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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1132.86022" ID-Pensoft-Pub="1313-2970-1132-127" ID-Pensoft-UUID="E55644A3C1EC5872B08C63B5E2F3912F" ID-ZooBank="46E68453A2FA4D22A0D74509DFFB7C1B" ModsDocID="1313-2970-1132-127" checkinTime="1669812760590" checkinUser="pensoft" docAuthor="Kato, Daichi, Watanabe, Kozo &amp; Kolcsar, Levente-Peter" docDate="2022" docId="12777ED8864B5DC6BD1A68DC255C9E78" docLanguage="en" docName="ZooKeys 1132: 127-162" docOrigin="ZooKeys 1132" docPubDate="2022-11-29" docSource="http://dx.doi.org/10.3897/zookeys.1132.86022" docTitle="Ormosia (Oreophila) confluenta Alexander 1922" docType="treatment" docVersion="1" id="E55644A3C1EC5872B08C63B5E2F3912F" lastPageNumber="127" masterDocId="E55644A3C1EC5872B08C63B5E2F3912F" masterDocTitle="Japanese species of Ormosia Rondani (Diptera, Limoniidae): revision of the subgenera Oreophila Lackschewitz and Parormosia Alexander" masterLastPageNumber="162" masterPageNumber="127" pageNumber="127" updateTime="1669812760590" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Japanese species of Ormosia Rondani (Diptera, Limoniidae): revision of the subgenera Oreophila Lackschewitz and Parormosia Alexander</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kato, Daichi</mods:namePart>
<mods:affiliation>Echigo-Matsunoyama Museum of Natural Science, ' Kyororo', 1712 - 2 Matsunoyama-Matsuguchi, Tokamachi, 942 - 1411, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watanabe, Kozo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7062-595X</mods:nameIdentifier>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kolcsar, Levente-Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7784-2386</mods:nameIdentifier>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kolcsar.peter@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1132</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>127</mods:start>
<mods:end>162</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1132.86022</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1132.86022</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1132-127</mods:identifier>
<mods:identifier type="ZooBank">46E68453A2FA4D22A0D74509DFFB7C1B</mods:identifier>
<mods:identifier type="Pensoft-UUID">E55644A3C1EC5872B08C63B5E2F3912F</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:12777ED8864B5DC6BD1A68DC255C9E78" httpUri="http://treatment.plazi.org/id/12777ED8864B5DC6BD1A68DC255C9E78" lastPageNumber="127" pageId="0" pageNumber="127">
<subSubSection pageId="0" pageNumber="127" type="nomenclature">
<paragraph pageId="0" pageNumber="127">
<taxonomicName LSID="12777ED8-864B-5DC6-BD1A-68DC255C9E78" authority="Alexander, 1922" authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Oreophila">Ormosia (Oreophila) confluenta Alexander, 1922</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of male A Ormosia (Oreophila) confluenta Alexander, 1922 B Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure1" httpUri="https://binary.pensoft.net/fig/775343" pageId="0" pageNumber="127">Figs 1A</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Wing of Ormosia (Oreophila) confluenta Alexander, 1922. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure2" httpUri="https://binary.pensoft.net/fig/775344" pageId="0" pageNumber="127">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure4" httpUri="https://binary.pensoft.net/fig/775346" pageId="0" pageNumber="127">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ormosia (Oreophila) and Ormosia (Parormosia). A Ormosia (Oreophila) confluenta Alexander, 1922 (red), Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. (blue) B Ormosia (Oreophila) sootryeni Lackschewitz, 1935 C Ormosia (Parormosia) diversipes Alexander, 1919 D Ormosia (Parormosia) nippoalpina Alexander, 1941 (red), Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. (blue)." figureDoi="10.3897/zookeys.1132.86022.figure5" httpUri="https://binary.pensoft.net/fig/775347" pageId="0" pageNumber="127">, 5A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="reference_group">
<paragraph pageId="0" pageNumber="127">
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta">Ormosia confluenta</taxonomicName>
in
<bibRefCitation author="Alexander, CP" journalOrPublisher="Insecutor Inscitiae Menstruus" pageId="0" pageNumber="127" pagination="177 - 188" refId="B3" refString="Alexander, CP, 1922. New species of Japanese crane-flies. Part III. (Diptera, Tipulidae). Insecutor Inscitiae Menstruus 10: 177 - 188" title="New species of Japanese crane-flies. Part III. (Diptera, Tipulidae)." volume="10" year="1922">Alexander 1922</bibRefCitation>
: 183: original description (type locality: Japan, Honshu,
<normalizedToken originalValue="Ôsaka">Osaka</normalizedToken>
, Mt. Minomo).
