263 lines
32 KiB
XML
263 lines
32 KiB
XML
<document id="5269A33BB9966D63CACA759B8482E526" ID-DOI="10.1111/j.1096-3642.2011.00717.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5441686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630451578892" checkinUser="valdenar" docAuthor="Soto, Matías, Pol, Diego & Perea, Daniel" docDate="2011" docId="5D268794FF806A1EC77BFBF5584CFF0B" docLanguage="en" docName="j.1096-3642.2011.00717.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (5)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00717.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Uruguaysuchus terrai Rusconi 1933" docType="treatment" docVersion="4" lastPageNumber="645" masterDocId="A11FFFECFF936A08C426FF52596CFFF1" masterDocTitle="A new specimen of Uruguaysuchus aznarezi (Crocodyliformes: Notosuchia) from the middle Cretaceous of Uruguay and its phylogenetic relationships" masterLastPageNumber="645" masterPageNumber="644" pageNumber="644" updateTime="1699275399670" updateUser="plazi">
|
||
<mods:mods id="8E6941E9298CF112AEF1B01BA7A3C919" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0250F9C4CE35DF283D8A897F045240C0">
|
||
<mods:title id="D497DF9BE6A38134DAAAE0CED0C92DDC">A new specimen of Uruguaysuchus aznarezi (Crocodyliformes: Notosuchia) from the middle Cretaceous of Uruguay and its phylogenetic relationships</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="DDB810B38675A6273A5ECED48DBC3807" type="personal">
|
||
<mods:role id="B049A2E776A300DC3D81793E18802B3C">
|
||
<mods:roleTerm id="06A1743B93FC45E98542906FC54326F1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="72F2BFA96538D334288AA179C64B64E7">Soto, Matías</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0D287E534A9FCEE678985AF6117B75E3" type="personal">
|
||
<mods:role id="400DA5681997F6B24C7440BE298091EE">
|
||
<mods:roleTerm id="C569425A2EC0CB97DD260FB9AC5A00CF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5732504101536181D6CB403B4D44AEE1">Pol, Diego</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9A02E2860AEADD7FE4E6AAAAEC0EB97E" type="personal">
|
||
<mods:role id="4B51EA2666FF9868AAA14F6FD0801D92">
|
||
<mods:roleTerm id="CC456C33E5AEEF2AC9C3A85DAC54BA18">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B80A796D6D900521559EB717EADA8795">Perea, Daniel</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="52FE97FC70C92020D139392501593CD4">text</mods:typeOfResource>
|
||
<mods:relatedItem id="462D99BEE984D123A26F07B5E7B9986F" type="host">
|
||
<mods:titleInfo id="48E70AADCA8FCFC4B5D6AE429D182DE9">
|
||
<mods:title id="10637C4610CB49E4E99AB0A1A044E478">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1BC3F406AA956C35B7A5CE2090454CA6">
|
||
<mods:date id="B8F0AC4D3A76138981BBCCBEBD6E9178">2011</mods:date>
|
||
<mods:detail id="ED11285752A3EA15D472FCB14FAE10C9" type="pubDate">
|
||
<mods:number id="C1A35512272CAF0EF01BB1B3CAFE95F0">2012-01-12</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D56F11ECDCE60A4EE90364D32733C901" type="volume">
|
||
<mods:number id="F6E506068B5E18E5BAB70EC0649FA41B">163</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="FB9A174BC06DD2C8BC218FCF832FA7AD" type="issue">
|
||
<mods:number id="FEED3259D4418500FCA2A82A05F715A1">5</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B014534CA4E1DF0D349517EEB970B2C7" unit="page">
|
||
<mods:start id="5E17E8D1608E7D581DCFFC2F46BB6D03">644</mods:start>
|
||
<mods:end id="FE6FBC01DE886DED3B0463183AB5CC09">645</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="81618104A607E7E24F3C5E9F8BC941B4">
|
||
<mods:url id="653B15722B8A510C08A7929555A5463E">http://dx.doi.org/10.1111/j.1096-3642.2011.00717.x</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="A00E3ADEFCB8B8FCAC8657F8764FABE9">journal article</mods:classification>
|
||
<mods:identifier id="A5974D2D8B772364E3F714D5F453BCD5" type="DOI">10.1111/j.1096-3642.2011.00717.x</mods:identifier>
|
||
<mods:identifier id="85630D770904574C23DE552399E78522" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="DB671BAE4A186BC78C2F8658FD472296" type="Zenodo-Dep">5441686</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5D268794FF806A1EC77BFBF5584CFF0B" ID-DOI="http://doi.org/10.5281/zenodo.5492111" ID-Zenodo-Dep="5492111" LSID="urn:lsid:plazi:treatment:5D268794FF806A1EC77BFBF5584CFF0B" httpUri="http://treatment.plazi.org/id/5D268794FF806A1EC77BFBF5584CFF0B" lastPageId="22" pageId="19">
|
||
<subSubSection id="9D956509FF806A1BC77BFBF55C11FB4F" box="[861,1405,1191,1215]" pageId="19" type="nomenclature">
|
||
<paragraph id="D5303682FF806A1BC77BFBF55C11FB4F" blockId="19.[861,1405,1191,1215]" box="[861,1405,1191,1215]" pageId="19">
|
||
<heading id="8E7881EEFF806A1BC77BFBF55C11FB4F" allCaps="true" box="[861,1405,1191,1215]" centered="true" fontSize="10" level="1" pageId="19" reason="1">
|
||
STATUS OF
|
||
<taxonomicName id="128F4D01FF806A1BC022FBF55C11FB4F" authorityName="Rusconi" authorityYear="1933" box="[1028,1405,1191,1214]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF806A1BC022FBF55C11FB4F" box="[1028,1405,1191,1214]" italics="true" pageId="19">URUGUAYSUCHUS TERRAI</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D956509FF806A1EC71CFB84584CFF0B" lastPageId="22" pageId="19" type="description">
|
||
<paragraph id="D5303682FF806A1BC71CFB845AC8F946" blockId="19.[826,1442,1238,1903]" pageId="19">
|
||
As already stated,
|
||
<bibRefCitation id="B11E4B73FF806A1BC036FB845DD5FB1D" author="Rusconi C" box="[1040,1209,1238,1260]" pageId="19" pagination="1 - 64" refId="ref15855" refString="Rusconi C. 1933. Sobre reptiles cretaceos del Uruguay (Uruguaysuchus Aznarezi [sic] n. g. n. sp.) y sus relaciones con los notosuquidos de Patagonia. Boletin del Instituto Geologico del Uruguay 19: 1 - 64." type="journal article" year="1933">Rusconi (1933)</bibRefCitation>
|
||
recognized a second species of
|
||
<taxonomicName id="128F4D01FF806A1BC796FBA75D0FFAFB" box="[944,1123,1269,1290]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="E7FBEA90FF806A1BC796FBA75D0FFAFB" box="[944,1123,1269,1290]" italics="true" pageId="19">Uruguaysuchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="128F4D01FF806A1BC054FBA75DB9FAFB" authorityName="Rusconi" authorityYear="1933" box="[1138,1237,1269,1290]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF806A1BC054FBA75DB9FAFB" box="[1138,1237,1269,1290]" italics="true" pageId="19">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
, based mainly on differences in the tooth count in the upper tooth row. Both
|
||
<bibRefCitation id="B11E4B73FF806A1BC75FFA605A97FAB9" author="Soto M" box="[889,1019,1330,1352]" pageId="19" refId="ref15901" refString="Soto M. 2005. Especulaciones sobre Uruguaysuchus (Crocodyliformes, Meso-eucrocodylia). VIII Jornadas de Zoologia del Uruguay, Montevideo. Actas, 108." type="book" year="2005">Soto (2005)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="B11E4B73FF806A1BC011FA605C33FAB9" author="Andrade MB & Bertini RJ" box="[1079,1375,1330,1352]" pageId="19" pagination="21 - 22" refId="ref14265" refString="Andrade MB, Bertini RJ. 2005. Bibliographic revision of Uruguaysuchus (Mesoeucrocodylia, Crocodylomorpha): is Uruguaysuchus terrai a valid species? II Congresso Latino- Americano de Paleontologia de Vertebrados, Rio de Janeiro. Boletim de Resumos, 21 - 22." type="book chapter" year="2005">Andrade & Bertini (2005)</bibRefCitation>
|
||
independently questioned the taxonomic validity of maintaining the distinction of
|
||
<taxonomicName id="128F4D01FF806A1BC041FA225DA0FA74" authorityName="Rusconi" authorityYear="1933" box="[1127,1228,1392,1413]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF806A1BC041FA225DA0FA74" box="[1127,1228,1392,1413]" italics="true" pageId="19">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="128F4D01FF806A1BC133FA225CF6FA74" authorityName="Rusconi" authorityYear="1933" box="[1301,1434,1392,1413]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF806A1BC133FA225CF6FA74" box="[1301,1434,1392,1413]" italics="true" pageId="19">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="128F4D01FF806A1BC71CFADC5ACEFA52" authorityName="Rusconi" authorityYear="1933" box="[826,930,1422,1443]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF806A1BC71CFADC5ACEFA52" box="[826,930,1422,1443]" italics="true" pageId="19">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
apparently possesses four maxillary teeth more than
|
||
<taxonomicName id="128F4D01FF806A1BC79DFAFF5D52FA33" authorityName="Rusconi" authorityYear="1933" box="[955,1086,1453,1474]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF806A1BC79DFAFF5D52FA33" box="[955,1086,1453,1474]" italics="true" pageId="19">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
: two incisiviforms, one caniniform, and ten post-caniniforms versus one incisiviform, one caniniform and seven post-caniniforms, respectively (
|
||
<figureCitation id="4DB42A07FF806A1BC7F3F95A5D4AF9EF" box="[981,1062,1544,1566]" captionStart="Figure 16" captionStartId="21.[164,243,1169,1188]" captionTargetBox="[169,1439,201,1134]" captionTargetId="figure-279@21.[164,1445,196,1140]" captionTargetPageId="21" captionText="Figure 16. Schematic diagram (not intended to reflect the real size or shape differences among teeth) depicting maxillary teeth in U. terrai and three specimens of U. aznarezi (the holotype being probably the only adult individual) drawn at the same size. Based on descriptions and drawings provided by Rusconi (1933) in all cases except for FC-DPV 2320. Question marks indicate the possibility of an additional tooth." figureDoi="http://doi.org/10.5281/zenodo.5441718" httpUri="https://zenodo.org/record/5441718/files/figure.png" pageId="19">Fig. 16</figureCitation>
|
||
). However, as already stated, it must be noted that the lack of preparation of the
|
||
<typeStatus id="0A348820FF806A1BC71CF9175AF7F9AA" box="[826,923,1605,1627]" pageId="19" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="128F4D01FF806A1BC7E6F9145D28F9AA" authorityName="Rusconi" authorityYear="1933" box="[960,1092,1606,1627]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF806A1BC7E6F9145D28F9AA" box="[960,1092,1606,1627]" italics="true" pageId="19">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
does not allow an assessment of the total number of post-caniniform teeth (i.e. whether there are teeth beyond the ninth maxillary position).
|
||
</paragraph>
|
||
<paragraph id="D5303682FF806A1DC775F9925BE5FA74" blockId="19.[826,1442,1238,1903]" lastBlockId="21.[164,779,1361,1903]" lastPageId="21" pageId="19">
|
||
Moreover, the maxilla of Rusconi’s specimen no. 4 (
|
||
<figureCitation id="4DB42A07FF806A1BC764F98C5AFEF905" box="[834,914,1758,1780]" captionStart="Figure 16" captionStartId="21.[164,243,1169,1188]" captionTargetBox="[169,1439,201,1134]" captionTargetId="figure-279@21.[164,1445,196,1140]" captionTargetPageId="21" captionText="Figure 16. Schematic diagram (not intended to reflect the real size or shape differences among teeth) depicting maxillary teeth in U. terrai and three specimens of U. aznarezi (the holotype being probably the only adult individual) drawn at the same size. Based on descriptions and drawings provided by Rusconi (1933) in all cases except for FC-DPV 2320. Question marks indicate the possibility of an additional tooth." figureDoi="http://doi.org/10.5281/zenodo.5441718" httpUri="https://zenodo.org/record/5441718/files/figure.png" pageId="19">Fig. 16</figureCitation>
|
||
), a juvenile
|
||
<taxonomicName id="128F4D01FF806A1BC01EF98D5DAFF905" authorityName="Rusconi" authorityYear="1933" box="[1080,1219,1759,1780]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF806A1BC01EF98D5DAFF905" box="[1080,1219,1759,1780]" italics="true" pageId="19">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
according to this author, exhibits two incisiforms and one caniniform (being the third maxillary tooth), the same condition as in
|
||
<taxonomicName id="128F4D01FF806A1BC7AEF8695A9EF8A1" authorityName="Rusconi" authorityYear="1933" box="[904,1010,1851,1872]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF806A1BC7AEF8695A9EF8A1" box="[904,1010,1851,1872]" italics="true" pageId="19">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
. Curiously,
|
||
<bibRefCitation id="B11E4B73FF806A1BC0ACF8685C56F8A1" author="Rusconi C" box="[1162,1338,1850,1872]" pageId="19" pagination="1 - 64" refId="ref15855" refString="Rusconi C. 1933. Sobre reptiles cretaceos del Uruguay (Uruguaysuchus Aznarezi [sic] n. g. n. sp.) y sus relaciones con los notosuquidos de Patagonia. Boletin del Instituto Geologico del Uruguay 19: 1 - 64." type="journal article" year="1933">Rusconi (1933)</bibRefCitation>
|
||
did not explicitly mention this relevant point. Interestingly, his fig. 20 illustrates seven post-caniniform alveoli although only six are implied in the text.
|
||
</paragraph>
|
||
<caption id="81F0660AFF876A1CC4B6F85A589BF8A7" ID-DOI="http://doi.org/10.5281/zenodo.5441716" ID-Zenodo-Dep="5441716" httpUri="https://zenodo.org/record/5441716/files/figure.png" pageId="20" startId="20.[144,223,1800,1819]" targetBox="[150,1417,202,1763]" targetPageId="20">
|
||
<paragraph id="D5303682FF876A1CC4B6F85A589BF8A7" blockId="20.[144,1422,1800,1878]" pageId="20">
|
||
<emphasis id="E7FBEA90FF876A1CC4B6F85A5869F8EA" bold="true" box="[144,261,1800,1819]" pageId="20">Figure 15.</emphasis>
|
||
Strict consensus of the two MPTs found in the phylogenetic analysis, based on the dataset published by
|
||
<bibRefCitation id="B11E4B73FF876A1CC14AF85A5833F8C9" author="Pol D & Gasparini Z" pageId="20" pagination="163 - 197" refId="ref15732" refString="Pol D, Gasparini Z. 2009. Skull anatomy of Dakosaurus andiniensis (Thalattosuchia: Crocodylomorpha) and the phylogenetic position of Thalattosuchia. Journal of Systematic Palaeontology 7: 163 - 197." type="journal article" year="2009">Pol & Gasparini (2009)</bibRefCitation>
|
||
. The new information provided by FC-DPV 2320 allowed us to recover
|
||
<taxonomicName id="128F4D01FF876A1CC07BF8775C6DF8C9" box="[1117,1281,1829,1848]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="E7FBEA90FF876A1CC07BF8775C6DF8C9" box="[1117,1281,1829,1848]" italics="true" pageId="20">Uruguaysuchus</emphasis>
|
||
</taxonomicName>
|
||
as the sister taxon of the
|
||
<taxonomicName id="128F4D01FF876A1CC53CF81158DDF8A7" box="[282,433,1859,1878]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="E7FBEA90FF876A1CC53CF81158DDF8A7" box="[282,433,1859,1878]" italics="true" pageId="20">Araripesuchus</emphasis>
|
||
</taxonomicName>
|
||
clade.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="81F0660AFF866A1DC482FBC35A2FFB0D" ID-DOI="http://doi.org/10.5281/zenodo.5441718" ID-Zenodo-Dep="5441718" httpUri="https://zenodo.org/record/5441718/files/figure.png" pageId="21" startId="21.[164,243,1169,1188]" targetBox="[169,1439,201,1134]" targetPageId="21">
|
||
<paragraph id="D5303682FF866A1DC482FBC35A2FFB0D" blockId="21.[164,1442,1169,1277]" pageId="21">
|
||
<emphasis id="E7FBEA90FF866A1DC482FBC35875FB55" bold="true" box="[164,281,1169,1188]" pageId="21">Figure 16.</emphasis>
|
||
Schematic diagram (not intended to reflect the real size or shape differences among teeth) depicting maxillary teeth in
|
||
<taxonomicName id="128F4D01FF866A1DC527FBFD5831FB30" authorityName="Rusconi" authorityYear="1933" box="[257,349,1199,1217]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF866A1DC527FBFD5831FB30" box="[257,349,1199,1217]" italics="true" pageId="21">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
and three specimens of
|
||
<taxonomicName id="128F4D01FF866A1DC641FBFD5B8CFB30" authorityName="Rusconi" authorityYear="1933" box="[615,736,1199,1217]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC641FBFD5B8CFB30" box="[615,736,1199,1217]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
(the holotype being probably the only adult individual) drawn at the same size. Based on descriptions and drawings provided by
|
||
<bibRefCitation id="B11E4B73FF866A1DC742FB9E5D6EFB2E" author="Rusconi C" box="[868,1026,1228,1247]" pageId="21" pagination="1 - 64" refId="ref15855" refString="Rusconi C. 1933. Sobre reptiles cretaceos del Uruguay (Uruguaysuchus Aznarezi [sic] n. g. n. sp.) y sus relaciones con los notosuquidos de Patagonia. Boletin del Instituto Geologico del Uruguay 19: 1 - 64." type="journal article" year="1933">Rusconi (1933)</bibRefCitation>
|
||
in all cases except for FC-DPV 2320. Question marks indicate the possibility of an additional tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D5303682FF866A1DC49AFADC5B4FFA0E" blockId="21.[164,779,1361,1903]" pageId="21">
|
||
Furthermore, several measurements of
|
||
<taxonomicName id="128F4D01FF866A1DC687FADC5A67FA52" authorityName="Rusconi" authorityYear="1933" box="[673,779,1422,1443]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF866A1DC687FADC5A67FA52" box="[673,779,1422,1443]" italics="true" pageId="21">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
are consistently smaller than those of the only adult individual of
|
||
<taxonomicName id="128F4D01FF866A1DC574FA9E588CFA10" authorityName="Rusconi" authorityYear="1933" box="[338,480,1484,1505]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC574FA9E588CFA10" box="[338,480,1484,1505]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="0A348820FF866A1DC5DDFA995B0CFA10" box="[507,608,1483,1505]" pageId="21" type="holotype">holotype</typeStatus>
|
||
), approaching those of juveniles of
|
||
