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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.932.50314" ID-GBIF-Dataset="af05630e-cc44-4734-ac95-d28cd2a8d934" ID-PMC="PMC7237527" ID-Pensoft-Pub="1313-2970-932-1" ID-Pensoft-UUID="949CD324ED3652DEA8CA6B89F09F8E5E" ID-PubMed="32476972" ID-ZooBank="6C64E085E11A4AEE85EE3443E296C8DF" ModsDocID="1313-2970-932-1" checkinTime="1589335341202" checkinUser="pensoft" docAuthor="Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K. &amp; Nguyen, Anh D." docDate="2020" docId="BA7F06852F4C5D538BD32FF8DBC149CF" docLanguage="en" docName="ZooKeys 932: 1-25" docOrigin="ZooKeys 932" docSource="http://dx.doi.org/10.3897/zookeys.932.50314" docTitle="Amynthas phuquocensis Nguyen &amp; Lam &amp; Trinh &amp; Nguyen 2020, sp. nov." docType="treatment" docUuid="8165ADE3-0638-4DAF-8BA3-78092439F7B5" docUuidSource="ZooBank" docVersion="5" id="949CD324ED3652DEA8CA6B89F09F8E5E" lastPageNumber="1" masterDocId="949CD324ED3652DEA8CA6B89F09F8E5E" masterDocTitle="The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="1" updateTime="1668168819817" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Tung T.</mods:namePart>
<mods:affiliation>Department of Biology, School of Education, Can Tho University, Can Tho City, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lam, Dang H.</mods:namePart>
<mods:affiliation>Department of Biology, School of Education, Can Tho University, Can Tho City, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Trinh, Binh T. K.</mods:namePart>
<mods:affiliation>Department of Applied Biology, Faculty of Agriculture and Rural Development, Kien Giang University, Kien Giang, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Anh D.</mods:namePart>
<mods:affiliation>Duy Tan University, 254, Nguyen Van Linh, Da Nang, Vietnam &amp; Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier type="email">ducanh410@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>932</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.932.50314</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.932.50314</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-932-1</mods:identifier>
<mods:identifier type="ZooBank">6C64E085E11A4AEE85EE3443E296C8DF</mods:identifier>
<mods:identifier type="Pensoft-UUID">949CD324ED3652DEA8CA6B89F09F8E5E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="164127903" LSID="urn:lsid:zoobank.org:act:8165ADE3-0638-4DAF-8BA3-78092439F7B5" httpUri="http://treatment.plazi.org/id/BA7F06852F4C5D538BD32FF8DBC149CF" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="BA7F0685-2F4C-5D53-8BD3-2FF8DBC149CF" authority="Nguyen &amp; Lam &amp; Trinh &amp; Nguyen, 2020" authorityName="Nguyen &amp; Lam &amp; Trinh &amp; Nguyen" authorityYear="2020" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas phuquocensis" order="Opisthopora" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="phuquocensis" status="sp. nov.">Amynthas phuquocensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Variation of genital markings in the male region of A. phuquocensis sp. nov." figureDoi="10.3897/zookeys.932.50314.figure12" httpUri="https://binary.pensoft.net/fig/409118" pageId="0" pageNumber="1">Figs 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A. phuquocsensis sp. nov. Holotype (CTU-EW. 190. h 01) A 1, A 2 Male region, ventral view B 1, B 2 right spermathecae C 1, C 2 spermathecal region, ventral view D testis sacs and vesicles E intestinal caecum F 1, F 2 right prostatic gland G transection via male porophore H transection via genital marking. Scale bar: 1 mm." figureDoi="10.3897/zookeys.932.50314.figure13" httpUri="https://binary.pensoft.net/fig/409119" pageId="0" pageNumber="1">, 13</figureCitation>
,
<figureCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Character comparison between A. phuquocensis sp. nov., A. phimpheti and A. antethecus." pageId="0" pageNumber="1">Tables 2</figureCitation>
,
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Character comparison between A. phuquocensis sp. nov., A. binhgiaensis, A. platycorpus." pageId="0" pageNumber="1">3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: 1 mature (CTU-EW.188.h01), Kien Giang Province, Phu Quoc island, natural forests (
<geoCoordinate degrees="10" direction="north" minutes="20" orientation="latitude" precision="1" seconds="51.3" value="10.347583">10°20'51.3&quot;N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="58" orientation="longitude" precision="1" seconds="58.0" value="103.98278">103°58'58.0&quot;E</geoCoordinate>
), 44.2 m, 7 November 2016, coll. Lam HD &amp; Trinh TKB.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 9 matures (CTU-EW.188.p02) same data as for holotype.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Non-types</emphasis>
: 9 matures (CTU-EW.188.03) same data as for holotype; 4 matures (CTU-EW.188.04), Kien Giang Province, Phu Quoc island, natural forests (
<geoCoordinate degrees="10" direction="north" minutes="20" orientation="latitude" precision="15" seconds="50" value="10.347222">10°20'50&quot;N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="55" orientation="longitude" precision="15" seconds="07" value="103.91861">103°55'07&quot;E</geoCoordinate>
), 57 m, 7 November 2016, coll. Lam HD &amp; Trinh TKB.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Medium-sized, length 74-145 mm, average diameter 2.5-4.7 mm, segments 116-145. Prostomium epilobous. Two pairs of spermathecal pores in ventral intersegments 7/8/9. Male pores in xviii, without copulatory pouches. Two pairs of genital markings present in ventral 17/18 and 18/19. Intestinal caeca simple. Holandric. Testis sacs in xi, separated. Septa 8/9/10/11 absent.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
&quot;
<taxonomicName authorityName="Nguyen &amp; Lam &amp; Trinh &amp; Nguyen" authorityYear="2020" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="phuquocensis" order="Opisthopora" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="phuquocensis">
<emphasis italics="true" pageId="0" pageNumber="1">phuquocensis</emphasis>
</taxonomicName>
&quot;, named for the type locality.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Body cylindrical, medium size, length 74-145 mm, average diameter 2.5-4.7 mm, segments 116-145, weight 0.6-3.7 g. Body uniformly greyish brown except darkish brown clitellum. Prostomium 2/3 epilobous. First dorsal pore in 12/13. Pre-clitellum setae stouter and sparser than post-clitellum ones; setal numbers: 42-61 in viii, 50-77 in xxx, 5-9 between male porophores in xviii; setal distance: aa&gt; ab, zz&gt; zy. Clitellum within xiv-xvi, without setae and dorsal pores. Female pore single, in mid-ventral xiv.</paragraph>
<paragraph pageId="0" pageNumber="1">
Two pairs of spermathecal pores in ventral intersegments 7/8/9. Genital markings absent in the spermathecal region. Male pores located in the setal ring xviii, without copulatory pouches; ventral distance between two male pores about 0.28
<normalizedToken originalValue="×">x</normalizedToken>
body circumference. Genital markings roundly pad-shaped and variable in the male region, normally with two pairs in 17/18 and 18/19, or a pair in xix (near 19/20).
</paragraph>
<paragraph pageId="0" pageNumber="1">Septa 5/6/7/8 thick, 8/9/10/11 absent, 11/12/13 thick. Oesophageal gizzard within viii-xi. Intestinal origin at xv. Intestinal caeca simple, within xxvii-xxiv. Last hearts in xiii. Pharyngeal micronephridia in 5/6/7. Typhlosole lamelliform. Lymph lobuled, from 15/16.</paragraph>
<paragraph pageId="0" pageNumber="1">Two pairs of spermathecae in viii and ix. Ampulla large, heart-shaped with transverse wrinkles; ducts short and stout. Diverticula waved, attached to the base of ampulla; seminal chamber ellipsoid. Accessory glands bean-shaped, surrounding spermathecal ducts.</paragraph>
<paragraph pageId="0" pageNumber="1">Holandric. Testis sacs in xi, separated, the anterior pair poorly developed. Seminal vesicles well developed in xi and xii. Ovaries in 12/13. Ovisacs invisible. Prostate glands deeply lobuled in xvi-xx; ducts strongly coiled. Accessory glands present.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitats.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was occasionally found in natural forests along the main road to the northern part of Phu Quoc island. They were collected in the upper soil layer (0-10cm).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="vietnamese name">
<paragraph pageId="0" pageNumber="1">Vietnamese name.</paragraph>
<paragraph pageId="0" pageNumber="1">
Giun
<normalizedToken originalValue="phú">phu</normalizedToken>
quốc.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variations.</paragraph>
<paragraph pageId="0" pageNumber="1">
The body size and number of genital markings in the male region are variable among collected specimens. Specimens can be divided into two groups based on body size. The group of smaller specimens (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 15) present the following ranges: length = 71-85 mm, diameter = 2.5-2.9 mm, segments = 116-145. The larger specimens (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 8) present: length = 96-145 mm, diameter = 3.1-4.7 mm, segments = 124-145. Both two groups were together at the same site.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Genital markings are also variable in the male region. Genital markings have two pairs located in 17/18 and 18/19 (Type 1), but reduced to three genital markings (Type 2), one pair either in xvii or in xiv (Type 3 or Type 4), or a single one in xix (Type 5) (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Variation of genital markings in the male region of A. phuquocensis sp. nov." figureDoi="10.3897/zookeys.932.50314.figure12" httpUri="https://binary.pensoft.net/fig/409118" pageId="0" pageNumber="1">12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The new species can be assigned into the
<taxonomicName class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amynthas aeruginosus" order="Opisthopora" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="aeruginosus">
<emphasis italics="true" pageId="0" pageNumber="1">Amynthas aeruginosus</emphasis>
</taxonomicName>
group, which is characterized by having two pairs of spermathecal pores in 7/8/9 and holandric (
<bibRefCitation DOI="https://doi.org/10.1111/j.1095-8312.1972.tb00694.x" author="Sims, RW" journalOrPublisher="Biological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="169 - 268" refId="B19" refString="Sims, RW, Easton, EG, 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society 4: 169 - 268, DOI: https://doi.org/10.1111/j.1095-8312.1972.tb00694.x" title="A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition." url="https://doi.org/10.1111/j.1095-8312.1972.tb00694.x" volume="4" year="1972">Sims and Easton 1972</bibRefCitation>
). This group is known to have about 70 species distributed mainly in Southeast and East Asia. Of these, about 11 species have been recorded in Vietnam (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4140.1.1" author="Nguyen, TT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 92" refId="B14" refString="Nguyen, TT, Nguyen, AD, Tran, TT, Blakemore, RJ, 2016. A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae). Zootaxa 4140: 1 - 92, DOI: https://doi.org/10.11646/zootaxa.4140.1.1" title="A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae)." url="https://doi.org/10.11646/zootaxa.4140.1.1" volume="4140" year="2016">Nguyen et al. 2016</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Within the
<taxonomicName lsidName="A. aeruginosus" pageId="0" pageNumber="1" rank="species" species="aeruginosus">
<emphasis italics="true" pageId="0" pageNumber="1">A. aeruginosus</emphasis>
</taxonomicName>
group, the new species is fairly similar to
<taxonomicName lsidName="A. phimpheti" pageId="0" pageNumber="1" rank="species" species="phimpheti">
<emphasis italics="true" pageId="0" pageNumber="1">A. phimpheti</emphasis>
</taxonomicName>
Hong, Inkavilay &amp; James, 2018 and
<taxonomicName lsidName="A. antethecus" pageId="0" pageNumber="1" rank="species" species="antethecus">
<emphasis italics="true" pageId="0" pageNumber="1">A. antethecus</emphasis>
</taxonomicName>
Hong, Inkavilay &amp; James, 2018 in having two pairs of genital markings in the male region, but absent in the spermathecal region, and a simple intestinal caeca. However, these species can be distinguished by the positions of the first dorsal pore and the spermathecal pores and genital markings in the male region, the presence of septum 10/11, presence of accessory glands, and position of testis sacs (summarized in Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Character comparison between A. phuquocensis sp. nov., A. phimpheti and A. antethecus." httpUri="http://table.plazi.org/id/4B9E76BD4E6E1CC9774E27AE95BEE428" pageId="0" pageNumber="1" tableUuid="4B9E76BD4E6E1CC9774E27AE95BEE428">2</tableCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Regarding the arrangement of genital markings in the male region, the new species is slightly similar to
<taxonomicName lsidName="A. platycorpus" pageId="0" pageNumber="1" rank="species" species="platycorpus">
<emphasis italics="true" pageId="0" pageNumber="1">A. platycorpus</emphasis>
</taxonomicName>
(Thai, 1982) and
<taxonomicName lsidName="A. binhgiaensis" pageId="0" pageNumber="1" rank="species" species="binhgiaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. binhgiaensis</emphasis>
</taxonomicName>
