emphasis>
|
||
genital area (ventral view)
|
||
<emphasis id="503704D35D3CE1A8DEA6EFFF3E3C7160" bold="true" pageId="0" pageNumber="295">H</emphasis>
|
||
left leg I (lateral view)
|
||
<emphasis id="00D2BD82DC2D96B18F171D0799502142" bold="true" pageId="0" pageNumber="295">I</emphasis>
|
||
left leg IV (lateral view). Scale bars: 0.20 mm (
|
||
<emphasis id="0AE21FFF47559EDED300FF494929D388" bold="true" pageId="0" pageNumber="295">A, E, G-I</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis id="384856D2DA64DD02C465C03076CC6D6C" bold="true" pageId="0" pageNumber="295">B-D, F</emphasis>
|
||
). Abbreviations:
|
||
<emphasis id="27C5709C83FDC7C605465035B3068BF4" bold="true" pageId="0" pageNumber="295">b</emphasis>
|
||
= basal;
|
||
<emphasis id="16C3977B2E2646DCD51BBF8D51605C64" bold="true" pageId="0" pageNumber="295">sb</emphasis>
|
||
= sub-basal;
|
||
<emphasis id="14F6D18F0A96851597E06C3AD8935DAF" bold="true" pageId="0" pageNumber="295">st</emphasis>
|
||
= sub-terminal;
|
||
<emphasis id="DB532C14E6A29CDBED3C35C2C65C701E" bold="true" pageId="0" pageNumber="295">t</emphasis>
|
||
= terminal;
|
||
<emphasis id="7C0019DF80DD5E00F69901E2FECB45A7" bold="true" pageId="0" pageNumber="295">ib</emphasis>
|
||
= interior basal;
|
||
<emphasis id="653A5D585894ACEFB34320EE61795AAE" bold="true" pageId="0" pageNumber="295">isb</emphasis>
|
||
= interior sub-basal;
|
||
<emphasis id="BD5E0D2075BD249969D88BD029391C2B" bold="true" pageId="0" pageNumber="295">ist</emphasis>
|
||
= interior sub-terminal;
|
||
<emphasis id="F28522093610B5C3EEA088C1ED36C56D" bold="true" pageId="0" pageNumber="295">it</emphasis>
|
||
= interior terminal;
|
||
<emphasis id="74718415424EF77F36A2FF355499CCF7" bold="true" pageId="0" pageNumber="295">eb</emphasis>
|
||
= exterior basal;
|
||
<emphasis id="F3A6BB8F6CF6BDFA0F7DA694D0126D30" bold="true" pageId="0" pageNumber="295">esb</emphasis>
|
||
= exterior sub-basal;
|
||
<emphasis id="2CFF428961F355EB046646BEF6F3F2DC" bold="true" pageId="0" pageNumber="295">est</emphasis>
|
||
= exterior sub-terminal;
|
||
<emphasis id="3D14F246BCFB744370CC746B77B4009C" bold="true" pageId="0" pageNumber="295">et</emphasis>
|
||
= exterior terminal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B3362F5952A0830381848EF0E60AE3F5" pageId="0" pageNumber="295">
|
||
<emphasis id="1FEE4686366675386D8A47EF12EE6D83" bold="true" pageId="0" pageNumber="295">Dimensions</emphasis>
|
||
(length/breadth or, in the case of the legs, length/depth in mm). Male: body length 3.34. Carapace 0.99/0.75. Pedipalp: trochanter 0.47/0.27, femur 0.83/0.34, patella 0.81/0.41, chela (with pedicel) 1.41/0.50, chela (without pedicel) 1.31/0.50, hand (with pedicel) 0.85/0.50, hand (without pedicel) 0.75/0.50, movable chelal finger length 0.56. Leg I: trochanter 0.20/0.16, femur 0.26/0.20, patella 0.42/0.19, tibia 0.44/0.13, tarsus 0.38/0.09. Leg IV: femoropatella 0.80/0.30, tibia 0.60/0.17, tarsus 0.42/0.11.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A72EE3B04E42B5907F9CA8FE05367B74" pageId="0" pageNumber="295" type="remarks">
|
||
<paragraph id="32A6AB561268141AA2585E591BAA8A5C" pageId="0" pageNumber="295">Remarks.</paragraph>
|
||
<paragraph id="3CF5C6796E49D0FE009F090F5BA42E02" pageId="0" pageNumber="295">
|
||
<taxonomicName id="521E290C0A5AA4B1EDA924FA73F435B4" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus tibetanus" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="D2A69FDE961A0360F44D7E5971BA48CA" italics="true" pageId="0" pageNumber="295">Catatemnus tibetanus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName id="2056E335EA4D9EAD296FB3B72D805CA3" class="Arachnida" kingdom="Animalia" lsidName="C. birmanicus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis id="B43E08BC48E4B30A8420720661B88FA6" italics="true" pageId="0" pageNumber="295">C. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
by the smaller body size (♂) (e.g., body length 3.34 mm vs 4.27 mm; palpal femur length 0.83 mm vs 1.09 mm; movable chelal finger length 0.56 mm vs 0.78 mm) and the trait of eyespots (with two distinct eyespots vs with two faint eyespots); from
|
||
<taxonomicName id="5B20D96D035963BA561551FCED6DE0CA" class="Arachnida" kingdom="Animalia" lsidName="C. concavus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="concavus">
|
||
<emphasis id="E338E11495B702015E26E3EC5A578B34" italics="true" pageId="0" pageNumber="295">C. concavus</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs with two faint eyespots) and the traits of pedipalps (the dorsal surfaces of femur and hand almost smooth vs with minute low granules everywhere except on fingers); from
