170 lines
16 KiB
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170 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.386.6913" ID-GBIF-Dataset="3a2df501-1763-4f11-a2ea-f9abaac99c00" ID-PMC="PMC3970064" ID-Pensoft-Pub="1313-2970-386-29" ID-PubMed="24693214" ID-ZBK="357FE9805295436EB40CFDD307D00D48" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-386-29" ModsDocOrigin="ZooKeys 386" ModsDocTitle="The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species" checkinTime="1451246199718" checkinUser="pensoft" docAuthor="Kerr, Peter H." docDate="2014" docId="1610145B46F0F8957244CD1E4EFAE0ED" docLanguage="en" docName="ZooKeys 386: 29-83" docOrigin="ZooKeys 386" docSource="http://dx.doi.org/10.3897/zookeys.386.6913" docTitle="Megophthalmidia misericordia Kerr, 2014, sp. n." docType="treatment" docUuid="F10D2F4C-08BC-4EA6-9994-7EC0F1F36BB4" docUuidSource="ZooBank" docVersion="4" lastPageNumber="49" masterDocId="FFE4CC07FFFAC50AFFB0FFF4FFD6FFF4" masterDocTitle="The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species" masterLastPageNumber="83" masterPageNumber="29" pageNumber="45" updateTime="1668157894205" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kerr, Peter H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>386</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.386.6913</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.386.6913</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-386-29</mods:identifier>
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<mods:identifier type="ZBK">357FE9805295436EB40CFDD307D00D48</mods:identifier>
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<mods:identifier type="ZooBank">357FE9805295436EB40CFDD307D00D48</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152051508" LSID="urn:lsid:zoobank.org:act:F10D2F4C-08BC-4EA6-9994-7EC0F1F36BB4" httpUri="http://treatment.plazi.org/id/1610145B46F0F8957244CD1E4EFAE0ED" lastPageId="20" lastPageNumber="49" pageId="16" pageNumber="45">
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<subSubSection pageId="16" pageNumber="45" type="nomenclature">
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<paragraph pageId="16" pageNumber="45">
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<taxonomicName LSID="http://zoobank.org/F10D2F4C-08BC-4EA6-9994-7EC0F1F36BB4" class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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<taxonomicNameLabel pageId="16" pageNumber="45">sp. n.</taxonomicNameLabel>
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Figs 41-50
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45" type="type material">
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<paragraph pageId="16" pageNumber="45">Type material.</paragraph>
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<paragraph pageId="16" pageNumber="45">
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Holotype: ♂, "USA: CA: Sonoma Co., Annadel SP, 0.9mi from park lot, Richardson trail,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="38.435165">38°26.11'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-122.61117">122°36.67'W</geoCoordinate>
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220masl 6m MT, 3-26.iv.2007 P. Kerr & S. Blank 07LOT029" / "HOLOTYPE 12J963,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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♂, Kerr, 2014" [red label]. Deposited in CSCA, specimen glued directly to the pin, complete specimen (Fig. 41). See Fig. 103 for image of type locality.
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</paragraph>
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<paragraph lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45">
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Paratypes (all bearing blue paratype labels): ♂, "USA: CA: Sonoma Co., Annadel SP, 0.9mi from park lot, ravine near Warren Richardson trail,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="38.435165">38°26.11'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-122.61117">122°36.67'W</geoCoordinate>
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, 220masl, 6m MT, 26.
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<normalizedToken originalValue="iv–">iv-</normalizedToken>
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17.v.2007 P. Kerr & S. Blank 07LOT049" [CSCA; specimen # 07Y264 (dissected)]; 9 ♂♂, "USA: CA: Sonoma Co., Annadel
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<pageBreakToken pageId="17" pageNumber="46" start="start">SP</pageBreakToken>
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, 0.9mi from park lot, Richardson trail,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="38.435165">38°26.11'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-122.61117">122°36.67'W</geoCoordinate>
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220masl, 6m MT#3, 17.
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<normalizedToken originalValue="v–">v-</normalizedToken>
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7.vi.2007 P. Kerr & S. Blank 07LOT196" [1 ♂ LACM; 8 ♂♂ CSCA including specimen numbers 12J984, 12J985 (Figs 42-50); four in alcohol, including # 12J958, # 12K739].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="46" type="diagnosis">
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<paragraph pageId="17" pageNumber="46">Diagnosis.</paragraph>
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<paragraph pageId="17" pageNumber="46">
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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sp. n. may be confused with Nearctic congeners that also have a brown thorax and short and broad (stout) apical epandrial processes. Among these, it is most similar to
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia ignea" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="ignea">Megophthalmidia ignea</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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but may be distinguished from these species by having setose apical epandrial processes (not bare; Fig. 43) and narrow, elongate bifurcations of the aedeagal fork (Fig. 49). The narrow, elongate bifurcations of the aedeagal fork in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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are similar to those found in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia browni" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="browni">Megophthalmidia browni</taxonomicName>
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, but
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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is distinguished by having stout apical epandrial processes (not elongate).
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia radiata" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="radiata">Megophthalmidia radiata</taxonomicName>
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also has stout apical epandrial processes, but these are much broader at their base than in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="19" lastPageNumber="48" pageId="17" pageNumber="46" type="description">
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<paragraph pageId="17" pageNumber="46">Description.</paragraph>
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<paragraph pageId="17" pageNumber="46">Male. Body length: 2.0-2.3, 2.1 [2.2] mm (n=6). Wing length: 2.0-2.2, 2.1 [2.1] mm (n=6).</paragraph>
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<paragraph pageId="18" pageNumber="47">
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<pageBreakToken pageId="18" pageNumber="47" start="start">Coloration</pageBreakToken>
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(Fig. 41). Head dark brown; antennal scape dark brown, pedicel and flagellomeres brown; face dark brown, clypeus and labrum brown to dark brown; palps and labellum cream-colored to pale yellow (palpomeres 1 and 2 usually slightly darker than others, palpomere 2 with light patch where sensilla present). Thorax brown to dark brown throughout; scutum setae brown. Coxae lighter in color than thorax, brown; femora light brown to brown, becoming gradually darker dorsoapically; tibiae light brown to brown (hind tibia darkest), hind tibial comb dark brown, tarsi light brown to brown. Wing hyaline without markings, wing veins brown; haltere stem and knob white to cream-colored. Abdominal segments concolorous brown to dark brown. Terminalia brown.
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</paragraph>
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<caption pageId="18" pageNumber="47">
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<paragraph pageId="18" pageNumber="47">
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Figure 41.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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sp. n., habitus [holotype male, # 12J963; female unknown]. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="18" pageNumber="47">
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Head. Ocelli slightly raised, median ocellus in line with anterior margin of lateral ocelli, median ocellus approx. 0.3
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<normalizedToken originalValue="–0.5×">-0.5x</normalizedToken>
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size of lateral ocelli; lateral ocellus located approx. 2
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<normalizedToken originalValue="×">x</normalizedToken>
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diameter of ocellus from eye margin, separated from median ocellus by approx. 2
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<normalizedToken originalValue="–2.2×">-2.2x</normalizedToken>
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its own diameter. Eyes with microsetae, which are approximately as long as width of facet. Frons microtrichose, without setae, flattened. Antennal length 0.7-0.8, 0.8 [0.7] mm (n=6). Face clearly longer than wide, setose; clypeus and labrum microtrichose, without setae. Palpus with four palpomeres; palpomere 1 barrel-shaped, without setae; other palpomeres with brown setae; palpomere 2 bearing small pocket of sensilla; palpomere 1 length longer than or subequal in length to palpomere 2; palpomere 3 length subequal to or slightly shorter than combined length of palpomeres 1 and 2; palpomere 4 length 0.8
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<normalizedToken originalValue="–1×">-1x</normalizedToken>
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combined lengths of palpomeres 1-3.
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</paragraph>
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<paragraph pageId="19" pageNumber="48">
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<pageBreakToken pageId="19" pageNumber="48" start="start">Thorax</pageBreakToken>
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. Dorsum with evenly-distributed, short, appressed setae, bearing longer setae only along lateral and posterior margins. Antepronotum, proepisternum, and laterotergite bearing setae; remaining lateral thoracic sclerites bare. Costal wing vein extends beyond R5, approx. two-thirds distance between R5 and M1; R1 approximately the same length as r-m; cubital fork proximad of r-m base (as in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia occidentalis" order="Diptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="occidentalis">Megophthalmidia occidentalis</taxonomicName>
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, Fig. 52); R1, M1, M2, CuA1, and CuA2 with setae on upper surface (lacking setae on M1 + M2). Wing veins A1 and CuP absent.
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</paragraph>
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<paragraph pageId="19" pageNumber="48">
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Male genitalia (Figs 42-50). Epandrium dorsal surface flat or nearly so, without setae medially, posterior margin narrowly emarginate at center (Fig. 44). Posterior processes of epandrium relatively wide, approx. 2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide, separated at base by approx. 0.5
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<normalizedToken originalValue="×">x</normalizedToken>
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width of process, length of setae at base of epandrial processes less than 0.5
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<normalizedToken originalValue="×">x</normalizedToken>
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width of process; apex of posterior process angled to dull point (Figs 43). Gonocoxites as in Figs 45-47. Aedeagal fork bifurcated into elongated tines of similar width; shorter tine broadly-curving upward and back, longer tine s-shaped (Figs 48-50).
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</paragraph>
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<caption pageId="19" pageNumber="48">
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<paragraph pageId="19" pageNumber="48">
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Figures 42-44.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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sp. n., male epandrium [paratype, # 12J985] 42 Lateral view 43 Posterior view 44 Dorsal view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="19" pageNumber="48">
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<paragraph pageId="19" pageNumber="48">
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Figures 45-47.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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sp. n., male hypandrium [paratype, # 12J985] 45 Lateral view 46 Dorsal view 47 Ventral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="19" pageNumber="48">
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<paragraph pageId="19" pageNumber="48">
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Figures 48-50.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia misericordia" order="Diptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="misericordia">Megophthalmidia misericordia</taxonomicName>
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sp. n., male aedeagus [paratype, # 12J985] 48 Lateral view 49 Dorsal view 50 Posterior view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="19" pageNumber="48">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="49" type="etymology">
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<paragraph pageId="20" pageNumber="49">
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<pageBreakToken pageId="20" pageNumber="49" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="20" pageNumber="49">
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The species epithet
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<normalizedToken originalValue="“misericordia”">"misericordia"</normalizedToken>
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is a noun in apposition, derived from the Latin word for pity/mercy. The species is known only from Annadel State Park, one of 70 California state parks that were scheduled to close in 2012 by California Governor Jerry Brown. Local support has kept this park in operation, but its economic foundation remains uncertain.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |