treatments-xml/data/F9/35/36/F93536231322FFF2FF315E9C00627557.xml

<document id="B802C206A64964D25F1BEF5700952174" ID-CLB-Dataset="55443" ID-DOI="10.11646/zootaxa.5138.4.5" ID-GBIF-Dataset="01147d04-47ed-4aa4-b4be-03220a2e267c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6571729" ID-ZooBank="26EA2794-C0AF-4A1E-A95B-06E59D60967B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653289068978" checkinUser="plazi" docAuthor="Koo, Jun-Mo &amp; Cho, Soowon" docDate="2022" docId="F93536231322FFF2FF315E9C00627557" docLanguage="en" docName="zootaxa.5138.4.5.pdf" docOrigin="Zootaxa 5138 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adela palella Koo &amp; Cho 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="438" masterDocId="050C4E5B1323FFF5FFA65B030477712A" masterDocTitle="Two species of metallic Microlepidoptera, one new species of Adela Latreille [1796] (Adeloidea, Adelidae) and one unrecorded species of Coleophora Hübner 1822 (Gelechioidea, Coleophoridae), from Korea" masterLastPageNumber="444" masterPageNumber="431" pageNumber="432" updateTime="1699337892706" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="268902D0DE08003D767775774F39A668">Two species of metallic Microlepidoptera, one new species of Adela Latreille [1796] (Adeloidea, Adelidae) and one unrecorded species of Coleophora Hübner 1822 (Gelechioidea, Coleophoridae), from Korea</mods:title>
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<mods:namePart id="63841BC6FEEFD40B9EB109C4A52F62DD">Koo, Jun-Mo</mods:namePart>
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<mods:affiliation id="030FC40AABADC73CADF18365D1D78679">Insect Systematic Bioinformatics Laboratory, Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea taran 9539 @ gmail. com; https: // orcid. org / 0000 - 0003 - 2639 - 6456</mods:affiliation>
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<mods:namePart id="C8A6BEB2A3D1C05361ACE947170DBD17">Cho, Soowon</mods:namePart>
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<mods:affiliation id="3F4C0D7D24C4E860860E8BEB6CDDAF9D">Insect Systematic Bioinformatics Laboratory, Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea taran 9539 @ gmail. com; https: // orcid. org / 0000 - 0003 - 2639 - 6456 &amp; chosoowon @ gmail. com; https: // orcid. org / 0000 - 0001 - 6085 - 0064</mods:affiliation>
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<taxonomicName id="B69CFCB61322FFF4FF315E9C05B87490" ID-CoL="9BWQ4" authority="Koo &amp; Cho" authorityName="Koo &amp; Cho" authorityYear="2022" box="[151,463,1439,1466]" class="Insecta" family="Adelidae" genus="Adela" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="432" phylum="Arthropoda" rank="species" species="palella" status="sp. nov.">
<emphasis id="43E85B271322FFF4FF315E9C05417490" bold="true" box="[151,310,1439,1466]" italics="true" pageId="1" pageNumber="432">Adela palella</emphasis>
Koo &amp; Cho
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, 
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<emphasis id="43E85B271322FFF4FF315EC7074374F4" bold="true" box="[151,820,1476,1502]" pageId="1" pageNumber="432">(Korean name: bo-la-bich-bi-dan-gin-su-yeom-na-bang)</emphasis>
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</paragraph>
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<paragraph id="712387351322FFF4FF315EEB066C7728" blockId="1.[151,1029,1439,1575]" box="[151,539,1512,1538]" pageId="1" pageNumber="432">
(
<figureCitation id="E9A79BB01322FFF4FF395EEB05577728" box="[159,288,1512,1538]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="1" pageNumber="432">Figs 1A–K</figureCitation>
; 
<figureCitation id="E9A79BB01322FFF4FE8D5EEB05187728" box="[299,367,1512,1538]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[180,1408,193,1787]" captionTargetId="figure-24@4.[164,1424,181,1798]" captionTargetPageId="4" captionText="FIGURE 2. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0141; B, dorsal view of genitalia; B', lateral view of genitalia; B'', ventral view of genitalia; C, juxta, gen. slide no. KJM0210; D, dorsal view of aedeagus; D', lateral view of anterior 2/5 of aedeagus; E, close-up of transtilla; E', close-up of left anterior process of transtilla; F, inner view of left valva. Abbreviations: ap—anterior process of transtilla, mp—medi-posterior process of transtilla, dc—dorsal corner of valva, tp—triangular process of valva, dm—distal margin of valva, vc—ventral corner of valva, vm—ventral margin of valva." figureDoi="http://doi.org/10.5281/zenodo.6571733" httpUri="https://zenodo.org/record/6571733/files/figure.png" pageId="1" pageNumber="432">2A–F</figureCitation>
; 
<figureCitation id="E9A79BB01322FFF4FEDC5EEB05B57728" box="[378,450,1512,1538]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="1" pageNumber="432">3A–H</figureCitation>
; 
<figureCitation id="E9A79BB01322FFF4FE6B5EEB05877728" box="[461,496,1512,1538]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="1" pageNumber="432">4A</figureCitation>
–F')
</paragraph>
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LSID: 
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<emphasis id="43E85B271322FFF4FF315D5705107744" bold="true" box="[151,359,1620,1646]" pageId="1" pageNumber="432">Type specimens.</emphasis>
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• 
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, genitalia in a tube containing glycerin under the specimen (except left valva, juxta and aedeagus), gen. slide no. KJM0210, wings slide no. KJM0141 (with the right antenna), 
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barcode 
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<collectorName id="DC69E2E31322FFF4FD3D5C2E077E766D" box="[667,777,1837,1863]" pageId="1" pageNumber="432">J.M. Koo</collectorName>
, gen. slide no. KJM0211 (mounted laterally), wings slide no. KJM0208, 
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barcode 
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accession no.: 
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848406
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), deposited in 
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<location id="7443D1EE1322FFF4FA8A5C5301EB7640" LSID="urn:lsid:plazi:treatment:F93536231322FFF2FF315E9C00627557:7443D1EE1322FFF4FA8A5C5301EB7640" box="[1324,1436,1872,1898]" country="South Korea" name="Specimen" pageId="1" pageNumber="432" stateProvince="Chungbuk">Specimen</location>
accession no.: 
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<collectingCountry id="098BC7A51322FFF4FDF45C7606EE76A5" box="[594,665,1909,1935]" name="South Korea" pageId="1" pageNumber="432">Korea</collectingCountry>
, 
<collectingMunicipality id="91471D4F1322FFF4FD035C77073F76A4" box="[677,840,1908,1934]" pageId="1" pageNumber="432">Secheon-dong</collectingMunicipality>
, 
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<collectorName id="DC69E2E31322FFF4FF315C9B04B57698" box="[151,194,1944,1970]" pageId="1" pageNumber="432">Alt.</collectorName>
<quantity id="B6642AD01322FFF4FF6A5C9B056F7699" box="[204,280,1944,1971]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="1" pageNumber="432" unit="m" value="548.0">
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, 
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<collectingDate id="1566581D1322FFF4FE815C9B05D67698" box="[295,417,1944,1970]" pageId="1" pageNumber="432" value="2017-04-30">30.iv.2017</collectingDate>
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, coll. 
<collectorName id="DC69E2E31322FFF4FE4C5C9A06217698" box="[490,598,1944,1971]" pageId="1" pageNumber="432">J.D. Kim</collectorName>
, gen. slide no. KJM0212 (mounted dorsally), wings slide no. KJM0207, 
<emphasis id="43E85B271322FFF4FF315CBF04B076FC" box="[151,199,1980,2006]" italics="true" pageId="1" pageNumber="432">COI</emphasis>
barcode 
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(
<collectorName id="DC69E2E31322FFF4FE655CBF065876FC" box="[451,559,1980,2006]" pageId="1" pageNumber="432">GenBank</collectorName>
accession no.: 
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848407
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), deposited in 
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(
<location id="7443D1EE1322FFF4FBD85CBF009976FC" LSID="urn:lsid:plazi:treatment:F93536231322FFF2FF315E9C00627557:7443D1EE1322FFF4FBD85CBF009976FC" box="[1150,1262,1980,2006]" country="South Korea" name="Specimen" pageId="1" pageNumber="432" stateProvince="Daejon">Specimen</location>
accession no.: 
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3986D4BE1321FFF7FF615B95061C7317" pageId="2" pageNumber="433" type="diagnosis">
<paragraph id="712387351321FFF7FF615B95006070FA" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="433">
<emphasis id="43E85B271321FFF7FF615B950535719A" bold="true" box="[199,322,150,176]" pageId="2" pageNumber="433">Diagnosis.</emphasis>
The species is superficially similar with 
<taxonomicName id="B69CFCB61321FFF7FC835B9400D7719B" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[805,1184,150,177]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FC835B9407A7719A" box="[805,976,150,177]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
(
<bibRefCitation id="150DFAC41321FFF7FC455B9500E0719B" author="Linnaeus, C." box="[995,1175,150,177]" pageId="2" pageNumber="433" refId="ref7165" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Ditio Decima Reformata. Impensis Direct, Laurentii Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
, but there are several distinguishable characters in the adult: 1) in the male of the new species vertex with raised black hairs and sparse few brownish-yellow hairs, but in the male of 
<taxonomicName id="B69CFCB61321FFF7FD025BDC073B71D2" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[676,844,222,249]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FD025BDC073B71D2" box="[676,844,222,249]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
</taxonomicName>
vertex with only black hairs; 2) in the female of the new species antenna not thickened basally, but in the female of 
<taxonomicName id="B69CFCB61321FFF7FCC15A0000787036" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[871,1039,258,285]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FCC15A0000787036" box="[871,1039,258,285]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
</taxonomicName>
antenna thickened in basal ¼. Furthermore, it can be distinguished from the latter by the male genitalia: 1) in the new species the area between dorsal corner and the triangular process of valva concave, while in 
<taxonomicName id="B69CFCB61321FFF7FCE75A48079E704E" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[833,1001,330,357]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FCE75A48079E704E" box="[833,1001,330,357]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
</taxonomicName>
the area flat; 2) in the new species the length of ventral margin of valva 0.5× shorter than the length of valva, while in 
<taxonomicName id="B69CFCB61321FFF7FBA85A6C00C270A2" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1038,1205,366,393]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FBA85A6C00C270A2" box="[1038,1205,366,393]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
</taxonomicName>
the length of ventral margin of valva about 0.3× shorter than the length of valva; 3) in the new species juxtal head of the same length as the width, while in 
<taxonomicName id="B69CFCB61321FFF7FED65AB4066070FA" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[368,535,438,465]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FED65AB4066070FA" box="[368,535,438,465]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
</taxonomicName>
juxtal head about 1.5× longer than the width.
</paragraph>
<paragraph id="712387351321FFF7FF615AD9061C7317" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="433">
<emphasis id="43E85B271321FFF7FF615AD9057F70DE" bold="true" box="[199,264,474,500]" pageId="2" pageNumber="433">Note:</emphasis>
When comparing the barcode sequences obtained from 
<specimenCount id="679A4CBC1321FFF7FC325AD9003C70DE" box="[916,1099,474,500]" pageId="2" pageNumber="433" type="generic">three specimens</specimenCount>
of the new species (showing 100% concordance rate) with those of 
<taxonomicName id="B69CFCB61321FFF7FDE15AFC06997332" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[583,750,510,537]" class="Insecta" family="Noctuidae" genus="Phalaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="433" phylum="Arthropoda" rank="species" species="reaumurella">
<emphasis id="43E85B271321FFF7FDE15AFC06997332" box="[583,750,510,537]" italics="true" pageId="2" pageNumber="433">A. reaumurella</emphasis>
</taxonomicName>
registered in NCBI and BOLD systems, it shows a similarity rate of 96.01–96.20% and 95.81–96.56%.
</paragraph>
</subSubSection>
<subSubSection id="3986D4BE1321FFF2FF6159450625753B" lastPageId="7" lastPageNumber="438" pageId="2" pageNumber="433" type="description">
<paragraph id="712387351321FFF7FF6159450171734B" blockId="2.[151,1437,150,2013]" box="[199,1286,582,609]" pageId="2" pageNumber="433">
<emphasis id="43E85B271321FFF7FF615945052E734A" bold="true" box="[199,345,582,608]" pageId="2" pageNumber="433">Description.</emphasis>
Adult of both sexes (
<figureCitation id="E9A79BB01321FFF7FDEF594506BD734A" box="[585,714,582,608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Figs 1A–K</figureCitation>
; 
<figureCitation id="E9A79BB01321FFF7FD735945068C734A" box="[725,763,582,608]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[180,1408,193,1787]" captionTargetId="figure-24@4.[164,1424,181,1798]" captionTargetPageId="4" captionText="FIGURE 2. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0141; B, dorsal view of genitalia; B', lateral view of genitalia; B'', ventral view of genitalia; C, juxta, gen. slide no. KJM0210; D, dorsal view of aedeagus; D', lateral view of anterior 2/5 of aedeagus; E, close-up of transtilla; E', close-up of left anterior process of transtilla; F, inner view of left valva. Abbreviations: ap—anterior process of transtilla, mp—medi-posterior process of transtilla, dc—dorsal corner of valva, tp—triangular process of valva, dm—distal margin of valva, vc—ventral corner of valva, vm—ventral margin of valva." figureDoi="http://doi.org/10.5281/zenodo.6571733" httpUri="https://zenodo.org/record/6571733/files/figure.png" pageId="2" pageNumber="433">2A</figureCitation>
; 
<figureCitation id="E9A79BB01321FFF7FCA059450739734A" box="[774,846,582,608]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">3A–H</figureCitation>
; 
<figureCitation id="E9A79BB01321FFF7FCFF59450709734A" box="[857,894,582,608]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="2" pageNumber="433">4A</figureCitation>
–B'). Forewing length 6.0– 
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.
</paragraph>
<paragraph id="712387351321FFF7FF61596905A4758A" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="433">
<emphasis id="43E85B271321FFF7FF615969057373AE" box="[199,260,618,644]" italics="true" pageId="2" pageNumber="433">Head</emphasis>
: Male (
<figureCitation id="E9A79BB01321FFF7FEFA596905C273AE" box="[348,437,618,644]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Figs 1A</figureCitation>
–D', F–K). Vertex with raised black hairs, mixed with sparse few brownish-yellow hairs; clypeus silvery black with black hairs laterally, and with sparse brownish-yellow hairs on upper margin and between antennae (
<figureCitation id="E9A79BB01321FFF7FEAD59B1052773E6" box="[267,336,690,716]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Figs 1</figureCitation>
A', B). Antenna filiform, about 2.8× longer than the length of forewing; scape clubhead-shaped, dark bronzy with dense black hairs ventrally; pedicel dark bronzy; flagellum dark bronzy in 1 
<superScript id="86E92A7D1321FFF7FB1F59D500B373CE" attach="none" box="[1209,1220,726,740]" fontSize="6" pageId="2" pageNumber="433">st</superScript>
–3 
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segments, then outer surface silvery white until basal half, then entirely silvery white distally; 1–2 hook-shaped flagellar pegs pointing anteriorly on the dorsal side of flagellar 9–10 
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segments, and each peg with two teeth in a row ventrally (one peg on flagellar 9 
<superScript id="86E92A7D1321FFF7FEC65841051A727A" attach="left" box="[352,365,834,848]" fontSize="6" pageId="2" pageNumber="433">th</superScript>
segment of the right antenna and one peg on each of flagellar 9 
<superScript id="86E92A7D1321FFF7FBE058410024727A" attach="left" box="[1094,1107,834,848]" fontSize="6" pageId="2" pageNumber="433">th</superScript>
and 10 
<superScript id="86E92A7D1321FFF7FB01584100C3727A" attach="left" box="[1191,1204,834,848]" fontSize="6" pageId="2" pageNumber="433">th</superScript>
segments of the left antenna, from 
<specimenCount id="679A4CBC1321FFF7FE9A586405D472AA" box="[316,419,870,897]" pageId="2" pageNumber="433" type="male">one male</specimenCount>
specimen examined in this study); antennal sensillae visible ventrally through the ventral scales-free zones; each zone isolated in a squarish shape by a row of scales until basal ¼ of flagellum, the squarish zones then interconnected and narrowed area of dispersed scales until the half of flagellum, then the interconnected zones re-isolated into single scale-free zones until the distal end of flagellum; the number of sensillae rapidly decreasing beyond the basal half of flagellum (
<figureCitation id="E9A79BB01321FFF7FD2C58F5075D753B" box="[650,810,1014,1041]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Figs 1A, F–K</figureCitation>
). Labial palpus short, same length as scape; 2 
<superScript id="86E92A7D1321FFF7FA8E58F5014F752E" attach="left" box="[1320,1336,1014,1028]" fontSize="6" pageId="2" pageNumber="433">nd</superScript>
segment dark bronzy with few silvery-white scales, and with white hairs sparsely on the inner and dorsal surface and black hairs on the outer and ventral surface, particularly dense hairs ventrally; 3 
<superScript id="86E92A7D1321FFF7FC695F3D07AA7566" attach="left" box="[975,989,1086,1100]" fontSize="6" pageId="2" pageNumber="433">rd</superScript>
segment dark bronzy with white scales, about 0.5× shorter than the length of 2 
<superScript id="86E92A7D1321FFF7FD995F610638755A" attach="left" box="[575,591,1122,1136]" fontSize="6" pageId="2" pageNumber="433">nd</superScript>
segment (
<figureCitation id="E9A79BB01321FFF7FD665F61076B7556" box="[704,796,1122,1148]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Figs 1D</figureCitation>
, D'). Proboscis covered with black hairs and white sensillae without scales (
<figureCitation id="E9A79BB01321FFF7FEC85F8505B2758A" box="[366,453,1158,1184]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="712387351321FFF7FF615FA9012274CE" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="433">
Female (
<figureCitation id="E9A79BB01321FFF7FE8E5FA905F375EE" box="[296,388,1194,1220]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3A</figureCitation>
–G'''). Vertex with raised brownish-yellow hairs, intermixed with few black sparse hairs; clypeus silvery black with black and brownish-yellow hairs laterally, and with sparse brownish-yellow hairs situated on upper margin and between antennae (
<figureCitation id="E9A79BB01321FFF7FD9D5FF106C07427" box="[571,695,1266,1293]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3B, E</figureCitation>
). Antenna filiform, about 1.6× longer than the length of forewing; scape and pedicel the same as in male, except hairless scape; flagellum dark bronzy in basal 3/5 dorsally and laterally, and in basal 4/5 ventrally, then silvery white distally; antennal sensillae visible ventrally through the scales-free zones; each zone well-isolated into a squarish shape by a row of scales, and progressively smaller distally; the number of sensillae rapidly decreasing beyond basal 3/5 (
<figureCitation id="E9A79BB01321FFF7FD4C5E8107F374B6" box="[746,900,1410,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3A, F, G</figureCitation>
–G'''). Labial palpus short, of the same length as scape; 2 
<superScript id="86E92A7D1321FFF7FF545EA50575749E" attach="left" box="[242,258,1446,1460]" fontSize="6" pageId="2" pageNumber="433">nd</superScript>
segment dark bronzy with silvery-white scales, with particularly dense black hairs ventrally; 3 
<superScript id="86E92A7D1321FFF7FA8C5EA5014F749E" attach="left" box="[1322,1336,1446,1460]" fontSize="6" pageId="2" pageNumber="433">rd</superScript>
segment the same as in male (
<figureCitation id="E9A79BB01321FFF7FE275EC9067674CF" box="[385,513,1482,1509]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3C, D</figureCitation>
). Proboscis with sensillae under dark bronzy scales, without hairs (
<figureCitation id="E9A79BB01321FFF7FB565EC9013F74CE" box="[1264,1352,1482,1508]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="712387351321FFF7FF615EED0096762E" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="433">
<emphasis id="43E85B271321FFF7FF615EED056F7722" box="[199,280,1518,1544]" italics="true" pageId="2" pageNumber="433">Thorax</emphasis>
: Male (
<figureCitation id="E9A79BB01321FFF7FED65EED05C47722" box="[368,435,1518,1544]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Figs 1</figureCitation>
A', E; 2A). Thorax bronze green with two elliptical patches side by side formed by black scales under long appressed black hairs (
<figureCitation id="E9A79BB01321FFF7FDC65D1106C17706" box="[608,694,1554,1580]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Fig. 1E</figureCitation>
). Tegula bronze green (
<figureCitation id="E9A79BB01321FFF7FC195D1100637706" box="[959,1044,1554,1580]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Fig. 1E</figureCitation>
). Forewing bronze green with purplish-lustered costal fringe and bronze-green dorsal fringe; costa weakly concave between basal 1/5 and distal ⅓ (
<figureCitation id="E9A79BB01321FFF7FF065D5904A9775E" box="[160,222,1626,1652]" captionStart="FIGURE 1" captionStartId="3.[151,250,1757,1781]" captionTargetBox="[154,1436,183,1733]" captionTargetId="figure-19@3.[151,1436,181,1734]" captionTargetPageId="3" captionText="FIGURE 1. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna)." figureDoi="http://doi.org/10.5281/zenodo.6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="2" pageNumber="433">Fig. 1</figureCitation>
A'); venation (
<figureCitation id="E9A79BB01321FFF7FE245D5905AD775E" box="[386,474,1626,1652]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[180,1408,193,1787]" captionTargetId="figure-24@4.[164,1424,181,1798]" captionTargetPageId="4" captionText="FIGURE 2. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0141; B, dorsal view of genitalia; B', lateral view of genitalia; B'', ventral view of genitalia; C, juxta, gen. slide no. KJM0210; D, dorsal view of aedeagus; D', lateral view of anterior 2/5 of aedeagus; E, close-up of transtilla; E', close-up of left anterior process of transtilla; F, inner view of left valva. Abbreviations: ap—anterior process of transtilla, mp—medi-posterior process of transtilla, dc—dorsal corner of valva, tp—triangular process of valva, dm—distal margin of valva, vc—ventral corner of valva, vm—ventral margin of valva." figureDoi="http://doi.org/10.5281/zenodo.6571733" httpUri="https://zenodo.org/record/6571733/files/figure.png" pageId="2" pageNumber="433">Fig. 2A</figureCitation>
) with free R veins; distance between origins of R 
<subScript id="ED1885701321FFF7FBA45D6E007D7751" attach="left" box="[1026,1034,1645,1659]" fontSize="6" pageId="2" pageNumber="433">3</subScript>
and R 
<subScript id="ED1885701321FFF7FBF55D6E002C7751" attach="left" box="[1107,1115,1645,1659]" fontSize="6" pageId="2" pageNumber="433">4</subScript>
of the same length as that of R 
<subScript id="ED1885701321FFF7FF0C5D9204C577B5" attach="left" box="[170,178,1681,1695]" fontSize="6" pageId="2" pageNumber="433">4</subScript>
and R 
<subScript id="ED1885701321FFF7FF5F5D93057677B4" attach="left" box="[249,257,1680,1694]" fontSize="6" pageId="2" pageNumber="433">5</subScript>
; R 
<subScript id="ED1885701321FFF7FE845D93055D77B4" attach="left" box="[290,298,1680,1694]" fontSize="6" pageId="2" pageNumber="433">5</subScript>
reaching before apex; M veins free; CuA 
<subScript id="ED1885701321FFF7FD535D92068A77B5" attach="left" box="[757,765,1681,1695]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
arising from lower corner of discal cell; origins of CuA 
<subScript id="ED1885701321FFF7FAC05D92011977B5" attach="left" box="[1382,1390,1681,1695]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
and CuA 
<subScript id="ED1885701321FFF7FF6A5DB604A377E9" attach="left" box="[204,212,1717,1731]" fontSize="6" pageId="2" pageNumber="433">2</subScript>
separated by the same distance as from M 
<subScript id="ED1885701321FFF7FD155DB606CC77E9" attach="left" box="[691,699,1717,1731]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
to CuA 
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; 1A+2A forked at about basal ¼; discal cell well-closed. Hindwing grayish brown with purplish luster, with one long frenulum; fringe bronze green, dark brown and orange white; Sc+R 
<subScript id="ED1885701321FFF7FE855DFE055C7621" attach="left" box="[291,299,1789,1803]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
long; Rs free from M 
<subScript id="ED1885701321FFF7FD865DFE065F7621" attach="left" box="[544,552,1789,1803]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
; M 
<subScript id="ED1885701321FFF7FDF65DFE062F7621" attach="left" box="[592,600,1789,1803]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
and M 
<subScript id="ED1885701321FFF7FD015DFE06D87621" attach="left" box="[679,687,1789,1803]" fontSize="6" pageId="2" pageNumber="433">2</subScript>
stalked at about basal 1/6; discal cell well-closed.
</paragraph>
<paragraph id="712387351321FFF7FF615C0D00EC76F6" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="433">
Female (
<figureCitation id="E9A79BB01321FFF7FE8F5C0D05BD7603" box="[297,458,1806,1833]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3A, B, H</figureCitation>
; 
<figureCitation id="E9A79BB01321FFF7FE705C0D058C7602" box="[470,507,1806,1832]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="2" pageNumber="433">4A</figureCitation>
–B'). Thorax bronze green without hairs and patches (
<figureCitation id="E9A79BB01321FFF7FBF25C0D00DB7602" box="[1108,1196,1806,1832]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Fig. 3B</figureCitation>
). Tegula concolorous (
<figureCitation id="E9A79BB01321FFF7FF065C3104807666" box="[160,247,1842,1868]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Fig. 3B</figureCitation>
). Forewing the same as in male except 1A+2A forked at about basal ⅓ (
<figureCitation id="E9A79BB01321FFF7FB865C3100087666" box="[1056,1151,1842,1868]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3A</figureCitation>
; 
<figureCitation id="E9A79BB01321FFF7FB2C5C3100B87667" box="[1162,1231,1842,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="2" pageNumber="433">4A, B</figureCitation>
). Hindwing of the same color as male, with 4–6 frenula (
<figureCitation id="E9A79BB01321FFF7FDED5C5506DD765A" box="[587,682,1878,1904]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Figs 3A</figureCitation>
; 
<figureCitation id="E9A79BB01321FFF7FD115C5506B1765A" box="[695,710,1878,1904]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="2" pageNumber="433">4</figureCitation>
A', B'), one in each of sockets arranged in a row; venation same as male except M 
<subScript id="ED1885701321FFF7FEFA5C8E051376B1" attach="left" box="[348,356,1933,1947]" fontSize="6" pageId="2" pageNumber="433">1</subScript>
and M 
<subScript id="ED1885701321FFF7FE125C8E05CB76B1" attach="left" box="[436,444,1933,1947]" fontSize="6" pageId="2" pageNumber="433">2</subScript>
stalked at base. Hindtibia bronze green with purplish-lustered scales, paler inner surface than outer surface, with long dark brown bristles dorsally; two pairs of spurs, one pair at basal half and the other pair at apex (
<figureCitation id="E9A79BB01321FFF7FF515CC1052676F6" box="[247,337,1986,2012]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Fig. 3H</figureCitation>
). Hindtarsus covered with bronze-green and silvery-white scales (
<figureCitation id="E9A79BB01321FFF7FB925CC100F976F6" box="[1076,1166,1986,2012]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="2" pageNumber="433">Fig. 3H</figureCitation>
).
</paragraph>
<caption id="25E3D7BD1320FFF6FF315DDE01557680" ID-DOI="http://doi.org/10.5281/zenodo.6571731" ID-Zenodo-Dep="6571731" httpUri="https://zenodo.org/record/6571731/files/figure.png" pageId="3" pageNumber="434" startId="3.[151,250,1757,1781]" targetBox="[154,1436,183,1733]" targetPageId="3">
<paragraph id="712387351320FFF6FF315DDE01557680" blockId="3.[151,1436,1757,1964]" pageId="3" pageNumber="434">
<emphasis id="43E85B271320FFF6FF315DDE056477DF" bold="true" box="[151,275,1757,1781]" pageId="3" pageNumber="434">FIGURE 1.</emphasis>
Male (holotype) of 
<taxonomicName id="B69CFCB61320FFF6FE795DDD069177DC" authority="Koo &amp; Cho" authorityName="Koo &amp; Cho" authorityYear="2022" box="[479,742,1758,1782]" class="Insecta" family="Adelidae" genus="Adela" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="434" phylum="Arthropoda" rank="species" species="palella" status="sp. nov.">
<emphasis id="43E85B271320FFF6FE795DDD061177DC" box="[479,614,1758,1782]" italics="true" pageId="3" pageNumber="434">Adela palella</emphasis>
Koo &amp; Cho
</taxonomicName>
, 
<emphasis id="43E85B271320FFF6FD565DDD073577DC" bold="true" box="[752,834,1758,1782]" pageId="3" pageNumber="434">
<taxonomicNameLabel id="58DBE65C1320FFF6FD565DDD073577DC" box="[752,834,1758,1782]" pageId="3" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, adult; A', close-up of adult; B, anterior view of head; C, proboscis; D, outer view of labial palpus; D', inner view of labial palpus; E, dorsal view of thorax; F, outer view of basal (left) antenna; F', close-up of flagellar pegs on left antenna; G, inner view of basal (right) antenna; G', close-up of flagellar peg on right antenna; H, close-up ventral view of partial antenna between base and basal ¼; I, close-up ventral view of partial antenna between basal ¼ and half; J, close-up ventral view of partial antenna between basal half and ¾; K, close-up ventral view of partial antenna between basal ¾ and apex. Abbreviations: sz—scales-free zone, fp—flagellar peg (all for antenna).
</paragraph>
</caption>
<caption id="25E3D7BD1327FFF1FF315C1C048676C1" ID-DOI="http://doi.org/10.5281/zenodo.6571733" ID-Zenodo-Dep="6571733" httpUri="https://zenodo.org/record/6571733/files/figure.png" pageId="4" pageNumber="435" startId="4.[151,250,1823,1847]" targetBox="[180,1408,193,1787]" targetPageId="4">
<paragraph id="712387351327FFF1FF315C1C048676C1" blockId="4.[151,1436,1823,2027]" pageId="4" pageNumber="435">
<emphasis id="43E85B271327FFF1FF315C1C0562761D" bold="true" box="[151,277,1823,1847]" pageId="4" pageNumber="435">FIGURE 2.</emphasis>
Male (holotype) of 
<taxonomicName id="B69CFCB61327FFF1FE4C5C1C068D761D" authority="Koo &amp; Cho" authorityName="Koo &amp; Cho" authorityYear="2022" box="[490,762,1823,1847]" class="Insecta" family="Adelidae" genus="Adela" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="435" phylum="Arthropoda" rank="species" species="palella" status="sp. nov.">
<emphasis id="43E85B271327FFF1FE4C5C1C0603761D" box="[490,628,1823,1847]" italics="true" pageId="4" pageNumber="435">Adela palella</emphasis>
Koo &amp; Cho
</taxonomicName>
, 
<emphasis id="43E85B271327FFF1FCA05C1C072C761D" bold="true" box="[774,859,1823,1847]" pageId="4" pageNumber="435">
<taxonomicNameLabel id="58DBE65C1327FFF1FCA05C1C072C761D" box="[774,859,1823,1847]" pageId="4" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, venation, wings slide no. KJM0141; B, dorsal view of genitalia; B', lateral view of genitalia; B'', ventral view of genitalia; C, juxta, gen. slide no. KJM0210; D, dorsal view of aedeagus; D', lateral view of anterior 2/5 of aedeagus; E, close-up of transtilla; E', close-up of left anterior process of transtilla; F, inner view of left valva. Abbreviations: ap—anterior process of transtilla, mp—medi-posterior process of transtilla, dc—dorsal corner of valva, tp—triangular process of valva, dm—distal margin of valva, vc—ventral corner of valva, vm—ventral margin of valva.
</paragraph>
</caption>
<caption id="25E3D7BD1326FFF0FF315DB800AF7615" ID-DOI="http://doi.org/10.5281/zenodo.6571735" ID-Zenodo-Dep="6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="5" pageNumber="436" startId="5.[151,250,1723,1747]" targetBox="[151,1434,183,1682]" targetPageId="5">
<paragraph id="712387351326FFF0FF315DB800AF7615" blockId="5.[151,1436,1723,1857]" pageId="5" pageNumber="436">
<emphasis id="43E85B271326FFF0FF315DB8056377F9" bold="true" box="[151,276,1723,1747]" pageId="5" pageNumber="436">FIGURE 3.</emphasis>
Female (paratype) of 
<taxonomicName id="B69CFCB61326FFF0FE5D5DB8077F77F9" authority="Koo &amp; Cho" authorityName="Koo &amp; Cho" authorityYear="2022" box="[507,776,1723,1747]" class="Insecta" family="Adelidae" genus="Adela" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="436" phylum="Arthropoda" rank="species" species="palella" status="sp. nov.">
<emphasis id="43E85B271326FFF0FE5D5DB806F377F9" box="[507,644,1723,1747]" italics="true" pageId="5" pageNumber="436">Adela palella</emphasis>
Koo &amp; Cho
</taxonomicName>
, 
<emphasis id="43E85B271326FFF0FCB55DB8071077F9" bold="true" box="[787,871,1723,1747]" pageId="5" pageNumber="436">
<taxonomicNameLabel id="58DBE65C1326FFF0FCB55DB8071077F9" box="[787,871,1723,1747]" pageId="5" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs.
</paragraph>
</caption>
<caption id="25E3D7BD1325FFF3FF315C51071476D0" ID-DOI="http://doi.org/10.5281/zenodo.6571737" ID-Zenodo-Dep="6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="6" pageNumber="437" startId="6.[151,250,1874,1898]" targetBox="[219,1365,191,1843]" targetPageId="6">
<paragraph id="712387351325FFF3FF315C51071476D0" blockId="6.[151,1436,1874,2042]" pageId="6" pageNumber="437">
<emphasis id="43E85B271325FFF3FF315C5105637640" bold="true" box="[151,276,1874,1898]" pageId="6" pageNumber="437">FIGURE 4.</emphasis>
Female (paratype) of 
<taxonomicName id="B69CFCB61325FFF3FE5F5C5107727640" authority="Koo &amp; Cho" authorityName="Koo &amp; Cho" authorityYear="2022" box="[505,773,1874,1898]" class="Insecta" family="Adelidae" genus="Adela" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="437" phylum="Arthropoda" rank="species" species="palella" status="sp. nov.">
<emphasis id="43E85B271325FFF3FE5F5C5106F57640" box="[505,642,1874,1898]" italics="true" pageId="6" pageNumber="437">Adela palella</emphasis>
Koo &amp; Cho
</taxonomicName>
, 
<emphasis id="43E85B271325FFF3FCB65C5107147640" bold="true" box="[784,867,1874,1898]" pageId="6" pageNumber="437">
<taxonomicNameLabel id="58DBE65C1325FFF3FCB65C5107147640" box="[784,867,1874,1898]" pageId="6" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7 
<superScript id="86E92A7D1325FFF3FDA95CBD066D76E6" attach="left" box="[527,538,1982,1996]" fontSize="6" pageId="6" pageNumber="437">th</superScript>
tergite; E', dorsal view of abdominal 7 
<superScript id="86E92A7D1325FFF3FC005CBD07C676E6" attach="left" box="[934,945,1982,1996]" fontSize="6" pageId="6" pageNumber="437">th</superScript>
tergite; F, lateral view of abdominal 7 
<superScript id="86E92A7D1325FFF3FA9F5CBD013376E6" attach="left" box="[1337,1348,1982,1996]" fontSize="6" pageId="6" pageNumber="437">th</superScript>
sternite; F', dorsal view of abdominal 7 
<superScript id="86E92A7D1325FFF3FE6A5CE105A076DA" attach="left" box="[460,471,2018,2032]" fontSize="6" pageId="6" pageNumber="437">th</superScript>
sternite. Abbreviation: gw—guy-wire.
</paragraph>
</caption>
<paragraph id="712387351324FFF2FF615B9407F9719B" blockId="7.[151,1437,151,1149]" box="[199,910,151,177]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF615B940546719B" box="[199,305,151,177]" italics="true" pageId="7" pageNumber="438">Abdomen</emphasis>
(
<figureCitation id="E9A79BB01324FFF2FEE65B9405ED719B" box="[320,410,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1723,1747]" captionTargetBox="[151,1434,183,1682]" captionTargetId="figure-19@5.[151,1436,181,1699]" captionTargetPageId="5" captionText="FIGURE 3. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, adult; B, dorsal view of head and thorax; C, lateral view of head; D, antero-lateral view of head; E, anterior view of head; F, dorsal view of antenna; G, ventral view of antenna; G', close-up ventral view of partial antenna between base and basal ⅓; G'', close-up ventral view of partial antenna between basal ⅓ and ⅔; G''', close-up ventral view of partial antenna between basal ⅔ and apex, H, lateral view of hind legs." figureDoi="http://doi.org/10.5281/zenodo.6571735" httpUri="https://zenodo.org/record/6571735/files/figure.png" pageId="7" pageNumber="438">Fig. 3A</figureCitation>
). Abdomen covered with dark purple scales.
</paragraph>
<paragraph id="712387351324FFF2FF615BB8071371D3" blockId="7.[151,1437,151,1149]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF615BB8057F71FF" bold="true" box="[199,264,187,213]" pageId="7" pageNumber="438">Note:</emphasis>
The colour of wings under sunlight at the time of collection was bronze green, but after the specimen dried, no greenish color is observed, only bronze-orange colour.
</paragraph>
<paragraph id="712387351324FFF2FF615A0007F97381" blockId="7.[151,1437,151,1149]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF615A0005067037" box="[199,369,259,285]" italics="true" pageId="7" pageNumber="438">Male genitalia</emphasis>
(
<figureCitation id="E9A79BB01324FFF2FE225A0006737037" box="[388,516,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[180,1408,193,1787]" captionTargetId="figure-24@4.[164,1424,181,1798]" captionTargetPageId="4" captionText="FIGURE 2. Male (holotype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0141; B, dorsal view of genitalia; B', lateral view of genitalia; B'', ventral view of genitalia; C, juxta, gen. slide no. KJM0210; D, dorsal view of aedeagus; D', lateral view of anterior 2/5 of aedeagus; E, close-up of transtilla; E', close-up of left anterior process of transtilla; F, inner view of left valva. Abbreviations: ap—anterior process of transtilla, mp—medi-posterior process of transtilla, dc—dorsal corner of valva, tp—triangular process of valva, dm—distal margin of valva, vc—ventral corner of valva, vm—ventral margin of valva." figureDoi="http://doi.org/10.5281/zenodo.6571733" httpUri="https://zenodo.org/record/6571733/files/figure.png" pageId="7" pageNumber="438">Figs 2B–F</figureCitation>
): Uncus almost parallel-sided with concave posterior margin. Tegumen domeshaped, nearly of the same length as valva. Socius elliptical dome-shaped; the length of major axis about 1.9× longer than that of minor axis. Vinculum 2× longer than the length of valva, with slightly convex posterior margin; anterior margin obtuse; ventro-lateral margin dented 3 times in a row in posterior ¼ of vinculum; vinculum widest at posterior half; the longest axis of vinculum 1.7× longer than the widest axis. Transtilla with elongated triangular medi-posterior process and two anterior processes having rounded apex; medi-posterior process nearly of the same length as anterior process; anterior process slightly twisted. Valva trapezoidal, slightly convex in basal ⅔, then near straight distally; dorsal corner rounded with distinct triangular process; distal margin concave in dorsal ⅓, convex in ventral ⅔; ventral margin straight, 0.5× shorter than the length of valva with extremely pointed ventral corner. Juxta arrow-shaped, 0.5× shorter than the length of aedeagus; juxtal head shovelhead-shaped, about the same length as the width, with pointed barbs and extraordinarily obtuse apex. Aedeagus straight horn-shaped, of the same length as vinculum, slightly curved upwards towards anteriorly in basal ⅓.
</paragraph>
<paragraph id="712387351324FFF2FF6159B0076372AB" blockId="7.[151,1437,151,1149]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF6159B005F073E7" box="[199,391,691,717]" italics="true" pageId="7" pageNumber="438">Female genitalia</emphasis>
(
<figureCitation id="E9A79BB01324FFF2FE3059B0058573E7" box="[406,498,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="7" pageNumber="438">Figs 4C</figureCitation>
–D'). Apophyses posteriores nearly of the same length as apophyses anteriores. Vagina cup-shaped with one pair of guy-wires arising from one on each latero-anterior corner. Vestibulum without vestibulum lamella. Ductus seminalis arising from vestibulum, spiraled medially with internal canal. Spermatheca ovoidal, nearly of the same length and width as corpus bursae. Ductus bursae broadened from basal ¼ to half, then gradually narrower until distal ¼. Corpus bursae ovoidal, shorter about 0.5× than the length of ductus bursae. Common oviduct ventral, sharing its base with ductus bursae.
</paragraph>
<paragraph id="712387351324FFF2FF6158880625753B" blockId="7.[151,1437,151,1149]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF6158880662728E" box="[199,533,907,933]" italics="true" pageId="7" pageNumber="438">Female abdominal structures</emphasis>
(
<figureCitation id="E9A79BB01324FFF2FD81588806F3728F" box="[551,644,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1874,1898]" captionTargetBox="[219,1365,191,1843]" captionTargetId="figure-24@6.[208,1379,181,1850]" captionTargetPageId="6" captionText="FIGURE 4. Female (paratype) of Adela palella Koo &amp; Cho, sp. nov. A, venation, wings slide no. KJM0207; A', close-up of frenula (one frenulum missing); B, venation of hindwing, wings slide no. KJM0208; B', close-up of frenula; C, lateral view of genitalia, gen. slide no. KJM0211; D, dorsal view of genitalia, gen. slide no. KJM0212; D', close-up of vagina with two guywires; E, lateral view of abdominal 7th tergite; E', dorsal view of abdominal 7th tergite; F, lateral view of abdominal 7th sternite; F', dorsal view of abdominal 7th sternite. Abbreviation: gw—guy-wire." figureDoi="http://doi.org/10.5281/zenodo.6571737" httpUri="https://zenodo.org/record/6571737/files/figure.png" pageId="7" pageNumber="438">Figs 4E</figureCitation>
–F'): 7 
<superScript id="86E92A7D1324FFF2FD6B588906AD72B2" attach="left" box="[717,730,906,920]" fontSize="6" pageId="7" pageNumber="438">th</superScript>
tergite elongated triangular with rounded base angles; vertex angle pointing posteriorly. 7 
<superScript id="86E92A7D1324FFF2FE6958AD05AB7296" attach="left" box="[463,476,942,956]" fontSize="6" pageId="7" pageNumber="438">th</superScript>
sternite broad with setae on posterior margin, forming incomplete cylinder in posterior half, with sharply pointed posterior end. 8 
<superScript id="86E92A7D1324FFF2FDCE58D1060272CA" attach="left" box="[616,629,978,992]" fontSize="6" pageId="7" pageNumber="438">th</superScript>
segment roundly tapered arc-shaped, enclosed by 7 
<superScript id="86E92A7D1324FFF2FB1258D100B672CA" attach="left" box="[1204,1217,978,992]" fontSize="6" pageId="7" pageNumber="438">th</superScript>
tergite and sternite, ventro-medial anterior margin concave.
</paragraph>
</subSubSection>
<subSubSection id="3986D4BE1324FFF2FF615F190757751C" box="[199,800,1050,1078]" pageId="7" pageNumber="438" type="distribution">
<paragraph id="712387351324FFF2FF615F190757751C" blockId="7.[151,1437,151,1149]" box="[199,800,1050,1078]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF615F190516751E" bold="true" box="[199,353,1050,1076]" pageId="7" pageNumber="438">Distribution.</emphasis>
<collectingCountry id="098BC7A51324FFF2FEC15F180583751F" box="[359,500,1051,1077]" name="South Korea" pageId="7" pageNumber="438">South Korea</collectingCountry>
(Cheongju-si &amp; 
<collectingRegion id="B35849D71324FFF2FD175F180763751C" box="[689,788,1051,1078]" country="South Korea" name="Daejon" pageId="7" pageNumber="438">Daejeon</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="3986D4BE1324FFF2FF615F3C00627557" pageId="7" pageNumber="438" type="etymology">
<paragraph id="712387351324FFF2FF615F3C00627557" blockId="7.[151,1437,151,1149]" pageId="7" pageNumber="438">
<emphasis id="43E85B271324FFF2FF615F3C05397573" bold="true" box="[199,334,1087,1113]" pageId="7" pageNumber="438">Etymology.</emphasis>
The specific epithet is derived from the Latin, 
<emphasis id="43E85B271324FFF2FCF15F3C07FE7573" box="[855,905,1087,1113]" italics="true" pageId="7" pageNumber="438">pala</emphasis>
(=shovel, spade) with the Latin diminutive suffix, 
<emphasis id="43E85B271324FFF2FF625F60048F7557" box="[196,248,1123,1149]" italics="true" pageId="7" pageNumber="438">-ella</emphasis>
, referring to the shovelhead-shaped head of juxta in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>