(Le, 1994) by having two pairs of genital markings in 17/18 and 18/19. However, these species can be distinguished by number and position of spermathecal pores, first dorsal pore, genital markings in the spermathecal region, presence of septum 10/11, type of intestinal caeca and testis sacs (summarized in Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Character comparison between A. phuquocensis sp. nov., A. binhgiaensis, A. platycorpus." httpUri="http://table.plazi.org/id/92EAA1787FE7882E8C6C90321D28ADCB" pageId="0" pageNumber="1" tableUuid="92EAA1787FE7882E8C6C90321D28ADCB">3</tableCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.932.50314.figure12" httpUri="https://binary.pensoft.net/fig/409118" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
Variation of genital markings in the male region of
<taxonomicName lsidName="A. phuquocensis" pageId="0" pageNumber="1" rank="species" species="phuquocensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. phuquocensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.932.50314.figure13" httpUri="https://binary.pensoft.net/fig/409119" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<taxonomicName lsidName="A. phuquocsensis" pageId="0" pageNumber="1" rank="species" species="phuquocsensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. phuquocsensis</emphasis>
</taxonomicName>
sp. nov. Holotype (CTU-EW.190.h01)
<emphasis bold="true" pageId="0" pageNumber="1">A1, A2</emphasis>
Male region, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">B1, B2</emphasis>
right spermathecae
<emphasis bold="true" pageId="0" pageNumber="1">C1, C2</emphasis>
spermathecal region, ventral view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
testis sacs and vesicles
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
intestinal caecum
<emphasis bold="true" pageId="0" pageNumber="1">F1, F2</emphasis>
right prostatic gland
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
transection via male porophore
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
transection via genital marking. Scale bar: 1mm.
</paragraph>
</caption>
<caption ID-Table-UUID="4B9E76BD4E6E1CC9774E27AE95BEE428" httpUri="http://table.plazi.org/id/4B9E76BD4E6E1CC9774E27AE95BEE428" pageId="0" pageNumber="1" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 2.</emphasis>
Character comparison between
<taxonomicName lsidName="A. phuquocensis" pageId="0" pageNumber="1" rank="species" species="phuquocensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. phuquocensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="A. phimpheti" pageId="0" pageNumber="1" rank="species" species="phimpheti">
<emphasis italics="true" pageId="0" pageNumber="1">A. phimpheti</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. antethecus" pageId="0" pageNumber="1" rank="species" species="antethecus">
<emphasis italics="true" pageId="0" pageNumber="1">A. antethecus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="A. phuquocensis" pageId="0" pageNumber="1" rank="species" species="phuquocensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. phuquocensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<taxonomicName lsidName="A. phimpheti" pageId="0" pageNumber="1" rank="species" species="phimpheti">A. phimpheti</taxonomicName>
*
</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<taxonomicName lsidName="A. antethecus" pageId="0" pageNumber="1" rank="species" species="antethecus">A. antethecus</taxonomicName>
*
</emphasis>
</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Length (mm)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">118-145</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">76-100</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">88-122</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Diameter (mm)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.7-4.7</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.2-4.8</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.0-4.1</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Segments</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">143-145</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">99-104</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">82-109</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Spermathecal pores</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">lateroventral</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">lateroventral</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ventral</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">First dorsal pore</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12/13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5/6</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5/6</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">GM in male region</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17/18 and 18/19 or xvii or xix</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17/18 and 18/19</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17/18 and xix</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Septum 10/11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Testis sacs</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">separated in xi</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">connected, x and xi</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">connected, x and xi</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Accessory glands in spermathecal region</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">surrounding spermathecal ducts</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">no</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Accessory glands in male region</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<tableNote pageId="0" pageNumber="1">
Source: *:
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4496.1.19" author="Hong, Y" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="238 - 250" refId="B8" refString="Hong, Y, Inkhavilay, K, James, SW, 2018. Nine new species of the aeruginosus -group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos. Zootaxa 4496 (1): 238 - 250, DOI: https://doi.org/10.11646/zootaxa.4496.1.19" title="Nine new species of the aeruginosus - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos." url="https://doi.org/10.11646/zootaxa.4496.1.19" volume="4496" year="2018">Hong et al. (2018)</bibRefCitation>
.
</tableNote>
</paragraph>
<caption ID-Table-UUID="92EAA1787FE7882E8C6C90321D28ADCB" httpUri="http://table.plazi.org/id/92EAA1787FE7882E8C6C90321D28ADCB" pageId="0" pageNumber="1" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 3.</emphasis>
Character comparison between
<taxonomicName lsidName="A. phuquocensis" pageId="0" pageNumber="1" rank="species" species="phuquocensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. phuquocensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="A. binhgiaensis" pageId="0" pageNumber="1" rank="species" species="binhgiaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. binhgiaensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="A. platycorpus" pageId="0" pageNumber="1" rank="species" species="platycorpus">
<emphasis italics="true" pageId="0" pageNumber="1">A. platycorpus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="A. phuquocensis" pageId="0" pageNumber="1" rank="species" species="phuquocensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. phuquocensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="A. binhgiaensis" pageId="0" pageNumber="1" rank="species" species="binhgiaensis">
<emphasis italics="true" pageId="0" pageNumber="1">A. binhgiaensis</emphasis>
</taxonomicName>
1
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="A. platycorpus" pageId="0" pageNumber="1" rank="species" species="platycorpus">
<emphasis italics="true" pageId="0" pageNumber="1">A. platycorpus</emphasis>
</taxonomicName>
2
</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Length (mm)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">168-220</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">72</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">38-45</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Diameter (mm)</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.50-7.58</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3-4</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2-3</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Segments</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">99-133</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">100</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40-51</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Spermathecal pores</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6/7/8/9, lateroventral</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5/6/7/8, dorsal</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6/7/8, lateral</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">First dorsal pore</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12/13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12/13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4/5</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">GM in spermathecal region</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4 pairs in vi-ix</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Septum 10/11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Intestinal caeca</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">lobuled</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">simple</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">manicate</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Male sexual system</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">holandric</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">holandric</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">holandric</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Testis sacs</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">xi, separated</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">x, xi, separated</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">x, xi, connected</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<tableNote pageId="0" pageNumber="1">
Source: 1:
<bibRefCitation author="Do, VN" journalOrPublisher="Tap Chi Sinh Hoc" pageId="0" pageNumber="1" pagination="22 - 28" refId="B3" refString="Do, VN, Le, VT, Tran, MK, 1994. New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from Lang Son and Nghe An Provinces. Tap Chi Sinh Hoc 16 (1): 22 - 28" title="New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from Lang Son and Nghe An Provinces." volume="16" year="1994">Do et al. (1994)</bibRefCitation>
; 2:
<bibRefCitation author="Thai, TB" journalOrPublisher="Zoologicheskii Zhurnal" pageId="0" pageNumber="1" pagination="817 - 830" refId="B20" refString="Thai, TB, 1982. The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Zhurnal 61 (6): 817 - 830" title="The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima." volume="61" year="1982">Thai (1982)</bibRefCitation>
.
</tableNote>
</paragraph>
</subSubSection>
</treatment>
</document>
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