<taxonomicName id="128F4D01FF866A1DC5B1FAB85B70FA0E" authorityName="Rusconi" authorityYear="1933" box="[407,540,1514,1535]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC5B1FAB85B70FA0E" box="[407,540,1514,1535]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D5303682FF866A1DC49AF95A5D5CFA10" blockId="21.[164,779,1361,1903]" lastBlockId="21.[826,1441,1361,1903]" pageId="21">
|
||
As proposed by
|
||
<bibRefCitation id="B11E4B73FF866A1DC549F95A589DF9EF" author="Soto M" box="[367,497,1544,1566]" pageId="21" refId="ref15901" refString="Soto M. 2005. Especulaciones sobre Uruguaysuchus (Crocodyliformes, Meso-eucrocodylia). VIII Jornadas de Zoologia del Uruguay, Montevideo. Actas, 108." type="book" year="2005">Soto (2005)</bibRefCitation>
|
||
, minor differences in the dental formulae can be explained by intraspecific variation in tooth count, which is rather common among crocodyliforms, either fossil or extant (C. Brochu, com. pers., 2008). In particular, if it is true that
|
||
<taxonomicName id="128F4D01FF866A1DC4F9F9F0582AF946" authorityName="Rusconi" authorityYear="1933" box="[223,326,1698,1719]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF866A1DC4F9F9F0582AF946" box="[223,326,1698,1719]" italics="true" pageId="21">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="128F4D01FF866A1DC57EF9F058B3F946" authorityName="Rusconi" authorityYear="1933" box="[344,479,1698,1719]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC57EF9F058B3F946" box="[344,479,1698,1719]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
no. 4 and FC-DPV 2320 represent juvenile individuals, it may be necessary to invoke ontogenetic loss of tooth positions to explain the fact that the adult individual of
|
||
<taxonomicName id="128F4D01FF866A1DC659F9AC5A66F8E2" authorityName="Rusconi" authorityYear="1933" box="[639,778,1790,1811]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC659F9AC5A66F8E2" box="[639,778,1790,1811]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="0A348820FF866A1DC48DF84E5861F8C3" box="[171,269,1820,1842]" pageId="21" type="holotype">holotype</typeStatus>
|
||
) has only one incisiviform maxillary tooth instead of two. Such a phenomenon has already been recorded in several crocodylian species (e.g.
|
||
<taxonomicName id="128F4D01FF866A1DC71CFA035D37FA97" box="[826,1115,1361,1382]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="cataphractus">
|
||
<emphasis id="E7FBEA90FF866A1DC71CFA035D37FA97" box="[826,1115,1361,1382]" italics="true" pageId="21">Crocodylus cataphractus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="128F4D01FF866A1DC056FA035D82FA97" authorityName="Schneider" authorityYear="1801" box="[1136,1262,1361,1382]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="porosus">
|
||
<emphasis id="E7FBEA90FF866A1DC056FA035D82FA97" box="[1136,1262,1361,1382]" italics="true" pageId="21">C. porosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="128F4D01FF866A1DC122FA035CF6FA97" authorityName="Schneider" authorityYear="1801" box="[1284,1434,1361,1382]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="siamensis">
|
||
<emphasis id="E7FBEA90FF866A1DC122FA035CF6FA97" box="[1284,1434,1361,1382]" italics="true" pageId="21">C. siamensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="128F4D01FF866A1DC751FA225D03FA75" baseAuthorityName="Muller" baseAuthorityYear="1838" box="[887,1135,1391,1413]" class="Reptilia" family="Crocodylidae" genus="Tomistoma" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="schlegelii">
|
||
<emphasis id="E7FBEA90FF866A1DC751FA225D03FA75" box="[887,1135,1391,1413]" italics="true" pageId="21">Tomistoma schlegelii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="B11E4B73FF866A1DC0AEFA3D5C74FA75" author="Mook CC" box="[1160,1304,1391,1413]" pageId="21" pagination="51 - 66" refId="ref15506" refString="Mook CC. 1921. Individual and age variation in the skulls of recent Crocodilia. Bulletin of the American Museum of Natural History 64: 51 - 66." type="journal article" year="1921">Mook, 1921</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B11E4B73FF866A1DC109FA3D5A1FFA55" author="Wermuth H" pageId="21" pagination="375 - 514" refId="ref16039" refString="Wermuth H. 1953. Systematik der rezenter krokodile. Mitteilungen der Zoologische Museum Berlin 29: 375 - 514." type="journal article" year="1953">Wermuth, 1953</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B11E4B73FF866A1DC7A7FADC5D55FA55" author="Iordansky NN" box="[897,1081,1422,1444]" pageId="21" pagination="201 - 260" refId="ref15179" refString="Iordansky NN. 1973. The skull of the Crocodylia. In: Gans C, Parsons T, eds. Biology of the Reptilia. 4. London: Academic Press, 201 - 260." type="book chapter" year="1973">Iordansky, 1973</bibRefCitation>
|
||
) as well as in the tyrannosaurid theropods
|
||
<taxonomicName id="128F4D01FF866A1DC7C2FAFF5DD2FA33" box="[996,1214,1453,1474]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="21" phylum="Chordata" rank="species" species="rex">
|
||
<emphasis id="E7FBEA90FF866A1DC7C2FAFF5DD2FA33" box="[996,1214,1453,1474]" italics="true" pageId="21">Tyrannosaurus rex</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="128F4D01FF866A1DC124FAFE5D40FA10" authority="(Carr, 1999)" baseAuthorityName="Carr" baseAuthorityYear="1999" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="21" phylum="Chordata" rank="species" species="libratus">
|
||
<emphasis id="E7FBEA90FF866A1DC124FAFE5AF8FA11" italics="true" pageId="21">Albertosaurus libratus</emphasis>
|
||
(
|
||
<bibRefCitation id="B11E4B73FF866A1DC780FA995D48FA10" author="Carr TD" box="[934,1060,1483,1505]" pageId="21" pagination="497 - 520" refId="ref14534" refString="Carr TD. 1999. Craniofacial ontogeny in Tyrannosauridae (Dinosauria, Coelurosauria). Journal of Vertebrate Paleontology 19: 497 - 520." type="journal article" year="1999">Carr, 1999</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D5303682FF866A1DC775FAB85D05F905" blockId="21.[826,1441,1361,1903]" pageId="21">
|
||
On the other hand, the hypothesis that
|
||
<taxonomicName id="128F4D01FF866A1DC138FAB85CCDFA0E" authorityName="Rusconi" authorityYear="1933" box="[1310,1441,1514,1535]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC138FAB85CCDFA0E" box="[1310,1441,1514,1535]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
could bear more post-caniniform teeth than recognized by
|
||
<bibRefCitation id="B11E4B73FF866A1DC79DF9755D1DF9CC" author="Rusconi C" box="[955,1137,1575,1597]" pageId="21" pagination="1 - 64" refId="ref15855" refString="Rusconi C. 1933. Sobre reptiles cretaceos del Uruguay (Uruguaysuchus Aznarezi [sic] n. g. n. sp.) y sus relaciones con los notosuquidos de Patagonia. Boletin del Instituto Geologico del Uruguay 19: 1 - 64." type="journal article" year="1933">Rusconi (1933)</bibRefCitation>
|
||
received support when FC-DPV 2320 was prepared. Indeed, the specimen described herein (
|
||
<figureCitation id="4DB42A07FF866A1DC02BF9365D37F98B" box="[1037,1115,1636,1658]" captionStart="Figure 16" captionStartId="21.[164,243,1169,1188]" captionTargetBox="[169,1439,201,1134]" captionTargetId="figure-279@21.[164,1445,196,1140]" captionTargetPageId="21" captionText="Figure 16. Schematic diagram (not intended to reflect the real size or shape differences among teeth) depicting maxillary teeth in U. terrai and three specimens of U. aznarezi (the holotype being probably the only adult individual) drawn at the same size. Based on descriptions and drawings provided by Rusconi (1933) in all cases except for FC-DPV 2320. Question marks indicate the possibility of an additional tooth." figureDoi="http://doi.org/10.5281/zenodo.5441718" httpUri="https://zenodo.org/record/5441718/files/figure.png" pageId="21">Fig. 16</figureCitation>
|
||
) showed the presence of at least 12 maxillary teeth (two incisiviforms, one caniniform, and at least nine post-caniniforms), reducing the apparent gap between the tooth count of
|
||
<taxonomicName id="128F4D01FF866A1DC71CF98D5AD3F905" authorityName="Rusconi" authorityYear="1933" box="[826,959,1759,1780]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC71CF98D5AD3F905" box="[826,959,1759,1780]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="128F4D01FF866A1DC7D8F98D5D0EF905" authorityName="Rusconi" authorityYear="1933" box="[1022,1122,1759,1780]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF866A1DC7D8F98D5D0EF905" box="[1022,1122,1759,1780]" italics="true" pageId="21">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D5303682FF866A1EC775F9AF584CFF0B" blockId="21.[826,1441,1361,1903]" lastBlockId="22.[144,758,198,250]" lastPageId="22" pageId="21">
|
||
In conclusion, given that no real differences in the maxillary dentition exist, we regard
|
||
<taxonomicName id="128F4D01FF866A1DC0DEF84E5C0CF8C0" authorityName="Rusconi" authorityYear="1933" box="[1272,1376,1820,1841]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF866A1DC0DEF84E5C0CF8C0" box="[1272,1376,1820,1841]" italics="true" pageId="21">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
as a juvenile individual of
|
||
<taxonomicName id="128F4D01FF866A1DC01CF8695DD2F8A1" authorityName="Rusconi" authorityYear="1933" box="[1082,1214,1851,1872]" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="aznarezi">
|
||
<emphasis id="E7FBEA90FF866A1DC01CF8695DD2F8A1" box="[1082,1214,1851,1872]" italics="true" pageId="21">U. aznarezi</emphasis>
|
||
</taxonomicName>
|
||
. Thus, as proposed by
|
||
<bibRefCitation id="B11E4B73FF866A1DC742F80B5A80F89E" author="Soto M" box="[868,1004,1881,1903]" pageId="21" refId="ref15901" refString="Soto M. 2005. Especulaciones sobre Uruguaysuchus (Crocodyliformes, Meso-eucrocodylia). VIII Jornadas de Zoologia del Uruguay, Montevideo. Actas, 108." type="book" year="2005">Soto (2005)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="B11E4B73FF866A1DC013F80B5C1FF89E" author="Andrade MB & Bertini RJ" box="[1077,1395,1881,1903]" pageId="21" pagination="21 - 22" refId="ref14265" refString="Andrade MB, Bertini RJ. 2005. Bibliographic revision of Uruguaysuchus (Mesoeucrocodylia, Crocodylomorpha): is Uruguaysuchus terrai a valid species? II Congresso Latino- Americano de Paleontologia de Vertebrados, Rio de Janeiro. Boletim de Resumos, 21 - 22." type="book chapter" year="2005">Andrade & Bertini (2005)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="128F4D01FF866A1EC1A0F80B59BCFF2A" authorityName="Rusconi" authorityYear="1933" class="Reptilia" family="Uruguaysuchidae" genus="Uruguaysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="22" order="Crocodylia" pageId="21" phylum="Chordata" rank="species" species="terrai">
|
||
<emphasis id="E7FBEA90FF866A1EC1A0F80B59BCFF2A" italics="true" lastPageId="22" pageId="21">U. terrai</emphasis>
|
||
</taxonomicName>
|
||
must be considered a junior synonym of the latter taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |