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<mods:title id="84A71876C0A619739E947D1F31DFA779">A taxonomic revision of the Cardiocondyla nuda group (Hymenoptera: Formicidae)</mods:title>
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<taxonomicName id="C8E00693FFF2FFDA30EFD84AFDDDF9FC" authority="Seifert 2003" authorityName="Seifert" authorityYear="2003" box="[151,609,1540,1566]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="346" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFF2FFDA30EFD84AFDDDF9FC" bold="true" box="[151,609,1540,1566]" pageId="22" pageNumber="346">
<emphasis id="3D94A102FFF2FFDA30EFD84AFE79F9FC" bold="true" box="[151,453,1540,1566]" italics="true" pageId="22" pageNumber="346">Cardiocondyla paranuda</emphasis>
<bibRefCitation id="6B7100E1FFF2FFDA31B4D84AFDDDF9FC" author="Seifert" box="[460,609,1540,1566]" pageId="22" pageNumber="346" refString="Seifert, B. (2003) The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae) - a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi and C. minutior species groups. Annals of the Natural History Museum Wien, 104 (B), 203 - 338." type="journal article" year="2003">Seifert 2003</bibRefCitation>
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<tableCitation id="426248ABFFF2FFDA30EFD868FF62F9DD" box="[151,222,1574,1599]" captionStart="TABLE 2" captionStartId="22.[151,239,152,175]" captionText="TABLE 2. Morphometric data of workers of C. nuda species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals. Data with reduced number of measured individuals in C. nuda are given in italics." pageId="22" pageNumber="346">Tab. 2</tableCitation>
,
<figureCitation id="97DB6195FFF2FFDA3092D868FED1F9DD" box="[234,365,1574,1599]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="23.[151,250,1258,1280]" captionStartId-1="24.[151,250,1213,1235]" captionStartId-2="25.[151,250,1226,1248]" captionTargetBox-0="[276,1303,179,1233]" captionTargetBox-1="[340,1247,195,1191]" captionTargetBox-2="[151,1436,195,1205]" captionTargetId-0="figure@23.[276,1303,179,1233]" captionTargetId-1="figure@24.[340,1247,195,1191]" captionTargetId-2="figure@25.[151,1436,195,1205]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURE 14. Gaster of Cardiocondyla paranuda in dorsal aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." captionText-1="FIGURE 15. Head of Cardiocondyla paranuda in dorsal aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." captionText-2="FIGURE 16. Cardiocondyla paranuda in lateral aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." httpUri-0="https://zenodo.org/record/829099/files/figure.png" httpUri-1="https://zenodo.org/record/829101/files/figure.png" httpUri-2="https://zenodo.org/record/829103/files/figure.png" pageId="22" pageNumber="346">
Figs. 14
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16
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<emphasis id="3D94A102FFF2FFDA30EFD820FE26F966" box="[151,410,1646,1668]" italics="true" pageId="22" pageNumber="346">Cardiocondyla paranuda</emphasis>
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: 246
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.
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worker: alledgedly
<collectingCountry id="77F73D80FFF2FFDA33E2D820FC54F966" box="[922,1000,1646,1668]" name="Tunisia" pageId="22" pageNumber="346">Tunisia</collectingCountry>
,
<location id="0A3F2BCBFFF2FFDA338DD820FBE9F966" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFF2FFDA338DD820FBE9F966" box="[1013,1109,1646,1668]" country="Tunisia" name="Chabania" pageId="22" pageNumber="346">Chabania</location>
[
<collectionCode id="69F1E5D5FFF2FFDA341BD820FB12F966" box="[1123,1198,1646,1668]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="22" pageNumber="346">SMNG</collectionCode>
, antweb.org images of specimen
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<collectionCode id="69F1E5D5FFF2FFDA314ED8C3FE30F941" box="[310,396,1677,1699]" pageId="22" pageNumber="346">FOCOL</collectionCode>
0739
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] (examined)
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.
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<typeStatus id="D05BC3B2FFF2FFDA3224D8C3FD06F941" box="[604,698,1677,1699]" pageId="22" pageNumber="346" type="holotype">Holotype</typeStatus>
labels &quot;
<collectingCountry id="77F73D80FFF2FFDA3364D8C3FC3BF941" box="[796,903,1677,1699]" name="Tunisia" pageId="22" pageNumber="346">TUNISIA</collectingCountry>
: Medinine-
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SE Chabania-
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NW leg.
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1976“, &quot;
<typeStatus id="D05BC3B2FFF2FFDA31D4D8E2FDB0F920" box="[428,524,1708,1730]" pageId="22" pageNumber="346" type="holotype">Holotype</typeStatus>
<taxonomicName id="C8E00693FFF2FFDA3264D8E2FCC0F920" authority="Seifert" authorityName="Seifert" box="[540,892,1708,1730]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="346" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFF2FFDA3264D8E2FC99F920" box="[540,805,1708,1730]" italics="true" pageId="22" pageNumber="346">Cardiocondyla paranuda</emphasis>
Seifert
</taxonomicName>
&quot;, &quot;GBIF-D/
<collectorName id="A21518C6FFF2FFDA3384D8E2FB80F920" box="[1020,1084,1708,1730]" pageId="22" pageNumber="346">FoCol</collectorName>
0
<specimenCount id="19E6B699FFF2FFDA3420D8E2FB51F920" box="[1112,1261,1708,1730]" pageId="22" pageNumber="346" type="generic">739 specimen</specimenCount>
and label data documented&quot;, and &quot;
<bibRefCitation id="6B7100E1FFF2FFDA31EDD884FD90F902" author="Seifert" box="[405,556,1738,1760]" pageId="22" pageNumber="346" refString="Seifert, B., d'Eustaccio, D., Kaufmann, B., Centorame, M., Lorite, P. &amp; Modica, M. V. (2017) Four cryptic species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News, 24, 123 - 144." type="journal article" year="2017">Seifert (2017)</bibRefCitation>
: Confusion of label by Collingwood. Terra Typica by morphometric analysis most probably Australia&quot;.
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<emphasis id="3D94A102FFF2FFDA30EFD962FEC5F8A7" bold="true" box="[151,377,1836,1861]" pageId="22" pageNumber="346">Material examined</emphasis>
A total of 30 nest samples with 52 workers were subject to NUMOBAT investigation.
</paragraph>
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<emphasis id="3D94A102FFF3FFDB30EFDAA4FE9FFB1D" bold="true" box="[151,291,1258,1280]" pageId="23" pageNumber="347">FIGURE 14.</emphasis>
Gaster of
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<emphasis id="3D94A102FFF3FFDB31EFDAA4FD26FAE2" box="[407,666,1258,1280]" italics="true" pageId="23" pageNumber="347">Cardiocondyla paranuda</emphasis>
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in dorsal aspect. Australia: Kunoth Paddock [23.517°S, 133.210°E], leg. Greenslade 24 October 1974.
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<emphasis id="3D94A102FFF3FFDB30BFDB0BFE8BFABC" bold="true" box="[199,311,1349,1374]" pageId="23" pageNumber="347">Australia</emphasis>
:
<materialsCitation id="BF88774DFFF3FFDB313EDB0BFC75FABC" ID-GBIF-Occurrence="1572325952" box="[326,969,1349,1374]" country="Australia" location="Australia" pageId="23" pageNumber="347" specimenCount="1" typeStatus="holotype">
<collectingCountry id="77F73D80FFF3FFDB313EDB0BFE0DFABC" box="[326,433,1349,1374]" name="Australia" pageId="23" pageNumber="347">Australia</collectingCountry>
: without site and date,
<typeStatus id="D05BC3B2FFF3FFDB32CCDB0BFCAAFABC" box="[692,790,1349,1374]" pageId="23" pageNumber="347" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C8E00693FFF3FFDB3343DB08FC75FABC" box="[827,969,1349,1374]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="347" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFF3FFDB3343DB08FC75FABC" box="[827,969,1349,1374]" italics="true" pageId="23" pageNumber="347">C. paranuda</emphasis>
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;
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<collectingRegion id="CD24B3F2FFF3FFDB33A0DB09FB19FABC" box="[984,1189,1349,1374]" country="Australia" name="New South Wales" pageId="23" pageNumber="347">
New
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: Barham, 1960.03.23, [-35.62, 144.15]
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;
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New
<collectingRegion id="CD24B3F2FFF3FFDB31E0DB24FD94FA61" box="[408,552,1386,1411]" inferredBySuffix="true" pageId="23" pageNumber="347">South Wales</collectingRegion>
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:
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, 1991.02.16, [-34.53, 150.25]
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;
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<collectingRegion id="CD24B3F2FFF3FFDB34FBDB25FAECFA61" box="[1155,1360,1386,1411]" country="Australia" name="New South Wales" pageId="23" pageNumber="347">
New
<collectingRegion id="CD24B3F2FFF3FFDB34B8DB24FAECFA61" box="[1216,1360,1386,1411]" inferredBySuffix="true" pageId="23" pageNumber="347">South Wales</collectingRegion>
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:
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, 1997.xx.xx, [-35.28, 149.09]
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;
<materialsCitation id="BF88774DFFF3FFDB3209DBC1FA24FA44" ID-GBIF-Occurrence="1572325946" box="[625,1432,1421,1446]" country="Australia" location="New South Wales" pageId="23" pageNumber="347" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB3209DBC1FC83FA44" box="[625,831,1421,1446]" country="Australia" name="New South Wales" pageId="23" pageNumber="347">
New
<collectingRegion id="CD24B3F2FFF3FFDB32D5DBC3FC83FA44" box="[685,831,1421,1446]" inferredBySuffix="true" pageId="23" pageNumber="347">South Wales</collectingRegion>
</collectingRegion>
: Broken Hill, Parkland, 1971.05.18, [-31.96, 141,46]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB30EFDBFDFCE4FA29" ID-GBIF-Occurrence="1572325959" box="[151,856,1458,1483]" country="Australia" location="New South Wales" pageId="23" pageNumber="347" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB30EFDBFDFED9FA29" box="[151,357,1458,1483]" country="Australia" name="New South Wales" pageId="23" pageNumber="347">
New
<collectingRegion id="CD24B3F2FFF3FFDB30ACDBFCFED9FA29" box="[212,357,1458,1483]" inferredBySuffix="true" pageId="23" pageNumber="347">South Wales</collectingRegion>
</collectingRegion>
: Fowlers Gap, 1979.02.19, [-31.02, 146.60]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB331CDBFDFC33FA0C" ID-GBIF-Occurrence="1572325958" country="Australia" location="Lake Menindee" pageId="23" pageNumber="347" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB331CDBFDFB8EFA29" box="[868,1074,1458,1483]" country="Australia" name="New South Wales" pageId="23" pageNumber="347">
New
<collectingRegion id="CD24B3F2FFF3FFDB33D9DBFCFB8EFA29" box="[929,1074,1458,1483]" inferredBySuffix="true" pageId="23" pageNumber="347">South Wales</collectingRegion>
</collectingRegion>
:
<location id="0A3F2BCBFFF3FFDB3446DBFCFB4DFA29" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFF3FFDB3446DBFCFB4DFA29" box="[1086,1265,1458,1483]" country="Australia" name="Lake Menindee" pageId="23" pageNumber="347" stateProvince="New South Wales">Lake Menindee</location>
, 1971.05.19, [- 32.32, 142.40]; Sydney, Concord, 1960.05.0 1, [-33.86, 151.10]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB33DADB9BFD0CF9F1" ID-GBIF-Occurrence="1572325955" collectorName="No I" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB33DADB9BFBC2FA0C" box="[930,1150,1493,1518]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB33DADB9BFBBBFA0C" box="[930,1031,1493,1518]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
: Alice Springs, Kunoth Paddock, 1974.10.22,
<collectorName id="A21518C6FFF3FFDB31EDDBB5FE77F9F0" box="[405,459,1531,1554]" pageId="23" pageNumber="347">No I</collectorName>
, [-23.517, 133.583]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB32C6DBB4FE08F9D4" ID-GBIF-Occurrence="1572325949" collectorName="No I" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB32C6DBB4FC2FF9F1" box="[702,915,1530,1555]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB32C6DBB4FC9FF9F1" box="[702,803,1530,1555]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
: Alice Springs, Kunoth Paddock, 1974.10.24,
<collectorName id="A21518C6FFF3FFDB30EFD851FF71F9D4" box="[151,205,1567,1590]" pageId="23" pageNumber="347">No I</collectorName>
, [-23.517, 133.583]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB31B9D853FB3EF9D4" ID-GBIF-Occurrence="1572325965" box="[449,1154,1565,1590]" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB31B9D853FD2BF9D4" box="[449,663,1565,1590]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB31B9D853FD9AF9D4" box="[449,550,1565,1590]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
: Ayers Rock, 1981.10.xx, [-25.35, 131.03]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB34F7D853FD7FF9B8" ID-GBIF-Occurrence="1572325960" collectorName="SW Katherine" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB34F7D853FADAF9D4" box="[1167,1382,1565,1590]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB34F7D853FB48F9D4" box="[1167,1268,1565,1590]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
:
<collectorName id="A21518C6FFF3FFDB350AD850FEB4F9B9" pageId="23" pageNumber="347">SW Katherine</collectorName>
, Manaulloo, 1978.04.xx, [-14.5, 132.2]
</materialsCitation>
,
<materialsCitation id="BF88774DFFF3FFDB32B7D80CFA24F9B8" ID-GBIF-Occurrence="1572325943" box="[719,1432,1602,1627]" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB32B7D80CFC1FF9B9" box="[719,931,1602,1627]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB32B7D80CFC88F9B9" box="[719,820,1602,1627]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
: Simpson Gap, 1972.xx.xx, [-23.71, 133.71]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB30EFD82BFC51F99C" ID-GBIF-Occurrence="1572325961" box="[151,1005,1637,1662]" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB30EFD82BFECDF99C" box="[151,369,1637,1662]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB30EFD82BFF40F99C" box="[151,252,1637,1662]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
: Ti Tree Well-
<quantity id="C818D0F5FFF3FFDB325DD82BFDD1F99C" box="[549,621,1637,1662]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="347" unit="km" value="11.0">11 km</quantity>
S, 1962.10.28, [-22.26, 133.38]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB3386D82BFD86F941" ID-GBIF-Occurrence="1572325957" country="Australia" location="Northern Territory" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB3386D82BFB64F99C" box="[1022,1240,1637,1662]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB3386D82BFBDFF99C" box="[1022,1123,1637,1662]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
: above Baroalba springs, 1972.11.17, [-12.47, 132.51]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB323DD8C4FE29F924" ID-GBIF-Occurrence="1572325947" country="Australia" location="Yulara" municipality="No" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB323DD8C4FCA6F941" box="[581,794,1674,1699]" country="Australia" name="Northern Territory" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB323DD8C4FD16F941" box="[581,682,1674,1699]" country="Sri Lanka" name="Northern" pageId="23" pageNumber="347">Northern</collectingRegion>
Territory
</collectingRegion>
:
<location id="0A3F2BCBFFF3FFDB335CD8C4FCCCF941" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFF3FFDB335CD8C4FCCCF941" box="[804,880,1674,1699]" country="Australia" municipality="No" name="Yulara" pageId="23" pageNumber="347" stateProvince="Northern Territory">Yulara</location>
, 2014.07.27,
<collectingMunicipality id="EF3BE76AFFF3FFDB347FD8C5FB95F940" box="[1031,1065,1675,1698]" pageId="23" pageNumber="347">No</collectingMunicipality>
AUS39 (GenBank
<accessionNumber id="10B3E0F3FFF3FFDB357DD8C4FAC7F940" box="[1285,1403,1674,1698]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LT718213" pageId="23" pageNumber="347">LT718213</accessionNumber>
) [- 25.24361, 130.98639]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB31DDD8E3FBF9F924" ID-GBIF-Occurrence="1572325986" box="[421,1093,1709,1734]" country="Iceland" location="Queensland" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB31DDD8E3FD92F924" box="[421,558,1709,1734]" country="Australia" name="Queensland" pageId="23" pageNumber="347">Queensland</collectingRegion>
: Chilcott
<collectingCountry id="77F73D80FFF3FFDB32D9D8E3FD54F924" box="[673,744,1709,1734]" name="Iceland" pageId="23" pageNumber="347">Island</collectingCountry>
, 1967.08.xx, [-16.95, 149.91]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB342CD8E3FE45F909" ID-GBIF-Occurrence="1572325971" country="Iceland" location="Queensland" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB342CD8E3FB61F924" box="[1108,1245,1709,1734]" country="Australia" name="Queensland" pageId="23" pageNumber="347">Queensland</collectingRegion>
: Chilcott
<collectingCountry id="77F73D80FFF3FFDB3529D8E3FA24F924" box="[1361,1432,1709,1734]" name="Iceland" pageId="23" pageNumber="347">Island</collectingCountry>
, 1967.08.xx, [-16.25, 150.00]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB326AD89CFB4BF908" ID-GBIF-Occurrence="1572325987" box="[530,1271,1746,1771]" country="Iceland" location="Queensland" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB326AD89CFD27F909" box="[530,667,1746,1771]" country="Australia" name="Queensland" pageId="23" pageNumber="347">Queensland</collectingRegion>
: Coongie-
<quantity id="C818D0F5FFF3FFDB3364D89CFCCCF909" box="[796,880,1746,1771]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="347" unit="km" value="25.0">25 km</quantity>
S, 1975.08.xx, [-27.5, 140.0]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB3568D89CFDC3F8EC" ID-GBIF-Occurrence="1572325980" country="Iceland" location="Queensland" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB3568D89CFA25F909" box="[1296,1433,1746,1771]" country="Australia" name="Queensland" pageId="23" pageNumber="347">Queensland</collectingRegion>
: Cunnamulla, 1974.09.17, [-28.070, 145.67]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB32F2D8BBFAECF8EC" ID-GBIF-Occurrence="1572325979" box="[650,1360,1781,1806]" country="Iceland" location="Queensland" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB32F2D8BBFCAFF8EC" box="[650,787,1781,1806]" country="Australia" name="Queensland" pageId="23" pageNumber="347">Queensland</collectingRegion>
: Woodstock-
<quantity id="C818D0F5FFF3FFDB33DDD8BBFC56F8EC" box="[933,1002,1781,1806]" metricMagnitude="4" metricUnit="m" metricValue="5.2" pageId="23" pageNumber="347" unit="km" value="52.0">52 km</quantity>
S, 1976.04.11, [-20.07, 146.82]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB3523D8BBFBBBF8D0" ID-GBIF-Occurrence="1572325966" country="Iceland" location="South Australia" pageId="23" pageNumber="347" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB3523D8BBFEBEF8D1" country="Australia" name="South Australia" pageId="23" pageNumber="347">South Australia</collectingRegion>
: Alton Down, Birdsville-
<quantity id="C818D0F5FFF3FFDB3252D954FDC8F8D1" box="[554,628,1818,1843]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="23" pageNumber="347" unit="km" value="48.0">48 km</quantity>
SW, 1972.xx.xx, [-26.28, 139.10]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB346ED954FDD4F8B4" ID-GBIF-Occurrence="1572325964" country="Iceland" location="South Australia" pageId="23" pageNumber="347" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB346ED954FB71F8D1" box="[1046,1229,1818,1843]" country="Australia" name="South Australia" pageId="23" pageNumber="347">South Australia</collectingRegion>
: Flinders Ranges, Elatina Hut
<quantity id="C818D0F5FFF3FFDB315ED970FEDEF8B4" box="[294,354,1853,1878]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="347" unit="km" value="1.0">1 km</quantity>
NW, [-31.35, 138.63]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB320FD973FF4DF899" ID-GBIF-Occurrence="1572325967" country="Iceland" location="South Australia" pageId="23" pageNumber="347" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB320FD973FC92F8B4" box="[631,814,1853,1878]" country="Australia" name="South Australia" pageId="23" pageNumber="347">South Australia</collectingRegion>
: Flinders Ranges, Westwloona-
<quantity id="C818D0F5FFF3FFDB34E6D973FB59F8B4" box="[1182,1253,1853,1878]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="347" unit="km" value="14.0">14 km</quantity>
WSW, [-31.50, 138.50]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB3084D92CFB7CF899" ID-GBIF-Occurrence="1572325975" box="[252,1216,1890,1915]" country="Iceland" location="Flinders Range" pageId="23" pageNumber="347" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB3084D92CFE13F899" box="[252,431,1890,1915]" country="Australia" name="South Australia" pageId="23" pageNumber="347">South Australia</collectingRegion>
:
<location id="0A3F2BCBFFF3FFDB31C2D92CFDDBF899" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFF3FFDB31C2D92CFDDBF899" box="[442,615,1890,1915]" country="Iceland" name="Flinders Range" pageId="23" pageNumber="347" stateProvince="South Australia">Flinders Range</location>
, 1999.01.0 6, (GenBank
<accessionNumber id="10B3E0F3FFF3FFDB3301D92CFC4BF899" box="[889,1015,1890,1915]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/DQ023068" pageId="23" pageNumber="347">
DQ
<date id="7B5E5BD0FFF3FFDB33DAD92CFC4BF899" box="[930,1015,1890,1915]" pageId="23" pageNumber="347" value="68-02-30">023068</date>
</accessionNumber>
) [-31.37, 138.63]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB34B3D92DFA25F899" ID-GBIF-Occurrence="1572325982" box="[1227,1433,1890,1915]" country="Iceland" location="Western Australia" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB34B3D92DFA25F899" box="[1227,1433,1890,1915]" country="Australia" name="Western Australia" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB34B3D92DFA9AF898" box="[1227,1318,1891,1914]" country="American Samoa" name="Western" pageId="23" pageNumber="347">Western</collectingRegion>
Australia
</collectingRegion>
</materialsCitation>
:
<materialsCitation id="BF88774DFFF3FFDB30EFD9CBFDD1F87C" ID-GBIF-Occurrence="1572325983" box="[151,621,1925,1950]" country="Iceland" location="Derby" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Derby" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB30EFD9CBFF61F87C" box="[151,221,1925,1950]" country="United Kingdom" name="Derby" pageId="23" pageNumber="347">Derby</collectingRegion>
City, 1982.xx.xx, [-17.31, 123.62]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB3201D9C9FA24F87C" ID-GBIF-Occurrence="1572325973" box="[633,1432,1925,1950]" country="Iceland" location="Eurardy station" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB3201D9C9FCF4F87C" box="[633,840,1925,1950]" country="Australia" name="Western Australia" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB3201D9C9FD68F87C" box="[633,724,1927,1950]" country="American Samoa" name="Western" pageId="23" pageNumber="347">Western</collectingRegion>
Australia
</collectingRegion>
:
<location id="0A3F2BCBFFF3FFDB332CD9CBFBB8F87C" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFF3FFDB332CD9CBFBB8F87C" box="[852,1028,1925,1950]" country="Iceland" name="Eurardy station" pageId="23" pageNumber="347" stateProvince="Western Australia">Eurardy station</location>
, 2015.02.04/11, [-27.531, 114.667]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB30EFD9E5FC22F821" ID-GBIF-Occurrence="1572325968" box="[151,926,1962,1987]" country="Iceland" location="Western Australia" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB30EFD9E5FEDBF821" box="[151,359,1962,1987]" country="Australia" name="Western Australia" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB30EFD9E5FF4EF820" box="[151,242,1963,1986]" country="American Samoa" name="Western" pageId="23" pageNumber="347">Western</collectingRegion>
Australia
</collectingRegion>
: Perth: Kings Park, 1969.12.14, [-31.96, 115.87]
</materialsCitation>
;
<materialsCitation id="BF88774DFFF3FFDB33D4D9E5FE0BF804" ID-GBIF-Occurrence="1572325974" collectorName="J. Clark" country="Iceland" location="Perth" pageId="23" pageNumber="347" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<collectingRegion id="CD24B3F2FFF3FFDB33D4D9E5FBC0F821" box="[940,1148,1962,1987]" country="Australia" name="Western Australia" pageId="23" pageNumber="347">
<collectingRegion id="CD24B3F2FFF3FFDB33D4D9E5FBBBF820" box="[940,1031,1963,1986]" country="American Samoa" name="Western" pageId="23" pageNumber="347">Western</collectingRegion>
Australia
</collectingRegion>
:
<location id="0A3F2BCBFFF3FFDB34F1D9E4FB7BF821" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFF3FFDB34F1D9E4FB7BF821" box="[1161,1223,1962,1987]" country="Iceland" name="Perth" pageId="23" pageNumber="347" stateProvince="Western Australia">Perth</location>
, pre 1965 (coll.
<collectorName id="A21518C6FFF3FFDB35F2D9E5FF66F804" pageId="23" pageNumber="347">J. Clark</collectorName>
), [-31.97, 115.840].
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="5B9F2D98FFFCFFD430EFDAF3FE7CFB10" httpUri="https://zenodo.org/record/829101/files/figure.png" pageId="24" pageNumber="348" targetBox="[340,1247,195,1191]" targetPageId="24">
<paragraph id="0F5F7D10FFFCFFD430EFDAF3FE7CFB10" blockId="24.[151,1436,1213,1266]" pageId="24" pageNumber="348">
<emphasis id="3D94A102FFFCFFD430EFDAF3FE98FB30" bold="true" box="[151,292,1213,1235]" pageId="24" pageNumber="348">FIGURE 15.</emphasis>
Head of
<taxonomicName id="C8E00693FFFCFFD431F6DAF3FD2EFB31" box="[398,658,1213,1235]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="348" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFCFFD431F6DAF3FD2EFB31" box="[398,658,1213,1235]" italics="true" pageId="24" pageNumber="348">Cardiocondyla paranuda</emphasis>
</taxonomicName>
in dorsal aspect. Australia: Kunoth Paddock [23.517°S, 133.210°E], leg. Greenslade 24 October 1974.
</paragraph>
</caption>
<subSubSection id="47FA2E9BFFFCFFD530BFDB57FAC5FAA7" lastPageId="25" lastPageNumber="349" pageId="24" pageNumber="348" type="description">
<paragraph id="0F5F7D10FFFCFFD430BFDB57FB3CF85B" blockId="24.[151,1437,1305,1978]" pageId="24" pageNumber="348">
<emphasis id="3D94A102FFFCFFD430BFDB57FD82FAD0" bold="true" box="[199,574,1305,1330]" pageId="24" pageNumber="348">Redescription of worker caste.</emphasis>
Worker (
<tableCitation id="426248ABFFFCFFD432D6DB57FD47FAD3" box="[686,763,1305,1330]" captionStart="TABLE 2" captionStartId="22.[151,239,152,175]" captionText="TABLE 2. Morphometric data of workers of C. nuda species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals. Data with reduced number of measured individuals in C. nuda are given in italics." pageId="24" pageNumber="348">Tab. 2</tableCitation>
,
<figureCitation id="97DB6195FFFCFFD43373DB57FC29FAD0" box="[779,917,1305,1330]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="23.[151,250,1258,1280]" captionStartId-1="24.[151,250,1213,1235]" captionStartId-2="25.[151,250,1226,1248]" captionTargetBox-0="[276,1303,179,1233]" captionTargetBox-1="[340,1247,195,1191]" captionTargetBox-2="[151,1436,195,1205]" captionTargetId-0="figure@23.[276,1303,179,1233]" captionTargetId-1="figure@24.[340,1247,195,1191]" captionTargetId-2="figure@25.[151,1436,195,1205]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURE 14. Gaster of Cardiocondyla paranuda in dorsal aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." captionText-1="FIGURE 15. Head of Cardiocondyla paranuda in dorsal aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." captionText-2="FIGURE 16. Cardiocondyla paranuda in lateral aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." httpUri-0="https://zenodo.org/record/829099/files/figure.png" httpUri-1="https://zenodo.org/record/829101/files/figure.png" httpUri-2="https://zenodo.org/record/829103/files/figure.png" pageId="24" pageNumber="348">
Figs. 14
<emphasis id="3D94A102FFFCFFD43311DB55FCCAFAD2" bold="true" box="[873,886,1307,1328]" pageId="24" pageNumber="348"></emphasis>
16
</figureCitation>
): Head elongated, CL/CW 1.214. Postocular distance rather large, PoOc/CL 0.463. Eyes relatively small, EYE 0.234. Frontal carinae immediately caudal of the FRS level parallel or very slightly converging. Foveolae on vertex without interspaces, deeply impressed, with 13
<emphasis id="3D94A102FFFCFFD435F7DB2DFA20FA9A" bold="true" box="[1423,1436,1379,1400]" pageId="24" pageNumber="348"></emphasis>
19 µm diameter, and with an inner corona (a flat tubercle) of 7
<emphasis id="3D94A102FFFCFFD43317DBC9FCC0FA7E" bold="true" box="[879,892,1415,1436]" pageId="24" pageNumber="348"></emphasis>
9 µm diameter having the base of a decumbent pubescence hair in its center. This
<typeStatus id="D05BC3B2FFFCFFD43263DBE4FDF7FA20" box="[539,587,1450,1474]" pageId="24" pageNumber="348">type</typeStatus>
of sculpture can also be described as a strongly sculptured microreticulum. Longitudinal sculpture on vertex often completely absent (
<figureCitation id="97DB6195FFFCFFD43353DB83FCC3FA04" box="[811,895,1485,1510]" captionStart="FIGURE 15" captionStartId="24.[151,250,1213,1235]" captionTargetBox="[340,1247,195,1191]" captionTargetId="figure@24.[340,1247,195,1191]" captionTargetPageId="24" captionText="FIGURE 15. Head of Cardiocondyla paranuda in dorsal aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." httpUri="https://zenodo.org/record/829101/files/figure.png" pageId="24" pageNumber="348">Fig. 15</figureCitation>
). Weak semicircular rugae are found around the antennal fossae. Lateral mesosoma on whole surface regularly and strongly microreticulate-foveolate; longitudinal sculpture except for 4
<emphasis id="3D94A102FFFCFFD431F7D859FE20F9CE" bold="true" box="[399,412,1559,1580]" pageId="24" pageNumber="348"></emphasis>
6 weak and short carinulae on metapleuron completely absent (
<figureCitation id="97DB6195FFFCFFD43415D85BFB7EF9CC" box="[1133,1218,1557,1582]" captionStart="FIGURE 16" captionStartId="25.[151,250,1226,1248]" captionTargetBox="[151,1436,195,1205]" captionTargetId="figure@25.[151,1436,195,1205]" captionTargetPageId="25" captionText="FIGURE 16. Cardiocondyla paranuda in lateral aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." httpUri="https://zenodo.org/record/829103/files/figure.png" pageId="24" pageNumber="348">Fig. 16</figureCitation>
); dorsal mesosoma irregularly reticulate-foveolate-shagrinate. Sides of petiole with a deeply sculptured microreticulum, dorsal petiole and postpetiole with a weak and shallowly sculptured microreticulum. Cuticular surface of first gaster tergite rather smooth and shining but on its whole surface with a well-developed microreticulum (
<figureCitation id="97DB6195FFFCFFD4343CD8CFFB24F97B" box="[1092,1176,1665,1690]" captionStart="FIGURE 14" captionStartId="23.[151,250,1258,1280]" captionTargetBox="[276,1303,179,1233]" captionTargetId="figure@23.[276,1303,179,1233]" captionTargetPageId="23" captionText="FIGURE 14. Gaster of Cardiocondyla paranuda in dorsal aspect. Australia: Kunoth Paddock [23.517 ° S, 133.210 ° E], leg. Greenslade 24 October 1974." httpUri="https://zenodo.org/record/829099/files/figure.png" pageId="24" pageNumber="348">Fig. 14</figureCitation>
). The pubescence hairs on gaster tergites are the shortest within the
<taxonomicName id="C8E00693FFFCFFD432E4D8E8FD4BF95C" box="[668,759,1701,1726]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="348" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFFCFFD432E4D8E8FD4BF95C" box="[668,759,1701,1726]" italics="true" pageId="24" pageNumber="348">C. nuda</emphasis>
</taxonomicName>
group, PLG/CS is only 5.06%. Metanotal depression very shallow, MGr/CS 1.28%. Propodeal spines short but clearly longer than in the
<taxonomicName id="C8E00693FFFCFFD4345CD884FB6DF903" box="[1060,1233,1738,1761]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="348" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="3D94A102FFFCFFD4345CD884FB6DF903" box="[1060,1233,1738,1761]" italics="true" pageId="24" pageNumber="348">C. mauritanica</emphasis>
</taxonomicName>
species complex. Dorsal propodeum sloping down to base of spines under an angle of 20°. Petiole node slightly longer than wide. Postpetiole in dorsal view with only suggestedly angulate sides and straight anterior margin that is slightly shorter than posterior margin; postpetiolar sternite bulging, without any protrusions but on each side with a suggested paramedian, longitudinal carina. Head, mesosoma, waist and appendages often amber-colored, gaster significantly darker—this is the most frequently observed coloration but populations with dark headed specimens or such with concolorous amber specimens do occur. For morphometric data of
<specimenCount id="19E6B699FFFCFFD433FDD9EFFBBEF858" box="[901,1026,1953,1978]" pageId="24" pageNumber="348" type="worker">52 workers</specimenCount>
see
<tableCitation id="426248ABFFFCFFD4344BD9EFFBC0F85B" box="[1075,1148,1953,1978]" captionStart="TABLE 2" captionStartId="22.[151,239,152,175]" captionText="TABLE 2. Morphometric data of workers of C. nuda species complex in the sequence arithmetic mean ± standard deviation [minimum, maximum], n = number of measured individuals. Data with reduced number of measured individuals in C. nuda are given in italics." pageId="24" pageNumber="348">Tab. 2</tableCitation>
.
</paragraph>
<caption id="5B9F2D98FFFDFFD530EFDA84FE94FB1D" httpUri="https://zenodo.org/record/829103/files/figure.png" pageId="25" pageNumber="349" targetBox="[151,1436,195,1205]" targetPageId="25">
<paragraph id="0F5F7D10FFFDFFD530EFDA84FE94FB1D" blockId="25.[151,1435,1226,1279]" pageId="25" pageNumber="349">
<emphasis id="3D94A102FFFDFFD530EFDA84FEA3FB3D" bold="true" box="[151,287,1226,1248]" pageId="25" pageNumber="349">FIGURE 16.</emphasis>
<taxonomicName id="C8E00693FFFDFFD5315CDA84FD98FB02" box="[292,548,1226,1248]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFDFFD5315CDA84FD98FB02" box="[292,548,1226,1248]" italics="true" pageId="25" pageNumber="349">Cardiocondyla paranuda</emphasis>
</taxonomicName>
in lateral aspect. Australia: Kunoth Paddock [23.517°S, 133.210°E], leg. Greenslade 24 October 1974.
</paragraph>
</caption>
<paragraph id="0F5F7D10FFFDFFD530BFDB62FAC5FAA7" blockId="25.[151,1437,1324,2035]" box="[199,1401,1324,1349]" pageId="25" pageNumber="349">
<emphasis id="3D94A102FFFDFFD530BFDB62FE19FAA7" bold="true" box="[199,421,1324,1349]" pageId="25" pageNumber="349">Geographic range.</emphasis>
Australia, only species of the whole genus
<taxonomicName id="C8E00693FFFDFFD533F5DB62FB88FAA7" box="[909,1076,1324,1349]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="3D94A102FFFDFFD533F5DB62FB88FAA7" box="[909,1076,1324,1349]" italics="true" pageId="25" pageNumber="349">Cardiocondyla</emphasis>
</taxonomicName>
occurring in inner Australia.
</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFFDFFD530BFDB01FF40FA6F" pageId="25" pageNumber="349" type="diagnosis">
<paragraph id="0F5F7D10FFFDFFD530BFDB01FF40FA6F" blockId="25.[151,1437,1324,2035]" pageId="25" pageNumber="349">
<emphasis id="3D94A102FFFDFFD530BFDB01FEFDFA8A" bold="true" box="[199,321,1359,1384]" pageId="25" pageNumber="349">Diagnosis.</emphasis>
see key. The very short gastral pubescence is the most obvious difference to the sister species
<emphasis id="3D94A102FFFDFFD535FADB1EFF40FA6F" italics="true" pageId="25" pageNumber="349">
<taxonomicName id="C8E00693FFFDFFD535FADB1EFF44FA6F" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="atalanta">C. atalanta</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFFDFFD530BFDBD9FAB1F969" pageId="25" pageNumber="349" type="biology_ecology">
<paragraph id="0F5F7D10FFFDFFD530BFDBD9FAB1F969" blockId="25.[151,1437,1324,2035]" pageId="25" pageNumber="349">
<emphasis id="3D94A102FFFDFFD530BFDBD9FE9BFA52" bold="true" box="[199,295,1431,1456]" pageId="25" pageNumber="349">Biology.</emphasis>
<taxonomicName id="C8E00693FFFDFFD53155DBD6FE06FA52" box="[301,442,1431,1456]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFDFFD53155DBD6FE06FA52" box="[301,442,1431,1456]" italics="true" pageId="25" pageNumber="349">C. paranuda</emphasis>
</taxonomicName>
is apparently well adapted to arid and very hot climate and the only species of the whole genus
<taxonomicName id="C8E00693FFFDFFD5309CDBF2FE37FA37" box="[228,395,1468,1493]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="3D94A102FFFDFFD5309CDBF2FE37FA37" box="[228,395,1468,1493]" italics="true" pageId="25" pageNumber="349">Cardiocondyla</emphasis>
</taxonomicName>
occurring in inner
<collectingCountry id="77F73D80FFFDFFD5320ADBF2FD60FA37" box="[626,732,1468,1493]" name="Australia" pageId="25" pageNumber="349">Australia</collectingCountry>
. This is demonstrated by significant differences between
<taxonomicName id="C8E00693FFFDFFD535FADBF3FF48FA1A" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="atalanta">
<emphasis id="3D94A102FFFDFFD535FADBF3FF48FA1A" italics="true" pageId="25" pageNumber="349">C. atalanta</emphasis>
</taxonomicName>
and
<taxonomicName id="C8E00693FFFDFFD53149DB91FE0BFA1A" box="[305,439,1503,1528]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFDFFD53149DB91FE0BFA1A" box="[305,439,1503,1528]" italics="true" pageId="25" pageNumber="349">C.paranuda</emphasis>
</taxonomicName>
in the continentality of the sites. The mean distance from sea shore and mean annual rainfall are 23 ± 51 [0,252] km and 1430 ± 716 [500, 4500] mm in 27 sites of
<taxonomicName id="C8E00693FFFDFFD53468D84BFB33F9FF" box="[1040,1167,1540,1565]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="atalanta">
<emphasis id="3D94A102FFFDFFD53468D84BFB33F9FF" box="[1040,1167,1540,1565]" italics="true" pageId="25" pageNumber="349">C. atalanta</emphasis>
</taxonomicName>
and 329 ± 332 [0, 904] km and 588 ± 385 [150, 1250] mm in 27 sites of
<taxonomicName id="C8E00693FFFDFFD532B8D866FCF2F9A2" box="[704,846,1575,1600]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFDFFD532B8D866FCF2F9A2" box="[704,846,1575,1600]" italics="true" pageId="25" pageNumber="349">C. paranuda</emphasis>
</taxonomicName>
. These differences are significantly different in both sea shore distance (ANOVA F1,52=22.39, p&lt;0.0005) and annual rainfall (ANOVA F1,52=28.90, p&lt;0.0005). As yet only foragers have been collected and colony structure, male morphology, and behavior are unknown.
</paragraph>
</subSubSection>
<subSubSection id="47FA2E9BFFFDFFD630BFD8D8FBB3FE6A" lastPageId="26" lastPageNumber="350" pageId="25" pageNumber="349" type="discussion">
<paragraph id="0F5F7D10FFFDFFD630BFD8D8FBB3FE6A" blockId="25.[151,1437,1324,2035]" lastBlockId="26.[151,1437,151,392]" lastPageId="26" lastPageNumber="350" pageId="25" pageNumber="349">
<materialsCitation id="BF88774DFFFDFFD630BFD8D8FBB6FE6A" ID-GBIF-Occurrence="1572325977" collectingDate="1982" collectingDateMax="1990" collectingDateMin="1982" collectorName="Comments. There &amp; The &amp; C. A. Collingwood &amp; H. Heatwole" country="Tunisia" county="If" lastPageId="26" lastPageNumber="350" location="West Palaearctic" municipality="This" pageId="25" pageNumber="349" specimenCount="4" stateProvince="Queensland" typeStatus="holotype">
<collectorName id="A21518C6FFFDFFD530BFD8D8FE27F94D" box="[199,411,1686,1711]" pageId="25" pageNumber="349">
<emphasis id="3D94A102FFFDFFD530BFD8D8FEF3F94D" bold="true" box="[199,335,1686,1711]" pageId="25" pageNumber="349">Comments.</emphasis>
There
</collectorName>
is a serious problem with the site documentation in the
<typeStatus id="D05BC3B2FFFDFFD53445D8D8FB23F94D" box="[1085,1183,1686,1711]" pageId="25" pageNumber="349" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C8E00693FFFDFFD534B4D8D9FAE2F94D" box="[1228,1374,1686,1711]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFDFFD534B4D8D9FAE2F94D" box="[1228,1374,1686,1711]" italics="true" pageId="25" pageNumber="349">C. paranuda</emphasis>
</taxonomicName>
.
<collectorName id="A21518C6FFFDFFD53508D8D8FA20F94D" box="[1392,1436,1686,1711]" pageId="25" pageNumber="349">The</collectorName>
specimen was sent by
<collectorName id="A21518C6FFFDFFD531EAD8F5FDDCF931" box="[402,608,1722,1747]" pageId="25" pageNumber="349">C.A. Collingwood</collectorName>
to the senior author in the 1980s with the labelling &quot;
<emphasis id="3D94A102FFFDFFD534C8D8F5FCABF915" italics="true" pageId="25" pageNumber="349">
<collectingCountry id="77F73D80FFFDFFD534C8D8F5FAA7F930" box="[1200,1307,1723,1746]" name="Tunisia" pageId="25" pageNumber="349">TUNISIA</collectingCountry>
: Medinine-
<quantity id="C818D0F5FFFDFFD530EFD891FF5DF915" box="[151,225,1758,1783]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="25" pageNumber="349" unit="km" value="32.0">32 km</quantity>
SE Chabania-
<quantity id="C818D0F5FFFDFFD531EAD890FE73F915" box="[402,463,1758,1783]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="349" unit="km" value="6.0">6 km</quantity>
NW leg.
<collectorName id="A21518C6FFFDFFD53233D890FD6EF915" box="[587,722,1758,1783]" pageId="25" pageNumber="349">H.Heatwole</collectorName>
1976
</emphasis>
“.
<collectingCounty id="E63E059CFFFDFFD53340D890FCF7F915" box="[824,843,1758,1783]" pageId="25" pageNumber="349">If</collectingCounty>
run as a wild-card in a LDA considering all 16 morphometric characters and collecting all samples of the
<taxonomicName id="C8E00693FFFDFFD5335FD94DFC6DF8F9" box="[807,977,1795,1819]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="3D94A102FFFDFFD5335FD94DFC6DF8F9" box="[807,977,1795,1819]" italics="true" pageId="25" pageNumber="349">C. mauritanica</emphasis>
</taxonomicName>
species complex in class 1 and all of the
<taxonomicName id="C8E00693FFFDFFD530EFD969FF4CF8DD" box="[151,240,1830,1855]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFFDFFD530EFD969FF4CF8DD" box="[151,240,1830,1855]" italics="true" pageId="25" pageNumber="349">C. nuda</emphasis>
</taxonomicName>
complex in class 2, the
<typeStatus id="D05BC3B2FFFDFFD5327BD968FDD9F8DD" box="[515,613,1830,1855]" pageId="25" pageNumber="349" type="holotype">holotype</typeStatus>
<taxonomicName id="C8E00693FFFDFFD53215D969FD40F8DD" box="[621,764,1830,1855]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFDFFD53215D969FD40F8DD" box="[621,764,1830,1855]" italics="true" pageId="25" pageNumber="349">C. paranuda</emphasis>
</taxonomicName>
is allocated to the
<taxonomicName id="C8E00693FFFDFFD533ABD969FB90F8DD" box="[979,1068,1830,1855]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFFDFFD533ABD969FB90F8DD" box="[979,1068,1830,1855]" italics="true" pageId="25" pageNumber="349">C. nuda</emphasis>
</taxonomicName>
complex with p=1.0000.
<collectingMunicipality id="EF3BE76AFFFDFFD53528D968FA3EF8DD" box="[1360,1410,1830,1855]" pageId="25" pageNumber="349">This</collectingMunicipality>
is problematic because species of the
<taxonomicName id="C8E00693FFFDFFD5325FD905FD3DF881" box="[551,641,1866,1891]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFFDFFD5325FD905FD3DF881" box="[551,641,1866,1891]" italics="true" pageId="25" pageNumber="349">C. nuda</emphasis>
</taxonomicName>
species complex are completely absent from the
<location id="0A3F2BCBFFFDFFD534C9D905FAD0F881" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD534C9D905FAD0F881" box="[1201,1388,1866,1891]" country="Tunisia" county="If" municipality="This" name="West Palaearctic" pageId="25" pageNumber="349" stateProvince="Queensland">West Palaearctic</location>
and
<location id="0A3F2BCBFFFDFFD530EFD920FE90F865" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD530EFD920FE90F865" box="[151,300,1902,1927]" country="Tunisia" county="If" municipality="This" name="North Africa" pageId="25" pageNumber="349" stateProvince="Queensland">North Africa</location>
and it appears also most unlikely that ants from
<location id="0A3F2BCBFFFDFFD533FDD920FC57F865" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD533FDD920FC57F865" box="[901,1003,1902,1927]" country="Tunisia" county="If" municipality="This" name="Australia" pageId="25" pageNumber="349" stateProvince="Queensland">Australia</location>
should have been anthropogenically introduced to a site in the
<location id="0A3F2BCBFFFDFFD531BED9DCFDAEF849" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD531BED9DCFDAEF849" box="[454,530,1938,1963]" country="Tunisia" county="If" municipality="This" name="Sahara" pageId="25" pageNumber="349" stateProvince="Queensland">Sahara</location>
desert.
<location id="0A3F2BCBFFFDFFD53217D9DCFD42F849" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD53217D9DCFD42F849" box="[623,766,1938,1963]" country="Tunisia" county="If" municipality="This" name="Furthermore" pageId="25" pageNumber="349" stateProvince="Queensland">Furthermore</location>
, NC-clustering places the
<typeStatus id="D05BC3B2FFFDFFD53455D9DCFB33F849" box="[1069,1167,1938,1963]" pageId="25" pageNumber="349" type="holotype">holotype</typeStatus>
in a cluster of
<taxonomicName id="C8E00693FFFDFFD53539D9DDFA20F849" box="[1345,1436,1938,1963]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="3D94A102FFFDFFD53539D9DDFA20F849" box="[1345,1436,1938,1963]" italics="true" pageId="25" pageNumber="349">C. nuda</emphasis>
</taxonomicName>
group specimens that are treated as a single species that is restricted to the
<location id="0A3F2BCBFFFDFFD53385D9F8FBCEF82D" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD53385D9F8FBCEF82D" box="[1021,1138,1974,1999]" country="Tunisia" county="If" municipality="This" name="Australian" pageId="25" pageNumber="349" stateProvince="Queensland">Australian</location>
continent and sister to
<taxonomicName id="C8E00693FFFDFFD535FAD9F9FF48F811" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="349" phylum="Arthropoda" rank="species" species="atalanta">
<emphasis id="3D94A102FFFDFFD535FAD9F9FF48F811" italics="true" pageId="25" pageNumber="349">C. atalanta</emphasis>
</taxonomicName>
(
<figureCitation id="97DB6195FFFDFFD5317ED994FEF1F811" box="[262,333,2010,2035]" captionStart="FIGURE 8" captionStartId="11.[151,250,1829,1851]" captionTargetBox="[219,1372,198,1807]" captionTargetId="figure@11.[214,1372,193,1807]" captionTargetPageId="11" captionText="FIGURE 8. NC-Ward clustering of 61 samples of the two remaining Cardiocondyla nuda species complex after C. nuda has been removed from analysis. The lower of the two main branches belongs to C. atalanta Forel, the upper one to C. paranuda. The left column of bars indicates the final clustering hypothesis for K = 2 (C. atalanta vs. C. paranuda) determined by a controlling linear discriminant function (LDA), and the right column of bars the classification by non-hierarchical NC-K- Means clustering. The classification errors in in NC-Ward and NC-K-Means are both 3.3 %." httpUri="https://zenodo.org/record/829089/files/figure.png" pageId="25" pageNumber="349">Fig. 8</figureCitation>
). A wild-card run in a LDA confirms this allocation with p=0.9916 (see section 4.4).
<location id="0A3F2BCBFFFDFFD53549D994FAE1F811" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFDFFD53549D994FAE1F811" box="[1329,1373,2010,2035]" country="Tunisia" county="If" municipality="This" name="The" pageId="25" pageNumber="349" stateProvince="Queensland">The</location>
most probable explanation for this conflicting situation is a confusion of labels.
<location id="0A3F2BCBFFFEFFD63385DED9FB7DFF52" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD63385DED9FB7DFF52" box="[1021,1217,151,176]" country="Tunisia" county="If" municipality="This" name="Harold Heatwole" pageId="26" pageNumber="350" stateProvince="Queensland">Harold Heatwole</location>
collected in
<location id="0A3F2BCBFFFEFFD63523DED9FF5EFF37" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD63523DED9FF5EFF37" country="Tunisia" county="If" municipality="This" name="North Africa" pageId="26" pageNumber="350" stateProvince="Queensland">North Africa</location>
,
<location id="0A3F2BCBFFFEFFD63095DEF2FE9BFF37" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD63095DEF2FE9BFF37" box="[237,295,188,213]" country="Tunisia" county="If" municipality="This" name="Tibet" pageId="26" pageNumber="350" stateProvince="Queensland">Tibet</location>
and Australia—for instance, the two
<taxonomicName id="C8E00693FFFEFFD632B6DEF3FCE0FF37" box="[718,860,188,213]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="350" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFEFFD632B6DEF3FCE0FF37" box="[718,860,188,213]" italics="true" pageId="26" pageNumber="350">C. paranuda</emphasis>
</taxonomicName>
samples from
<collectingRegion id="CD24B3F2FFFEFFD6347CDEF2FB31FF37" box="[1028,1165,188,213]" country="Australia" name="Queensland" pageId="26" pageNumber="350">Queensland</collectingRegion>
:
<location id="0A3F2BCBFFFEFFD634E0DEF2FA82FF37" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD634E0DEF2FA82FF37" box="[1176,1342,188,213]" country="Tunisia" county="If" municipality="This" name="Chilcott Island" pageId="26" pageNumber="350" stateProvince="Queensland">Chilcott Island</location>
in 1967 were taken by him.
<location id="0A3F2BCBFFFEFFD631F9DEAEFE1EFF15" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD631F9DEAEFE1EFF15" box="[385,418,224,247]" country="Tunisia" county="If" municipality="This" name="He" pageId="26" pageNumber="350" stateProvince="Queensland">He</location>
usually gave his specimens to
<location id="0A3F2BCBFFFEFFD63377DE91FC1DFF1A" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD63377DE91FC1DFF1A" box="[783,929,223,248]" country="Tunisia" county="If" municipality="This" name="Collingwood" pageId="26" pageNumber="350" stateProvince="Queensland">Collingwood</location>
stored in tubes with ethanol.
<location id="0A3F2BCBFFFEFFD63486DEAEFAA1FF1A" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD63486DEAEFAA1FF1A" box="[1278,1309,224,248]" country="Tunisia" county="If" municipality="This" name="As" pageId="26" pageNumber="350" stateProvince="Queensland">As</location>
repeatedly witnessed by the senior author in personal contacts during laboratory work in
<collectingDate id="6B1AA238FFFEFFD63386DF4AFB1BFEFE" box="[1022,1191,260,285]" pageId="26" pageNumber="350" value="1982" valueMax="1990">
<date id="7B5E5BD0FFFEFFD63386DF4AFB1BFEFE" box="[1022,1191,260,285]" pageId="26" pageNumber="350" value="1982" valueMax="1990">1982 and 1990</date>
</collectingDate>
,
<location id="0A3F2BCBFFFEFFD634CADF4AFAF8FEFF" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD634CADF4AFAF8FEFF" box="[1202,1348,260,285]" country="Tunisia" county="If" municipality="This" name="Collingwood" pageId="26" pageNumber="350" stateProvince="Queensland">Collingwood</location>
had the dangerous habit of placing similar ethanol-stored ants from different tubes side-by-side under the microscope for better comparison and sometimes he confused from which tube he had taken the specimens.
<location id="0A3F2BCBFFFEFFD634D7DF03FB6FFE86" LSID="urn:lsid:plazi:treatment:8749CC06FFF2FFD630EFD84AFBB3FE6A:0A3F2BCBFFFEFFD634D7DF03FB6FFE86" box="[1199,1235,333,356]" country="Tunisia" county="If" municipality="This" name="We" pageId="26" pageNumber="350" stateProvince="Queensland">We</location>
conclude that the
<typeStatus id="D05BC3B2FFFEFFD630EFDF3EFF7BFE6A" box="[151,199,368,392]" pageId="26" pageNumber="350">type</typeStatus>
of
<taxonomicName id="C8E00693FFFEFFD63094DF3EFEC6FE6A" box="[236,378,367,392]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="350" phylum="Arthropoda" rank="species" species="paranuda">
<emphasis id="3D94A102FFFEFFD63094DF3EFEC6FE6A" box="[236,378,367,392]" italics="true" pageId="26" pageNumber="350">C. paranuda</emphasis>
</taxonomicName>
has most probably been collected somewhere in
<collectingCountry id="77F73D80FFFEFFD633D8DF21FBB6FE6A" box="[928,1034,367,392]" name="Australia" pageId="26" pageNumber="350">Australia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>