</paragraph>
<paragraph pageId="0" pageNumber="127">
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Ormosia) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Ormosia">Ormosia (Ormosia) confluenta</taxonomicName>
in
<bibRefCitation author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="127" pagination="21 - 75" refId="B11" refString="Alexander, CP, 1953a. Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I. Philippine Journal of Science 82: 21 - 75" title="Records and descriptions of Japanese Tipulidae (Diptera). Part I. The crane-flies of Shikoku. I." volume="82" year="1953 a">Alexander 1953a</bibRefCitation>
: 71: faunistic record.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<taxonomicName class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluent" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluent" subGenus="Oreophila">Ormosia (Oreophila) confluent</taxonomicName>
in
<bibRefCitation author="Nakamura, T" editor="Tochigi, Prefectural Museum" journalOrPublisher="Tochigi Prefectural Museum, Tochigi, Japan" pageId="0" pageNumber="127" pagination="167 - 170" refId="B25" refString="Nakamura, T, 2002. Diptera of the Nasu Imperial Villa, Tochigi, Japan. In: Tochigi, Prefectural Museum, Ed., Flora and Fauna of the Nasu Imperial Villa. Tochigi Prefectural Museum, Tochigi, Japan: 167 - 170" title="Diptera of the Nasu Imperial Villa, Tochigi, Japan." volumeTitle="Flora and Fauna of the Nasu Imperial Villa." year="2002">Nakamura 2002</bibRefCitation>
: 169: faunistic record;
<bibRefCitation author="Nakamura, T" editor="Nakamura, T" journalOrPublisher="Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza). Touka Shobo, Fukuoka, Japan" pageId="0" pageNumber="127" pagination="9 - 53" refId="B26" refString="Nakamura, T, 2014. Family Limoniidae. In: Nakamura, T, Saigusa, T, Suwa, M, Eds., Catalogue of the Insects of Japan. Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza). Touka Shobo, Fukuoka, Japan: 9 - 53" title="Family Limoniidae." volumeTitle="Catalogue of the Insects of Japan." year="2014">Nakamura 2014</bibRefCitation>
: 31: distribution;
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza). Touka Shobo, Fukuoka, Japan" pageId="0" pageNumber="127" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B27" refString="Oosterbroek, P, 2022. Catalogue of the Craneflies of the World (Diptera, Tipuloidea: Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae). https://ccw.naturalis.nl/index.php" title="Catalogue of the Craneflies of the World (Diptera, Tipuloidea: Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae)." url="https://ccw.naturalis.nl/index.php" year="2022">Oosterbroek 2022</bibRefCitation>
: distribution.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="type material">
<paragraph pageId="0" pageNumber="127">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">
<emphasis italics="true" pageId="0" pageNumber="127">Paratype</emphasis>
.
</emphasis>
Japan • ♀; Honshu, Mt. Minomo; 4 May 1921; K. Takeuchi leg.; USNM.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="type material">
<paragraph pageId="0" pageNumber="127">Non-type material examined.</paragraph>
<paragraph pageId="0" pageNumber="127">
Japan • 2 ♂; Honshu, Nagano, Ueda-shi, Sanada-machi-Osa, Kakuma Valley;
<geoCoordinate degrees="36.45378" direction="north" orientation="latitude" precision="1" value="36.45378">36.45378°N</geoCoordinate>
,
<geoCoordinate degrees="138.36592" direction="east" orientation="longitude" precision="1" value="138.36592">138.36592°E</geoCoordinate>
; alt. 1050 m; 16 May 2012; D. Kato leg.; BLKU. • 2 ♂; Honshu, Tokyo, near Tokyo, Mt. Mitake; 10 May 1931; B. Oda leg.; USNM. • 1 ♂; Honshu, Okayama, Maniwa-shi, Hiruzen-Shimotokuyama;
<geoCoordinate degrees="35.32931" direction="north" orientation="latitude" precision="1" value="35.32931">35.32931°N</geoCoordinate>
,
<geoCoordinate degrees="133.59725" direction="east" orientation="longitude" precision="1" value="133.59724">133.59725°E</geoCoordinate>
; alt. 780 m; 1 May 2016; D. Kato leg.; BLKU. • 1 ♀; Honshu, Hiroshima,
<normalizedToken originalValue="Akiôta-chô">Akiota-cho</normalizedToken>
,
<normalizedToken originalValue="Yokogô">Yokogo</normalizedToken>
;
<geoCoordinate degrees="34.59419" direction="north" orientation="latitude" precision="1" value="34.59419">34.59419°N</geoCoordinate>
,
<geoCoordinate degrees="132.14497" direction="east" orientation="longitude" precision="1" value="132.14497">132.14497°E</geoCoordinate>
; alt. 890 m; 18 May 2015; D. Kato leg.; BLKU. • 1 ♂; Shikoku, Tokushima, Mt. Tsurugi; alt. 1400-1950 m; 31 May 1950; Issiki and Ito leg.; USNM. • 1 ♂; Shikoku, Tokushima, Miyoshi-shi, Higashiiya-Ochiai, near Matsuogawa Dam;
<geoCoordinate degrees="33.96478" direction="north" orientation="latitude" precision="1" value="33.96478">33.96478°N</geoCoordinate>
,
<geoCoordinate degrees="133.93908" direction="east" orientation="longitude" precision="1" value="133.93909">133.93908°E</geoCoordinate>
; alt. 900 m; 15 May 2015; • 1 ♂, 1 ♀; same locality; 30 Apr. 2016; D. Kato leg., BLKU. • 6 ♂, 1 ♀; Shikoku, Tokushima, Miyoshi-shi, Higashiiya-Sugeoi, near Nagoro Dam;
<geoCoordinate degrees="33.85182" direction="north" orientation="latitude" precision="1" value="33.85182">33.85182°N</geoCoordinate>
,
<geoCoordinate degrees="134.0234" direction="east" orientation="longitude" precision="5" value="134.0234">134.0234°E</geoCoordinate>
; alt. 920 m; 29 Apr. 2016; • 3 ♂; same locality; 30 Apr. 2016; D. Kato leg. BLKU. • 4 ♂, 1 ♀; Shikoku, Ehime, Kumakogen, River Myogadani, springs;
<geoCoordinate degrees="33.56701" direction="north" orientation="latitude" precision="1" value="33.56701">33.56701°N</geoCoordinate>
,
<geoCoordinate degrees="132.9344" direction="east" orientation="longitude" precision="5" value="132.9344">132.9344 °E</geoCoordinate>
; 1420 m; 8 May 2022; L.-P.
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 1 ♂; Kyushu, Saga, Karatsu-shi, Kyuragi-Hirano, Mt. Sakurei-zan;
<geoCoordinate degrees="33.35701" direction="north" orientation="latitude" precision="1" value="33.35701">33.35701°N</geoCoordinate>
,
<geoCoordinate degrees="130.07038" direction="east" orientation="longitude" precision="1" value="130.07037">130.07038°E</geoCoordinate>
; alt. 860 m; 26 Apr. 2015; D. Kato leg.; BLKU.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="diagnosis">
<paragraph pageId="0" pageNumber="127">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="127">
General coloration yellow to pale brown (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of male A Ormosia (Oreophila) confluenta Alexander, 1922 B Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure1" httpUri="https://binary.pensoft.net/fig/775343" pageId="0" pageNumber="127">1A</figureCitation>
). Vertex often pale brownish grey. Antenna dark brown except scape. Wing yellowish tinged, unpatterned. Legs distal to mid-tibiae gradually becoming dark brown towards tips. Male terminalia: tergite 9 bearing pair of triangular lobes at caudal margin. Gonocoxite slightly produced beyond base of clasper of gonostylus. Clasper of gonostylus wider apically, 3/4 length of lobe of gonostylus, distal part 2
<normalizedToken originalValue="×">x</normalizedToken>
as wide as that of lobe of gonostylus. Interbases fused medially into roundish sac-like plate, without mesal-apical lobe. Female terminalia with cercus almost straight, slightly upcurved on distal part. Genital frame with lateral arm of genital fork roughly triangular, situated at posterior 1/3 of genital fork, with finger-shaped lobe on posterior end. Sternite 9 very small, fan-shaped.
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure1" httpUri="https://binary.pensoft.net/fig/775343" pageId="0" pageNumber="127" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 1.</emphasis>
Habitus of male
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Oreophila">Ormosia (Oreophila) confluenta</taxonomicName>
Alexander, 1922
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) komazawai" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="komazawai" subGenus="Oreophila">Ormosia (Oreophila) komazawai</taxonomicName>
Kato &amp;
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. Scale bars: 2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="redescription">
<paragraph pageId="0" pageNumber="127">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Male</emphasis>
. Body length 2.9-4.2 mm, wing length 4.1-5.9 mm.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Head</emphasis>
: covered with yellow to brown setae. Vertex dusky yellow to pale brownish grey, paler grey on anterior part, often widely dark brown on center of posterior part. Eyes small and widely separated, ~ 1/2 as wide as narrowest point of vertex, ~ 1/3 length of head including rostrum in dorsal view. Rostrum dusky yellow to pale brown, ~ 1/2 length of eye in lateral view. Palpus 5-segmented, 2/3 length of head, dusky yellow on basal two segments, dark brown on succeeding segments, palpomere 1 small, globular, palpomere 2 cylindrical, palpomeres 3-5 globular. Labellum dark brown. Antenna 15 to 16-segmented, relatively short, ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
length of head; scape dusky yellow to pale brown, 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; pedicel dark brown, oval, 2/3 length of scape; flagellomeres dark brown, barrel-shaped, gradually decreasing in size toward apical segment; each flagellomere with ca. six verticils, longest one except in apical segment ~ 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as each segment.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Thorax</emphasis>
: covered with yellow setae. Antepronotum dusky yellow to pale brown; postpronotum pale yellow. Mesonotum subnitidous, pale brown to brown, lateral margin yellow, postero-outer corner of scutal lobe often yellow. Scutellum or mediotergite sometimes yellowish. Prescutal pit indistinctly present, oval to bacilliform. Tuberculate pit indistinctly present, situated slightly anterior to level of prescutal pit. Pleuron dusky yellow to yellow. Wing (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Wing of Ormosia (Oreophila) confluenta Alexander, 1922. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure2" httpUri="https://binary.pensoft.net/fig/775344" pageId="0" pageNumber="127">2</figureCitation>
) yellowish tinged, narrow, 3.5-4
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; stigma absent; Sc ending between level of forks of Rs and R2+3+4; crossvein sc-r indistinct or absent, situated on level of basal 1/3 of Rs if present; R2+3+4 1/7-1/4 length of R3; R2 situated between fork of R2+3+4 and length of itself distal to it; M4 1-1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as M3+4; wing margin between tips of CuP and A1 1.3-2
<normalizedToken originalValue="×">x</normalizedToken>
as long as that between tips of CuP and CuA; A1 almost straight. Halter yellow, ~ 2/3 length of thorax. Legs yellow on coxae to femora; tibiae yellow basally, gradually turning to dark brown toward tips; tarsi dark brown (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of male A Ormosia (Oreophila) confluenta Alexander, 1922 B Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. Scale bars: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure1" httpUri="https://binary.pensoft.net/fig/775343" pageId="0" pageNumber="127">1A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure2" httpUri="https://binary.pensoft.net/fig/775344" pageId="0" pageNumber="127" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 2.</emphasis>
Wing of
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Oreophila">Ormosia (Oreophila) confluenta</taxonomicName>
Alexander, 1922. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Abdomen</emphasis>
: yellow to yellowish ochreous, densely covered with yellow setae.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Male terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3</figureCitation>
): Tergite 9 with pair of triangular lobes at caudal margin, ~ 1/3 length of middle of tergite 9; anterior margin of tergite 9 deeply and widely notched; tergite 9 slightly wider than long including caudal lobe (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3A</figureCitation>
). Sternite 9 almost straight at caudal margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3B</figureCitation>
). Gonocoxite almost same width in whole length, ca. as long as tergite 9, posteroventral margin slightly produced beyond base of clasper of gonostylus (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3A</figureCitation>
). Gonocoxal apodeme short, connected to anterolateral part of interbase (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3D</figureCitation>
). Clasper of gonostylus scabrous, darkened apically, 3/4 length of lobe of gonostylus, gradually wide distally, rounded at tip, distal part 2
<normalizedToken originalValue="×">x</normalizedToken>
as wide as that of lobe of gonostylus in apical view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3C</figureCitation>
). Lobe of gonostylus long finger-shaped, ~ 1/2 length of gonocoxite, slightly curved, distal part flattened in apical view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3C</figureCitation>
). Interbases fused medially into roundish sac-like plate, ca. as long as wide in dorsal view, posterior margin slightly concave, anterolateral part with short arm (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3D</figureCitation>
). Paramere roughly blade-shaped, ca. as long as interbase, anterior end curved inward (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3D</figureCitation>
). Aedeagus dorso-ventrally flattened, ~ 1/3 width of interbase, tip slightly beyond apex of interbase (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3D</figureCitation>
). Sperm pump long oval, anterior end reaching at anterior 1/3 of paramere (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3D, E</figureCitation>
). Ejaculatory apodeme poorly developed (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127">3D, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure3" httpUri="https://binary.pensoft.net/fig/775345" pageId="0" pageNumber="127" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 3.</emphasis>
Male terminalia of
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Oreophila">Ormosia (Oreophila) confluenta</taxonomicName>
Alexander, 1922
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
sternite 9, ventral view
<emphasis bold="true" pageId="0" pageNumber="127">C</emphasis>
gonostylus, outer surface
<emphasis bold="true" pageId="0" pageNumber="127">D</emphasis>
aedeagal complex, dorsal view (left gonocoxal apodeme omitted)
<emphasis bold="true" pageId="0" pageNumber="127">E</emphasis>
aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Female</emphasis>
. Body length 5.0-6.2 mm, wing length 5.6-6.7 mm. Generally resembling male.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Female terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure4" httpUri="https://binary.pensoft.net/fig/775346" pageId="0" pageNumber="127">4</figureCitation>
): yellow to yellowish ochreous, cercus and hypogynial valve amber-colored. Tergites 8 and 9 fused. Cercus almost straight, slightly upcurved on distal part, 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as tergite 10; hypogynial valve ~ 1.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as sternite 8, basal part distinctly wider than that of cercus, gradually narrowed on distal 1/2 toward tip, tip acute, ending near level of middle of cercus (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure4" httpUri="https://binary.pensoft.net/fig/775346" pageId="0" pageNumber="127">4A</figureCitation>
). Genital frame with genital fork, constricted at anterior 1/3, roughly heart-shaped on anterior end; lateral arm of genital fork roughly triangular distally, situated at anterior 2/3 of genital fork, with small finger-shaped lobe at posterior end; sternite 9 very small, narrower than posterior end of genital fork, fan-shaped distally, largely desclerotized on middle of posterior part. Three spermathecal ducts present, middle one very wide; spermathecae indistinct (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Female terminalia of Ormosia (Oreophila) confluenta Alexander, 1922 A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure4" httpUri="https://binary.pensoft.net/fig/775346" pageId="0" pageNumber="127">4B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure4" httpUri="https://binary.pensoft.net/fig/775346" pageId="0" pageNumber="127" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 4.</emphasis>
Female terminalia of
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Oreophila">Ormosia (Oreophila) confluenta</taxonomicName>
Alexander, 1922
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
genital frame, ventral view (left = anterior). Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="distribution">
<paragraph pageId="0" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="127">
Japan (Honshu, Shikoku, and Kyushu) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ormosia (Oreophila) and Ormosia (Parormosia). A Ormosia (Oreophila) confluenta Alexander, 1922 (red), Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. (blue) B Ormosia (Oreophila) sootryeni Lackschewitz, 1935 C Ormosia (Parormosia) diversipes Alexander, 1919 D Ormosia (Parormosia) nippoalpina Alexander, 1941 (red), Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. (blue)." figureDoi="10.3897/zookeys.1132.86022.figure5" httpUri="https://binary.pensoft.net/fig/775347" pageId="0" pageNumber="127">5A</figureCitation>
), Russia (Far East), and Kuril Is.
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure5" httpUri="https://binary.pensoft.net/fig/775347" pageId="0" pageNumber="127" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 5.</emphasis>
Distribution map of Japanese species of
<taxonomicName authority="" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila)" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="subGenus" subGenus="Oreophila">Ormosia (Oreophila)</taxonomicName>
and
<taxonomicName authority="" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia)" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="subGenus" subGenus="Parormosia">Ormosia (Parormosia)</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
<taxonomicName authorityName="Alexander" authorityYear="1922" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) confluenta" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="confluenta" subGenus="Oreophila">Ormosia (Oreophila) confluenta</taxonomicName>
Alexander, 1922 (red),
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) komazawai" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="komazawai" subGenus="Oreophila">Ormosia (Oreophila) komazawai</taxonomicName>
Kato &amp;
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. (blue)
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
<taxonomicName authorityName="Lackschewitz" authorityYear="1935" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) sootryeni" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="sootryeni" subGenus="Oreophila">Ormosia (Oreophila) sootryeni</taxonomicName>
Lackschewitz, 1935
<emphasis bold="true" pageId="0" pageNumber="127">C</emphasis>
<taxonomicName authorityName="Alexander" authorityYear="1919" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) diversipes" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="diversipes" subGenus="Parormosia">Ormosia (Parormosia) diversipes</taxonomicName>
Alexander, 1919
<emphasis bold="true" pageId="0" pageNumber="127">D</emphasis>
<taxonomicName authorityName="Alexander" authorityYear="1941" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) nippoalpina" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="nippoalpina" subGenus="Parormosia">Ormosia (Parormosia) nippoalpina</taxonomicName>
Alexander, 1941 (red),
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" subGenus="Parormosia">Ormosia (Parormosia) phalara</taxonomicName>
Kato &amp;
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. (blue).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="remarks">
<paragraph pageId="0" pageNumber="127">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="127">
This species resembles a Nearctic species,
<taxonomicName baseAuthorityName="Coquillett" baseAuthorityYear="1900" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) flaveola" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="flaveola" subGenus="Oreophila">Ormosia (Oreophila) flaveola</taxonomicName>
(Coquillett, 1900) (
<bibRefCitation author="Coquillett, DW" journalOrPublisher="Proceedings of the Washington Academy of Sciences" pageId="0" pageNumber="127" pagination="389 - 464" refId="B17" refString="Coquillett, DW, 1900. Papers from the Harriman Alaska expedition. IX. Entomological results (3): Diptera. Proceedings of the Washington Academy of Sciences 2: 389 - 464" title="Papers from the Harriman Alaska expedition. IX. Entomological results (3): Diptera." volume="2" year="1900">Coquillett 1900</bibRefCitation>
), and an Eastern Palaearctic species known from North Korea and China,
<taxonomicName authorityName="Alexander" authorityYear="1940" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) yankovskyi" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="yankovskyi" subGenus="Oreophila">Ormosia (Oreophila) yankovskyi</taxonomicName>
Alexander, 1940 (
<bibRefCitation author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="127" pagination="39 - 76" refId="B8" refString="Alexander, CP, 1940a. New or little-known Tipulidae from eastern Asia (Diptera). XLI. Philippine Journal of Science 71: 39 - 76" title="New or little-known Tipulidae from eastern Asia (Diptera). XLI." volume="71" year="1940 a">Alexander 1940a</bibRefCitation>
), but is differentiated from them by the following characters: femora entirely yellow (yellow basally with darker apical parts in
<taxonomicName baseAuthorityName="Coquillett" baseAuthorityYear="1900" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) flaveola" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="flaveola" subGenus="Oreophila">Ormosia (Oreophila) flaveola</taxonomicName>
and
<taxonomicName authorityName="Alexander" authorityYear="1940" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Oreophila) yankovskyi" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="yankovskyi" subGenus="Oreophila">Ormosia (Oreophila) yankovskyi</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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