|
||
<taxonomicName id="4DFE3528F1A4787A8F9A5A806075076E" class="Arachnida" kingdom="Animalia" lsidName="C. monitor" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="monitor">
|
||
<emphasis id="AB5693F265266FA0DC9B54ABA2DC5707" italics="true" pageId="0" pageNumber="295">C. monitor</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs no eyespots visible) and the stouter pedipalps (♂) (e.g., palpal femur 2.44
|
||
<normalizedToken id="3BD68AC4797471A40010DF3FC1E45E63" originalValue="×">x</normalizedToken>
|
||
vs 2.25
|
||
<normalizedToken id="AA94EBA59DDF4AD7EB1B7204ABF7ECAE" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.83 mm vs 1.01 mm); from
|
||
<taxonomicName id="93187A94A6FF7A994C623D12DD572EC7" class="Arachnida" kingdom="Animalia" lsidName="C. nicobarensis" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="BDCE70C441E7802BBC26E87C9902801F" italics="true" pageId="0" pageNumber="295">C. nicobarensis</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs with two indistinct eyespots) and the traits of the palpal trochanter (anterior surface granular vs smooth or almost smooth); from
|
||
<taxonomicName id="80B38E2CC2ABCA62EFCE12C336BA3740" class="Arachnida" kingdom="Animalia" lsidName="C. thorelli" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="thorelli">
|
||
<emphasis id="6A7EF1A4C258752A762F8812F09AA8E6" italics="true" pageId="0" pageNumber="295">C. thorelli</emphasis>
|
||
</taxonomicName>
|
||
by the trait of furrow on the carapace (distinctly curved vs almost straight) and the stouter pedipalps (e.g., palpal femur 2.44
|
||
<normalizedToken id="467310047DFADC704E20DB7DECB2947D" originalValue="×">x</normalizedToken>
|
||
vs 2.82
|
||
<normalizedToken id="9B7605A604C3F84779FB261E6D7CB2D0" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.83 mm vs 0.96 mm; movable chelal finger teeth length 0.56 mm vs 0.70 mm) (
|
||
<bibRefCitation id="5A18EEA6DAFB3AC42C2BBE916C30B228" DOI="https://doi.org/10.5962/bhl.title.10400" author="Thorell, T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" pagination="521 - 729" refId="B9" refString="Thorell, T, 1889. Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885-1887. Annali del Museo Civico di Storia Naturale di Genova 7 (2): 521 - 729, DOI: https://doi.org/10.5962/bhl.title.10400" title="Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885 - 1887." url="https://doi.org/10.5962/bhl.title.10400" volume="7" year="1889">Thorell 1889</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="379D4736F21809600DCC8F10260F11A9" author="Balzan, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" pagination="497 - 512" refId="B1" refString="Balzan, L, 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societe Entomologique de France 60: 497 - 512" title="Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones)." volume="60" year="1891">Balzan 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B63D250FD9B6796BEB3DDF19BFD8C8AC" author="With, CJ" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" refId="B10" refString="With, CJ, 1906. The Danish expedition to Siam 1899-1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7)3: 1-214." title="The Danish expedition to Siam 1899 - 1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7) 3: 1 - 214." year="1906">With 1906</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40A3C6200B3111A9FD219B25DF240C00" DOI="https://doi.org/10.1515/9783111385402" author="Beier, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" refId="B2" refString="Beier, M, 1932. Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1-294. https://doi.org/10.1515/9783111385402" title="Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1 - 294." url="https://doi.org/10.1515/9783111385402" year="1932">Beier 1932</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="495B3E1A99FF0A14A3F3D46A8EABE90F" pageId="0" pageNumber="295" type="distribution">
|
||
<paragraph id="D2A50F56DAA6EDF2DF67CE0CCF73F279" pageId="0" pageNumber="295">Distribution.</paragraph>
|
||
<paragraph id="E8AD6B96723B645F7E87E8B413B24E5F" pageId="0" pageNumber="295">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0EDF54590B53E512225716D26E872FCF" pageId="0" pageNumber="295" type="etymology">
|
||
<paragraph id="454041DCCBBA408BD512A74FC5EFBA1A" pageId="0" pageNumber="295">Etymology.</paragraph>
|
||
<paragraph id="37090F160C6CEBE7685B7F8BDE91AA03" pageId="0" pageNumber="295">Named after the type locality, Tibet (China).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |